It’s been too long since I’ve been able to go out with the WGNSS Botany Group on their weekly Monday outing—a consequence of travel and renovations on top of the frenetic-as-usual insect-collecting season. The result is that my attendance on the Botany Group outings is semi-regular during fall/winter but spotty at best during spring/summer. That may seem exactly the opposite of what would be optimum for studying plants, but as a naturalist to the core I have no trouble finding things of interest no matter the season. Especially when the destination is a place as fascinatingly diverse as Johnson’s Shut-Ins State Park—best known previously for its rhyolite “shut-ins” but now mostly for the gashing scour zone that was ripped across it in Dec 2005 when a catastrophic failure of the reservoir atop nearby Proffit Mountain released one billion gallons of water that tore through the landscape in a matter of 12 minutes. The geology exposed by the scour and the living experiment of biological succession that began afterwards are both fascinating, making the Scour Trail one of the Missouri Ozarks’ most interesting day hikes.
Our chief target for the day was Hamamelis virginiana (common or American witch-hazel), which blooms in November and December and is restricted in Missouri to a few counties in the St. Francois Mountains and the extreme southwestern corner of the state. Interestingly, there is a second species of witch hazel—H. vernalis (Ozark witch hazel), more common in Missouri but much more restricted globally—that occurs here, but as it blooms later in winter (January/February) we did not expect to see it on this trip. We found the former reliably, though not abundantly, and among the last plants we found in bloom were some with the freshest (and best-illuminated by the low-angled sun) flowers. At one point while we were still within the dry-mesic upland deciduous forest uphill from the scour zone, we saw a nice colony of the patch-forming Diarrhena obovata (beak grass). This is an attractive grass that does well in shade and should be utilized more as an ornamental.
The overlook provided a stunning overview of the scour zone from an elevated vantage—the since rebuilt Proffit Mountain Reservoir rising ominously above it as an almost deliberate reminder of its potential power—before the descent down into the scour zone. It’s an almost alien landscape with an irregular, unweathered floor of exposed bedrock strewn with rocks ranging from pebbles to boulders. Sycamore and willow are the early leaders in the now 17-year-old race to recolonize the barren swath of land, but lack of toeholds for roots to grow is a bigger problem for this future forest than lack of sunlight by taller neighbors. At one point, we spotted a large bush heavily laden with dense clusters of berries atop a pile of rocks. While the more astute botanists in the group recognized it for what it was, I was dumbfounded as to its identity until it was revealed to me to be none other than Toxicodendron radicans (poison ivy)—the largest, densest, most heavily berry-laden “bush” form of the species I have ever seen. So impressive it was that seven botanists gave it much more than just a trifling look.
About halfway down the scour zone we encountered the “great unconformity”—previously hidden by topsoil and forest but now exposed. Here, knobs of 1.3 billion-year-old granite are surrounded by 540 million-year-old dolomite deposited atop the granite in the shallow Cambrian seas that once covered all but the tallest of these by then already ancient knobs—mere nubs of the towering mountains they once were but worn down nearly to sea level by nearly a billion years of relentless rain and wind. The exposures of pink granite, their large embedded crystals glistening sharply in the sunlight, contrasted starkly with the dark gray dolomite surround them, representing an incomprehensible gap of nearly 800 million years in the record of Earth’s history preserved in the rocks. The entire history of multicellular life on Earth could be swallowed by such a gap!
As an entomologist, I cannot ever stop being on the lookout for insects, no matter what the season. Even though temps were well on the chilly side, I still managed to discern a couple of small wolf spiders, and somehow I managed to see a small ant cadaver on a twig that had succumbed to an insect-pathogenic fungus in the Ophiocordyceps unilateralis complex. Even the botanists around me started taking advantage of the opportunity for insect education. Len and Michael noticed a gall on a small Quercus muhlenbergii (chinquapin oak) which turned out to be the work of Disholcaspis quercusglobulus (round bullet gall wasp), and John noticed a colony of Prociphilus tessellatus (woolly alder aphid) on Alnus glutinosa (European alder). Closer inspection revealed an adult Harmonia axyridis (Asian lady beetle) preying upon the aphids.
It was as enjoyable an outing as I’d hoped (how can four hours in the woods be anything BUT enjoyable), and I hope not to let so much time pass before the next time I’m able to join the group!
After missing the past three weeks, I was finally able to rejoin the Webster Groves Nature Study Society Botany Group for their weekly Monday outing. It was a good outing for making my return, as the group visited one of Missouri’s most famous and unusual landmarks—Elephant Rocks State Park—on what turned out to be a sunny day with unseasonably balmy conditions. Located in Acadia Valley in the heart of the St. Francois Mountains, the park is named for its main feature—one of the mid-continent’s best examples of an unusual geological feature known as a “tor.” These piles of rounded, weathered granite boulders sitting atop a bedrock mass of the same rock resemble groups of elephants lumbering across the landscape. First shaped underground in 1.5-billion-year-old granite as vertical and horizontal fractures developed in the rock and percolating water softened and degraded the rock adjacent to the cracks, the “core stones” were eventually exposed as erosion removed the overlying layers and the disintegrated rock surround the fractures, exposing the giant boulders at the surface.
The group explored the area along the Braille Trail, which passes through dry-mesic upland deciduous forest as it circumnavigates the tor. Oaks and hickories—primarily Quercus alba (white oak), post oak (Q. stellata), blackjack oak (Q. marilandica), Carya ovata (shagbark hickory), and C. glabra (pignut hickory)—dominate the canopy, while the understory featured Viburnum rufidulum (rusty blackhaw), Cornus florida (flowering dogwood), Prunus serotina (black cherry). Unusually abundant also was Nyssa sylvatica (black or sour gum). This small tree reaches its northern limit of distribution near St. Louis, Missouri but is more common further south. Winter bud skills are necessary to recognize the species at this time of year, which can be recognized by their alternate arrangement with three reddish-brown scales and three bundle scars. The leaves of this tree turn a brilliant red in the fall, making them desirable for landscape planting.
Like Nyssa sylvatica, Viburnum rufidulum (rusty blackhaw) is also most common south of the Missouri River. However, in contrast to the former, the winter buds of the latterare immediately recognizable by their dark rusty-colored “velvety” buds and opposite arrangement. The tree we saw at the beginning of today’s outing was also heavily laden with fruits, a dark blue-black pruinose drupe.
As we examined the blackhaw tree, we noticed a robust vine entwining its trunk and ascending high into the canopy above. Heavily laden along its length was a crop of fruits that immediately identified the vine as Celastrus scandens (American bittersweet). This native species can be distinguished from the introduced C. orbiculatus (Oriental bittersweet) at this time of year by its terminal fruits with orange instead of yellow dehiscing valves.
The “fruit” theme of the day continued as we veered off the path to look at a rather magnificent specimen of Quercus marilandica (blackjack oak) and saw yet another vine bearing fruit inside its canopy. The opposite leaf remnants had us quickly thinking of some type of honeysuckle (genus Lonicera), and we arrived at Lonicera sempervirens (trumpet honeysuckle) once we noticed the fused perfoliate leaf pairs directly behind the fruits. This native honeysuckle is a desirable species and not to be confused with any of the several invasive introduced species of honeysuckle that can now be found in Missouri.
Another honeysuckle relative, Symphoricarpos orbiculatus (coralberry), was heavily laden with fruit in the shrub layer. Like Lonicera sempervirens, this species is also native to Missouri and should not be confused with the invasive introduced species—especially Lonicera mackii (bush honeysuckle), which it superficially resembles but can be immediately distinguished from during this time of year by its uniquely coral-colored fruits.
As the Braille Trail wrapped around the eastern side of its loop, we passed by a pile of granite boulders—obvious rubble fragments from the quarrying days of the area’s earlier history due to their sharp, angular shapes. Drill holes could be seen in and around the margins of some of the fragments, providing more evidence of their provenance from rock splitting operations before their eventual abandonment, perhaps not being of sufficient quality to warrant further cutting and shaping into building blocks or paving stones before shipment to St. Louis. Lichens growing sparingly on the cut faces indicated that some amount of time had passed since the stone had been cut, but it was a mere fraction of time compared to the densely colonized original exposed surfaces.
Lichens were not the only forms of life taking advantage of new habitat created by past quarrying activities. Two species of ferns were found growing in protected crevices between the boulders, especially those where water was able to collect or seep from. The first was Asplenium platyneuron (ebony spleenwort), with only sterile fronds present but distinguished by the shiny dark rachis (stem) and stipe (stem base) and alternate, basally auriculate (lobed) pinnae (leaflets). The second was at first thought to be another species of Asplenium, possibly A. trichomanes (maidenhair spleenwort), but later determined to be Woodsia obtusa (common woodsia) by virtue of its all green rachis and stipe and much more highly dissected pinnae arranged very nearly opposite on the rachis. We later found the two species again growing close to each other right along the trail—completely unnoticed despite the group having passed them three times already (i.e., 25 person passes!).
A short spur took us to the Engine House ruin—originally built to repair train engines and cars; its granite skeleton still in good condition—before passing by the park’s main geological attraction: the central tor with its famous “elephants”! Standing atop the exposed granite and boulders, I try to let my mind go back half a billion years—an utterly incomprehensible span of time—when the boulders before me are still part of a giant submerged batholith underneath volcanic peaks soaring 15,000 above the Precambrian ocean lapping at their feet; life already a billion years old and dizzyingly diverse yet still confined to those salty waters.
The landscape atop the tor seems sterile and barren, but like the rubble piles below it’s cracks and crevices abound with life. An especially fruticose stand of Vaccinium arboreum (farkleberry) found refuge in a protected area among some of the bigger boulders, their dark blue fruits continuing the “berry” theme of the day and providing an opportunity for the group to sample their flavor and compare to its cultivated blueberry cousins (I found their flavor to be quite pleasing, if somewhat subdued compared to what is my favorite fruit of all). Vaccinium arboreum is the largest of the three species in the genus occurring in Missouri, and the woody stems of larger plants make it quite unmistakable. Smaller plants, however, can be difficult to distinguish from the two other species, in which case the leaf venation can be used—that of V. arboreum being very open. This is another species that finds itself at the northwestern limit of its distribution in the craggy hills of the Ozark Highlands, where it shows a distinct preference for the dry acidic soils found in upland forests overlying igneous or sandstone bedrocks.
Despite this being a botany group outing, I rarely manage to go the entire time go by without finding and pointing out at least one interesting insect. Today, it was an adult Chilocorus stigma (twice-stabbed ladybird beetle) sitting on a Nyssa sylvatica trunk. This is a native ladybird, not to be confused with the introduced and now notorious Harmonia axyridis (Asian ladybird beetle), that lives primarily in forest habitats and is generally considered to be a beneficial species (although not sold for commercial use in orchards or on farms).
Don Robinson State Park comprises and protects much of the upper watershed of LaBarque Creek in northwestern Jefferson Co.—one of east-central Missouri’s most pristine and ecologically significant watersheds. The St. Peter’s sandstone bedrock underlying the area features box canyons, shelter caves, cliffs, and glades amidst high-quality upland and lowland deciduous forests. The property was originally purchased in the 1960s by businessman Don Robinson, who’s dream was to have a personal sanctuary as large as New York’s Central Park. Through his generosity, the property was bequeathed to the state to become part of Missouri’s state park system following his death a half-century later. The park opened to the public in 2017 and offers some of the highest-quality hiking trails within an hour’s drive from St. Louis. For those interested in more detail regarding the watershed’s geology, ecology, and conservation, an excellent summary can be found in the recently issued LaBarque Creek Watershed Conservation Plan by Friends of LaBarque Creek Watershed.
Here are a few photos from along the Sandstone Canyon Trail.
The flora along the riparian corridor inside the box canyons was of particular interest to me, as it contained nice stands of three tree species of note: Betula nigra (river birch), Ostrya virginiana (eastern hop hornbeam), and Carpinus caroliniana (blue beech, musclewood, American hornbeam). All three species belong to the family Betulaceae and have been associated with some interesting woodboring beetle species in Missouri. I have reared large series of Anthaxia (Haplanthaxia) cyanella from fallen branches of B. nigra (both blue and bronze color forms—see MacRae 2006), and in the course of doing so I also reared a series of an Agrilus species that turned out to be undescribed (to which I later gave the name Agrilus betulanigrae—see MacRae 2003). From O. virginiana, I have reared two specimens of Agrilus champlaini from galls on living trees (still the only known Missouri specimens of this species—see MacRae 1991). Finally, from dead branches of C. caroliniana, I have reared Agrilus ohioensis (see Nelson & MacRae 1990), and from a larger, punkier dead branch I reared a single Trachysida mutabilis—this also still the only known specimen from Missouri (see MacRae & Rice 2007). I think I’ll go back in late winter to early spring and see if I can find dead branches of each to place in rearing boxes or perhaps girdle some branches to leave in situ for a season before retrieving and placing in rearing boxes. Who knows, maybe I’ll get lucky with additional new finds.
MacRae, T. C. 1991. The Buprestidae (Coleoptera) of Missouri. Insecta Mundi 5(2):101–126.
MacRae, T. C. 2003.Agrilus (s. str.) betulanigrae MacRae (Coleoptera: Buprestidae), a new species from North America, with comments on subgeneric placement and a key to the otiosus species-group in North America. Zootaxa 380:1–9.
MacRae, T. C. 2006. Distributional and biological notes on North American Buprestidae (Coleoptera), with comments on variation in Anthaxia (Haplanthaxia) cyanella Gory and A. (H.) viridifrons Gory. The Pan-Pacific Entomologist 82(2):166–199.
MacRae, T. C. & M. E. Rice. 2007. Biological and distributional observations on North American Cerambycidae (Coleoptera). The Coleopterists Bulletin 61(2):227–263.
Nelson, G. H. & T. C. MacRae. 1990. Additional notes on the biology and distribution of Buprestidae (Coleoptera) in North America, Part III. The Coleopterists Bulletin 44:349–354.
Despite the relatively long drive from St. Louis, a healthy group of 15 showed up for this past Monday’s WGNSS Botany Group outing at Hughes Mountain Natural Area; participation no doubt helped out by a spectacular forecast (sunny with highs in the 70s) and near-peak fall colors. Hughes Mountain is situated in the northern portion of the St. Francois Mountains. At its summit is Devil’s Honeycomb—a barren expanse of uniquely fractured Precambrian rhyolite formed by the gradual cooling of magma inside a volcano that was then exposed over 1.5 billion years of erosion. Devil’s Honeycomb is one of Missouri’s geologic wonders, and it’s rocks are among the oldest exposed rocks in all of North America.
Rocks are not the only items of interest here; the igneous substrate results in acidic conditions that affect the flora in equally interesting ways. This is most pronounced in the igneous “glades” (more properly called xeric igneous prairies) where the soils are too thin and conditions too dry to support the growth of trees, offering refugia for grasses and other herbaceous plants more typical of the western grasslands to persist. Surrounding the glades are dry and dry-mesic upland deciduous forests of oak and hickory featuring a rich shrub layer and open woodland-adapted herbaceous plants.
Beginning on the trail from the parking lot, John Oliver pointed out a stand of tall, now leafless sumacs which nearly everybody (including this author) assumed to be Rhus glabra (smooth sumac) due to their size. In fact, despite their size, they proved to be R. copallinum (winged sumac), with the ID confirmed by a few persisting leaves and their distinctive axial “wings.” John pointed out that an easy winter ID tip for this species is the fruiting structures, which nod distinctively after first frost (those of R. glabra do not).
Ascending the trail through the dry-mesic forest towards the first set of glades, we noted the brilliant colors of small Acer rubrum (red maple) saplings in the understory. When their leaves finally drop, they will be more difficult to distinguish from A. saccharum; however, their rounded rather than elongated buds will still allow differentiation.
Several of the oaks were examined, with most thinking they were largely Quercus shumardii (Shumard’s oak) and Q. velutina (black oak)—both similar to each other but the latter bearing larger, grayer, pubescent, quadrangular terminal buds. Approaching the glades, Q. marilandica (blackjack oak), Carya texana (black hickory), and Ulmus alata (winged elm) became more abundant, all three much preferring the drier conditions found around the glade margins. An interesting feature of the latter (in addition to the distinctive, corky ridges on the twigs), is the leaves, which are smaller than those of most other elms but tend to grow larger towards the terminus of the twig. They also tend to be much less asymmetrical at their base than other elms.
Very little was left in bloom, but the remnants of recent bloomers were still evident. Solidago petiolaris (downy goldenrod) and Symphyotrichum anomalum (many-rayed aster) were common along the trail and still recognizable, their showy flowers gone and replaced by developing seeds. Hieracium sp. prob. gronovii (beaked hawkweed) was found nestled among mosses perched on a rhyolite shelf, the flowers gone but the leaves still green and distinctively hairy. Hypericum gentianoides (pineweed) was found on the glades proper, most with their stems and leaves turning red but the occasional plant still green enough to allow crushing its stems and enjoying its orange-like fragrance. Bucking the trend, however, was a small patch of Solidago nemoralis (old-field goldenrod), it’s yellow flowers fresh and bright in defiance of the calendar’s call to senescence. A small jumping spider in the genus Phidippus took advantage of the lingering greenery, hiding among the leaves in hopes of finding equally persistent prey.
The benefits of management efforts by the Missouri Department of Conservation in the area’s forests were more evident than ever. Between the first set of glades and the main glades surrounding the summit, a rich shrub layer dominated by Rhus aromatica (fragrant sumac) stretched endlessly under an open woodland of oak and hickory, the latter turning the canopy bright yellow in vivid contrast to the orange and red shrub layer beneath. Such open woodlands were once common in pre-settlement Missouri but are now rare due to the elimination of fire in the landscape and its mediating impacts.
Entering the main glades, the group made their way up towards the summit and Devil’s Honeycomb, while Ted and Sharon stayed back to take a closer look at and photograph a robust colony of Cladonia cristellata (British soldiers) growing under Juniperus virginiana (eastern red-cedar). Lichens, of course, are unique in the world of vegetation in that they are a composite organism—a fusion between a fungus and another organism (usually a green alga or cyanobacterium) capable of producing food via photosynthesis. None of these groups of organisms are considered plants in the modern sense, and, in fact, fungi are more closely related to animals than they are to plants. Nevertheless, the convergence in appearance, habitat, and ecology of lichens with plants puts their study much more in the realm of botany than zoology.
The group arrived at the summit just in time to enjoy spectacular vistas under crystal blue skies with wisps of clouds and the balmiest temperatures one could possibly hope for in early November.
Even though I ended the bait trapping season last weekend, I still plan to get out on a weekly basis to explore some areas that I haven’t been able to visit yet this season. Tops on the list for me is Hughes Mountain Natural Area, an exposed rhyolite dome in the St. Francois Mountains that features dry oak-hickory woodlands surrounding xeric igneous glades.
The main thing I was hoping to see was Tragidion coquus, a spectacular longhorned beetle that seems common in some areas (e.g., Texas) but is rarely seen in Missouri. I saw one here three years ago in late September at the woodland/glade interface, and a recent conversation with fellow cerambyciphile Dan Heffern, who mentioned that they seem to prefer recently burned oak woodlands, makes me think that is why I saw it here (the surrounding woodlands are managed with periodic prescribed burns to stave off woody encroachment of the glade proper).
I hiked along the trail through the forest leading to the main glade, noting an abundance of many-rayed aster (Symphyotrichum anomalum) in bloom and a few persisting blooms on now-rank plants of slender false foxglove (Agalinis tenuifolia).
After reaching the main glade, I stayed along the interface around its entire perimeter, hoping to see one of the beetles either resting on foliage or in flight. It was a good day to look—sunny and relatively warm, but no beetles were seen. In fact, even though we haven’t yet had any frost, there was not a lot of insect activity in general with the exception of marvelously cryptic lichen grasshoppers (Trimerotropis saxatilis), which were common on the glade along with a variety of other grasshoppers.
There was also little blooming on the glade, which made the chances of seeing the beetle even lower since they are known to be attracted to flowers such as thoroughwort (Eupatorium spp.) and blazingstar (Liatris spp.). I did find a few persisting blooms of the hot-pink largeflower fameflower (Phemeranthus calycinus), but most other plants were well past bloom. Eventually I completed the walk around the glade perimeter and worked my way back.
At one point, I found a clump of small shagbark hickories (Carya ovata) that were oozing sap at several points along the main trunks and noted a variety of insects feeding at the sap flows. I checked carefully, thinking that they might include T. coquus, but none were seen—just flies, butterflies, and a wheel bug assassin bug (Arilus cristatus). Shortly afterwards, I reached the car—my bottles empty but my soul nourished by another day surrounded by nature.
Alternative title: Rich and Ted’s “Excellenter” Adventure.
This is the ninth “Collecting Trip iReport”; this one covering a 10-day trip to western Texas from April 27 to May 6, 2021 with friend and local collecting buddy Rich Thoma. Rich and I have done many shorter collecting trips (up to five days) throughout Missouri and in the neighboring states of Kansas, Nebraska, and Oklahoma (in fact, our first joint trip was to Barber County, Kansas way back in May 1986!). This trip, however, was our first truly long one together—10 days of collecting plus a travel day on each end. To take full advantage of the amount of time we had, we chose western Texas; an area that I have visited several times from the mid-90s through 2004 but not since. We wanted to make the trip during early to mid-May, but scheduling conflicts forced us to go earlier. I reasoned that even if it was a bit too early in the season, I could still collect infested wood for rearing—as I did with great success during my April 2004 trip. For Rich, who is more of a general insect collector, the trip provided him an opportunity for extended collecting in an area that he’d not previously spent a lot of time.
Day 1 – Monahans State Park, Shin Oak Picnic Area First stop of the trip. I was hoping to see beetles on flowers and maybe some tiger beetles, but unfortunately the area hasn’t had any rain yet this spring (according to the ranger). The mesquite was in bloom, but the only beetles I beat from it were a few tiny weevils. A few other plants were in bloom, but only one—Hymenopappus flavescens—had beetles on it (mordellids, which I picked up for Enrico Ruzzier). After a lot of walking I noticed Quercus havardii (shin oak) with flagged branches of dead leaves—a bit of investigation revealed it had been attacked by what must be Chrysobothris mescalero, so I collected as many flagged branches as I could find (7 total) and will bring them back for rearing.
Monahans State Park, Sandhills Picnic Area The big dunes are in this area. We didn’t expect to see any insects but brought our nets anyway. As we were walking the ridge we saw two grouse-like birds in the distance. We tracked them for a bit before I decided to go back and get my binoculars. They kept us at bay, but eventually I was able to get close enough to get a good look at them—they turned out to be scaled quail, a new bird for me. We continued tracking them and eventually they were joined by two more individuals. Handsome birds!
Monahans State Park, Shin Oak Picnic Area After going into town and picking up some dinner, we came back out to the park and setup the ultraviolet lights. I didn’t have much optimism based on the lack of insect activity we saw during the day, but the temperatures were still plenty warm (well into the 80s) and we had nothing better to do. We returned to the Shin Oak Picnic Area since it had a mix of open and more vegetated dunes. Glad we did because two male Prionus arenarius, one Megacyllene antennata, and a tiny, unidentified elaphidiine came to the lights. I also found two small darkling beetles crawling on the sand nearby. I searched the surrounding sand hoping to find more males looking for females, or perhaps even a female herself, but found none. Wolf spiders, however, were common on the sand, their glowing eyes drawing attention beyond their abundance. I guess they are a species of Hogna, but I’m not certain—I photographed two individuals. Also got a large bostrichid (Apatides fortis?) at the light. Before we took down the lights, Rich called me over to see a tiny, slender, worm-like snake that we eventually determined was one of the blind snakes (Leptotyphlum sp.)—definitely a first for me.
Day 2 – Toyahyale We stopped here on a tip from Jason Hansen and Tyler Hedlund, who swept good numbers of Agrilus cochisei off of Ambrosia psilostachya (western ragweed)—albeit, a few weeks later during May. I found the plants, but they were very small and low to the ground. Nevertheless, adults could be swept abundantly from the plants, and I was able to take good photos of singles and a mating pair with the big camera (iPhone photo here just to show what they loook like). Also got a single specimen of an apparently undescribed Acmaeodera sp. while sweeping for A. cochisei and one of two A. cochisei adults that I saw on flowers of Sphaeralcea sp.
Davis Mountains, 15.8 mi NE Ft. Davis We stopped here to look for the undescribed species of Acmaeodera, which Jason had found in good numbers during May on blooms of Lygodesmia and Convolvulus. Both plants were present, but neither was in bloom. Still, I found one adult on flowers of Verbina sp. and swept another from roadside vegetation. Ambrosia polystachia (western ragweed) was also present—I looked visually for Agrilus cochisei and did not see any, but I did get one adult and a couple of cryptocephalines in the sweeping that produced the second Acmaeodera.
Day 3 – Point of Rocks Roadside Park The weather turned decidedly cool in the Davis Mountains—first time I’ve ever frozen camping out on a collecting trip. The high temps are expected to stay in the 50s to 60s with a chance of rain for the next few days, so we decided to head down to the Big Bend area where there is still a chance of rain but warmer temps (up to the high 70s). Maybe we’ll come back to this area next week. Before leaving, however, I wanted to check the Quercus vasseyana (vassey oak) at Point of Rocks, where in the past I collected a good series of Mastogenius texanus even earlier in April (it was actually undescribed at the time). I’ve also collected Elytroleptus lycid-mimicking cerambycids on soapberry flowers here in June, although I knew the soapberry would not be in bloom. There was nothing on the oaks, but I did collect a few miscellaneous beetles beating Prosopis glandulosa (mesquite) and found one vassey oak branch with evidence of wood-boring beetle larval feeding and which I collected for rearing.
17 mi S Alpine This is the first of two picnic areas along Hwy 118 going south from Alpine towards Big Bend National Park. There are lots of big Quercus vasseyana and Q. grisea here, so I stopped to see if I could find any infested wood. Bingo—one of the Q. grisea had a dead branch hanging from it that looked recently dead, and breaking apart a few of its smaller branches revealed fresh larval workings of some kind of buprestid (perhaps Polycesta arizonica). I cut of the branch and will bring back the bundle for rearing.
26 mi S Alpine This is the second of two picnic areas south of Alpine on Hwy 118 towards Big Bend National Park. Rich and I have both stopped here before, and Rich brought back infested wood (apparently Juglans sp.) from which I reared Chrysobothris comanche, so that was the plan again unless we saw active insects. We did not, so I scanned the trees and found a small Celtis laevigata (sugarberry) that had recently died—the bark was peeling, but there were no emergence holes that I could see. I started chopping into the trunk wood and quickly encountered a large buprestid larvae in its “pre-pupal fold”. This could be Texania fulleri based on host and location, so I cut a couple of bolts from the trunk to bring back for rearing. The branches also showed fresh larval workings, so I cut up one along with its smaller branchlets to also bring back for rearing.
3.3 mi W of Hwy 118 on Agua Fria Rd. Last stop of the day, which I was told could have water with tiger beetles. The creekbed was bone dry, and I collected but a single Chrysobothris sp. beating Prosopis glandulosa in flower—amazing given the proliferation of wildflowers that were in bloom. We did find a nautiloid/ammonite-type fossil in the bone-dry creekbed, which Rich says is of Cretaceous origin based on clam fossils in the underlying layer, and I tracked a common poorwill (Phalaenoptilus nuttallii) for a little bit, eventually getting close enough for the rare iPhone bird photo. Interesting position it assumed upon landing with its wings outstretched above its back.
Big Bend Ranch State Park, West Contrabondo Campground We arrived in Study Butte with just enough time to check into a motel and get dinner before heading out to Big Bend Ranch State Park. The drive through the park was incredible as we searched for a spot to setup the lights. After finding such spot, however, we were greeted as we got out of the car by a stiff, chilly wind. I knew there was no point in going through the trouble to setup, so instead we drove further down the 2-track to an amazing scenic overlook into an impressive box canyon. Words cannot describe the contortions this acrophobiac took to find good position for these photos, but it was well worth the views.
Day 4 – Big Bend National Park, Boquillas Canyon Trail Well, the rain and cold continue to follow us. Rather than trying once again to drive somewhere else to escape, we decided to just sit this day out and visit the national park (not a bad Plan B!). Boquillas Canyon is an amazing slice through the rocks along the course of the Rio Grande River, and we hiked as far into the canyon as we could before sheet rock on the left and deep water on the right prevented any further progress. We saw only two insects—a tiger beetle larva that I “fished” out of one of the many larval burrows we saw (definitely Tetracha, and likely T. carolina) and a velvet ant (black head and pronotum, red abdomen).
Big Bend National Park, near Panther Junction Driving towards the Chisos Mountains after hiking the Boquillas Canyon Trail, we encountered this fine adult male Aphonopelmis hentzii (Texas tarantula) crossing the road.
Big Bend National Park, Chisos Basin, Window Trail After lunching at Panther Junction, we headed up the into the Chisos Mountains towards Chisos Basin. Heavy clouds shrouded the peaks, so we weren’t sure what we would encounter up there, and once in the cloud zone and then heading down into the basin we could hardly see anything. Suddenly the western side of the basin came into view, still overcast and drizzly but at least free from the heavy fog that shrouded the eastern half of the basin. That made our decision of which trail to hike easy—the Lost Mines Trail under heavy fog versus the Window Trail with semi-clear views. I’ve hiked the Window Trail several times, but the last time was 17 years ago, and Rich in his single attempt a year or two later did not make it to the “Window” due to an impatient 10-year old son in tow. The views on the way down the canyon were spectacular—not despite the rain and clouds but because of it. It is a rare opportunity to see richly moist desert mountains shrouded in mist. At one point on the way down, a Woodhouse’s scrub jay (Aphelocoma woodhouseii) caught our attention—sitting very nearby in a tree before hopping down to the ground and nonchalantly pecking for bugs. Another soon joined him, first landing on a branch just a few feet above me and returning my captivated stare for a few moments before joining his mate on the trail ahead of us… followed shortly by a third individual. Their soft chirpings were a charming contrast to their more familiar raucous calls, and Rich and I soaked in the moment until they moved on. The trail is not an easy hike—nearly 7 miles round trip, dropping over a thousand feet on the way down, and then gaining over a thousand feet on the way back. The “Window,” however, is a sight to behold—a narrow gap in the rocks soaring high overhead with a view out onto the desert floor almost a thousand feet below. There is tempting danger at the window—its smooth, water-carved rocks are deceptively slippery even in dry conditions, and with the rain of the day they were especially so. I would not be surprised to learn that at least one person had made a fatal error in judging how close to the window one can get. They would have had plenty of time to think about that mistake on the way down! The views on the way back up were even more breathtaking, as fog enshrouded the high peaks towering above us. Periodically the sun attempted to push through the clouds, creating surreal lighting in a battle of sun versus rain, but eventually the rain won out and fell steadily on us for the last, switchback-laden mile back to the trailhead. As for insects, we actually did see some despite the rain—a few blister beetles resting torpidly on yellow composite flowers.
Day 5 – Big Bend National Park, Sotol Overlook We’re on our way to Santa Elena Canyon and stopped at this overlook. From a distance of 14 air miles, the canyon entrance looks like a tiny split in the rocks, belying the 1000-foot canyon walls that await us. Cacti were nicely in bloom, if a bit rain battered—two species of yellow-flowered Opuntia (pricklypear) and the always extraordinary pink flowers of Cylindropuntia imbricata (tree cholla). No insects were to be found, but we did find a live Orthoporus ornatus (desert millipede)—the first that we’ve seen on this trip—who obliged us by coiling into its classic defensive pose.
Big Bend National Park, Santa Elena Canyon From 14 air miles away, Santa Elena Canyon looks like a tiny split in a little cliff (see previous post). Up close, however, it’s soaring walls tower 1000 feet overhead! The hike into the canyon features a tortuous staircase to bypass a narrows, followed by a leisurely stroll along the canyon bottoms along the Rio Grande River. Rain last night has triggered an en masse millipede emergence, and even a few insects were seen: velvet ant; Acmaeodera mixta, Trichodes sp. and Gnathium sp. on yellow composite flowers; and Omorgus sp. crawling in the sand.
Big Bend National Park, Cerro Castellan (Castolon Peak) The layers visible in Cerro Castellan reveal millions of years of volcanic events. Stacked in this tower are several lava flows and volcanic tuffs (ash deposits), with layers of gravel and clay from periods of erosion between eruptions. Cerro Castellan’s cap rock is the same lava that formed the Chisos’ South Rim. The lighter orange and gray layers beneath are tuffs.
Big Bend National Park, Tuff CanyonTrail Some of the oldest layers of volcanic rocks lie at the bottom of Tuff Canyon. It is dry most of the time, but summer thunderstorm runoff churns through the canyon, cutting it deeper. This canyon is narrower and deeper than most others in Big Bend, partly because the light gray volcanic tuff is relatively cohesive. Swift, powerful floodwaters will cut down through any kind of bedrock, but the tuff is better able to resist the widening effects of sideward erosion.
4.6 mi W of Langtry After leaving Big Bend National Park we started making our way towards the Del Rio area in Val Verde Co., where we plan to meet up tomorrow with a few other beetle collectors (Dan Heffern, Brian Raber, and Ed Riley). I noted that our path took us right by the type locality of the recently described tiger beetle, Amblycheila katzi. I didn’t have much hope of actually seeing the species, given that the season seems to have even really started yet and the earliest record of the species is the 23rd of May. Nevertheless, since we happened by the spot right as dusk was falling it seems a good idea to at least try. First we walked the limestone 2-track just to see what was out and about (just a few darkling beetles), then we started checking the limestone ledges where the tiger beetle can be found. We checked about 100 m of ledge without seeing any, and I was about ready to call it a night when we finally spotted one. It was running in a seam about 2 m above the ground and was unmistakable. My attempt tiger an in situ photo failed at first, and it almost escaped deep into a crevice before I pulled out my long forceps and pulled him out by a tarsus. It gave me a healthy pinch when I grabbed it while fumbling in my pack for a bottle, but eventually I prevailed. Later on I placed it back on the ledge and covered it with the Nalgene bottle cap, waited for it to calm down, then carefully lifted the cap and got a couple of shots before it began scurrying again. We checked another 50 m of ledge without seeing any and decided to call it a night.
Day 6 – Comstock (prologue) The owner of the motel in which we stayed was super friendly and kind enough to leave a key in the door for our very late arrival last night. Settling up this morning, I saw this on the wall (right next to his vaccination card—two doses) and just had to get a pic. He was only too happy to oblige when I told him how awesome it was and could I get a picture. Hey, no reason to reveal true political leanings if it means we can all just get along.
Amistad National Recreation Area, Spur 406 Campground A quick stop here on the way to meet up with Dan, Brian, and Ed near Devil’s River. There were lots of dead and dying Acacia constricta (whitethorn acacias), off of which I beat a diversity of cerambycids and buprestids from both the dead and dying branches. I found one small sapling of the same with evidence of fresh woodboring beetle larval feeding, so I collected it as well for rearing. Other than that I just collected a few weevils off of living Prosopis glandulosa (mesquite) for weevil-specialist Bob Anderson.
Devils River near Dry Devils River We met up with Dan, Brian, and Ed north of Del Rio and, after exchanging peasantries, followed them into a private resort surrounding a stretch of the Devils River*—considered by some to be the most unspoiled river in Texas. Dan had arranged for access after befriending Dave Barker, a commercial herpetologist who had built a home on property overlooking the Devils River and also a guest cabin on property overlooking nearby Gold Mine Canyon. We met up at the cabin and then carpooled to a spot along the Devils River where Dan and Brian had placed a variety of traps that needed servicing. While they took care of that, Rich, Ed, and I collected in the area around where the traps had been placed. I started off beating dead branches of Vachellia farnesiana (huisache), sweeping blooming Salvia sp. (sage), and beating dead branches of Acacia rigidula (blackbrush acacia) down by the river but collected only a smattering of beetles. I then clambered up the rocks and found good numbers of Acmaeodera spp. visiting flowers of Echinocereus enneacanthus (strawberry cactus) and Opuntia engelmannii (Engelmann’s pricklypear). After collecting my fill of those beetles, I returned to the riverbanks and noticed some large Carya illinoensis (pecan), from which I beat a few Anthaxia (Haplanthaxia) sp. (hoping they are one of the recently described taxa). By then, Dan and Brian had finished servicing their traps and gave me a few specimens that had been collected in their ethanol-baited Lingren funnel trap.
* The accepted usage of the name is without an apostrophe, although the reason for this is a matter of debate.
After finishing along the Devils River, Dave invited us to his home for a few post-collecting beers. Spectacular views overlooking the river.
GoldMine Canyon We setup a variety of light stations at Dave’s cabin a little east of the river. It was warm and dry, so conditions were good, if a bit windy. My two ultraviolet light stations a bit north of the cabin ended up catching the lion’s share of cerambycids, although it was mostly elaphidiines and a Lepturges sp. We also picked up a few tenebrionids and a Carabidae crawling on the ground near the lights. Ed’s mercury vapor/ ultraviolet station on the road west of the cabin attracted a few more cerambycids, including a Lagocheirus sp. Dan, however, got the catch of the night—a Goes that came to Dan’s $6 battery-powered lantern on the road south of Ed’s station. We at first thought it might be G. novus, but that Dan later decided it was just a very lightly marked G. tesselatus. Zephyranthes chlorosolen (Brazos rain lily) blossoms were beautiful at night, their stark whiteness catching the beam of the headlamp.
Day 7 – Gold Mine Canyon Rich and I spent the morning walking the grounds around Dave’s cabin while the others packed up and got ready to leave. I found some oak (Quercus vasseyana) saplings infested with cerambycid larvae, which I cut and bundled to bring back for rearing. Acmaeodera were already coming to the flowers—a couple of small ones on an undetermined white composite, three different species on Opuntia engelmannii (Engelmann’s pricklypear) and Echinocereus enneacanthus (strawberry cactus) flowers, and a couple on Coreopsis? flowers. I beat some of the Diospyros texana (Texas persimmon) looking for Spectralia robusta but did not find any. It got hotter than blazes real quick!
After Dan, Brian, and Ed left, Rich and I went down to the canyon entrance to beat on the oaks and Texas persimmons that dot the sides of the canyon. Nothing was on either plant, however, and I ended up again concentrating on the diversity of Acmaeodera that were coming to flowers of Coreopsis sp., Opuntia engelmannii, and an unidentified yellow composite. I did beat a single Cleridae off a dead branch of mesquite.
Devils Rivernear Dry Devils River After finishing at Gold Mine Canyon, we came back to the Devils River crossing near the first stop we made here for our final stop of the day. Temperatures had maxed out at 99°F! and I wasn’t too motivated to collect much more today, but when we arrived at the spot I noticed some declining Platanus occidentalis (American sycamore) with large emergence holes suggestive of Mallodon dasystomus and old fallen branches with the same suggestive of Polycesta elata. The tree with the Mallodon holes was much too large to cut (and embedded within a thicket of poison ivy), so I occupied myself by collecting a few more Acmaeodera off of Opuntia engelmannii flowers. As I walked the roadway I noticed more sycamore with some smaller trees in the grove that looked recently dead. One was dead from about three feet up and had buprestid workings under loose, peeling bark. I cut just above the live portion (3–4” diameter) and took three 4-ft sections of the trunk above that point, each cut showing internal galleries. If P. elata emerges from these pieces of wood I will be “elated” [Later edit: I did rear the species!].
Gold Mine Canyon For blacklighting tonight we decided to bring the lights down to the mouth of the canyon where we collected this afternoon so we could have acces to anything associated with the oaks. Unfortunately it was a much slower night than last night and cooled off quickly despite the high heat earlier in the day. I only got three cerambycids (one Ecyrus and two Aneflomorpha) and a few clerids at the lights. I also walked the jeep track leading to the mouth of the canyon and the main road outside and didn’t see anything until I almost got back, when I noticed a beetle sitting on the trail that looked a bit odd. When I picked it up I realized it was a buprestid in the genus Melanophila—what the heck?! Totally unexpected to see this beetle at night and especially on the ground instead of on a tree. I suppose it is one of the juniper-feeding species (since pine doesn’t occur here).
Day 8 – Gold Mine Canyon Our plan today was to head over to some spots further west in Val Verde Co., but before leaving the cabin we did a bit of walking around and took a last few photographs.
22 mi N Del Rio I had noted a few scattered plants of Senna roemeriana (two-leaved senna) at this spot a couple of days ago when meeting up with Dan but didn’t have the chance to sample them for Agrilus obtusus—one of my target species for the trip. I found one on the second plant I checked, so I went back to the truck to get my big camera hoping to photograph one in situ. I didn’t see anything on the next plant, but when I tapped it over my net there was another one! I did that for the next hour or so—inspecting and tapping—and never saw another one. Rich did get one sweeping the S. roemeriana (in an area I’d already worked) and was gracious enough to give it to me. There were also tiny bruchids and clerids on the plant. Other than that I got a couple of Acmaeodera mixta sweeping, a couple of Canthon sp. in flight, and a Euphoria kerni on the flower of Zephyranthes chlorosolen.
Hwy 90 at Del Rio River I first visited this spot nearby 30 years ago based on a tip by Dan Heffern, who had reared a Polycesta elata from Fraxinus greggii (Gregg ash). I found the ash on that visit, though I didn’t find any wood infested with that species here, but what I did find was Diospyros texana (Texas persimmon) infested with Spectralia robusta and managed to rear out a few individuals. That was my quarry today, but when I arrived the abandoned road on the northwest side of the bridge was fenced and posted. I took a look on the southwest side and found open access up top and decided to hike down towards the ravine from that point. Things seems to be about as far along here as they were at Devils River, with not much activity except for Acmaeodera coming to the Opuntia engelmannii flowers, albeit not quite the diversity. I found a few more also on flowers of an undetermined yellow composite, Coreposis sp., and an undetermined white composite. Closer towards the ravine I found just a single large F. microphylla (with no signs of infestation) and several D. texana—two of which had the half-live/half-dead branches in which S. robusta larvae live and showing the emergence holes of adults. I collected both branches and will bring them back for rearing.
Amistad National Recreation Area, Pecos River Access Nature Trail Just a quick stop at the Pecos River Access on the east side to walk the short nature trail and gaze at the 300-ft high, 100 million-year-old (Cretaceous Period) limestone bluffs that the Pecos River has cut near the junction with the Rio Grande River (the latter can be seen on the left side of photo 2). The first photo also shows the old road that was originally used to cross the river snaking down the west bluff—traffic today uses the tall bridge in the right side of the photo.
Seminole Canyon State Park, Canyon Rim Trail We came here looking for oak potentially infested with Spectralia roburella. We didn’t find any oak on this trail, but I did find a Acacia rigidula (blackbrush acacia) showing signs of infestation by buprestid larvae (difficult to find such this year because the freeze in February apparently killed or severely knocked back most of this species). I cut up and bundled the wood to bring back for rearing.
Amistad National Recreation Area, Spur 406 Campground We got to Seminole Canyon State Park too late to check with the supervisor about setting up our blacklights at the park, so we came back to Spur 406 Campground where we’d collected a few things two days earlier. Temps were okay and there was no moon or wind, but it was still a very slow night—for me just a couple of elaphidiines, two trogids, two Digitinthophagus gazella (why do I continue to pick these things up?), and a bostrichid.
Day 9 – Seminole Canyon State Park, Windmill Trail We came back to the state park since we ran out of time to look for oak yesterday. The park staff were extraordinarily helpful—both in getting me checked in with my permit and in directing me to the spots where I might be able to find oak. Their first tip—along the Window Trail—paid off, where we found a nice cluster of Quercus fusiformis (plateau live oak) clinging to the upper canyon walls. Most of them had dead branches on them, and I did some beating to see if by some chance the beetles would be out already. They were not, but on the second tree that I examined I found a main branch from near the base with the outer 4–6 ft dead but the bark not peeling and small living sprouts about 2 ft from the base. Pulling apart the dead portion revealed buprestid larval workings, likely my quarry—Spectralia roburella, but these could be old. I cut the branch at the base, however, and found fresh larval galleries in the sapwood of the still-living portion even extending into the trunk—success! I’ll bring this back for rearing and will hopefully get S. roburella out of it. Further along the trail I found a single Senna roemeriana (two-leaved senna), inspected it carefully and didn’t see anything, then tapped the plant over my beating sheet and a single Agrilus obtusus fell onto it to add to the three that I got yesterday. I really wish I could see these things before I beat them off the plants so I could take an in situ photo!
Seminole Canyon State Park, Canyon Rim Trail Another place the park staff recommended to find oaks was along the Canyon Rim Trail. We hiked that trail yesterday for a bit and didn’t see any oaks, but it turns out they were farther down the trail then we went. We headed back out on the trail to find them, along the way checking Opuntia engelmannii flowers for Acmaeodera and seeing only one for the time being. Just past the first of two east-facing ravines where we expected to find oaks, we found one on the canyon edge that looked rather bedraggled. There were some completely dead branches with bark already sloughed but also one large fresher-looking dead branch that had one live branchlet coming out of it about a third of the way up (meaning there was at least a strip of live wood within the branch). I broke of one of the dead branches near the live/dead junction, and there in its gallery was a smallish buprestid larva that almost certainly is Spectralia roburella! I took the entire branch and cut it up to bring back for rearing. We continued hiking along the canyon rim and saw the most amazing views—sheer Cretaceous limestone walls towering 300 feet above the narrow canyon bottoms! Farther down the trail we finally started seeing Acmaeodera on O. engelmannii flowers. By then we’d hiked more than a mile and a half down the trail and temps were beginning to soar, so we turned back, picked up the wood we’d cut as we came back by, and finished the long, hot slog back to the truck.
Comstock As Rich and I were lunching after our last stop, I got a text from Ed Riley about a spot near Comstock where he’d collected what he believed to be Acmaeodera starrae—a species I’ve never encountered. It just so happened that we would be passing by Comstock on our way back east this afternoon, so we stopped to see if we could find it. Bingo—right where and in the flowers he said it would be (an undetermined white composite that I later determined to be Aphanostephus ramosissimus [lazy daisy]). Together we found about 15 specimens, and interestingly about 25% have red rather than yellow elytra markings. [EDIT: I’m not convinced these are A. starrae, but I do not yet know what they are.]
Day 10 – Garner State Park, Wild Horse Creek/Highway/Campos Trails It’s the final day of collecting for the trip, and for our last stop we picked Garner State Park along the Frío River. I was last here back in the mid 90s—nearly 30 years ago, Acmaeodera ornatoides and Polycesta elata being the two species of note that I remember finding. I remember during that first visit that the area reminded me of my beloved Ozark Mountains, especially the White River Hills region in southwestern Missouri—scraggly forests of oak and juniper on steep, rocky slopes over craggy hill and lazy dale. It still does, although the species are a bit different—Juniperus ashei (Ashe juniper) dominates instead of J. virginiana (eastern red-cedar), and a variety of other oaks replace the familiar Ozarkian Quercus stellata and Q. marilandica (post and blackjack oaks, respectively). We hiked a series of trails on the western side of the park, thinking the west-facing slopes would tend to be drier and result in more open, glade-like habitats, and for the most part this was true. Almost immediately after reaching the first glade along Wild Horse Creek Trail, we found A. ornatoides and at least two smaller congeners on flowers of Coreopsis sp. Flowers of Viguiera dentata have been uncharacteristically depauperate of buprestids on this trip, but I picked up a couple of Acmaeodera neglecta/neoneglecta nearby as well. On the Highway Trail a good series of Acmaeodera was found on flowers of an undetermined small white composite, and a few were also found on flowers of Senna roemeriana (two-leaved senna)—though no Agrilus obtusus. The Campos Trail ascended steeply and ruggedly to a nice overlook, where I found one Acmaeodera sp. on the flower of Zephyranthes chlorosolen and then the mother-load—the biggest diversity and abundance of Acmaeodera I’ve ever seen on cactus flowers occurred nearby in a single flowering Opuntia engelmannii. The final specimen of the day’s “Acmaeodera-a-thon” was taken a bit further up the trail on the flower of Echinocereus enneacanthus (strawberry cactus). Fortunately, the trail was all downhill from there (albeit a bit too steep and rocky at times for these no-longer-nimble legs!). We finished off the hike back along the Wild Horse Creek Trail by collecting a branch off a fallen oak that I hope proves fruitful in the rearing box back home and had some lunch. As we were getting ready to leave, I noticed most of the trees in the camping area were Carya illinoensis (pecans)—a great host for buprestids (especially Xenorhipis brendeli), so I picked up several fallen branches from under the trees to complete the wood collecting portion of the trip.
Garner State Park, Brazos River (epilogue) We visited the nearby Frio River for one last look at the park, took a shower, and settled in for the 15-hour trek back to St. Louis.
This month’s destination for the WGNSS Natural History Photography Group was Orchid Valley Natural Area in Hawn State Park. This natural area is south of the main park and not normally open to the public, but we were granted permission to enter by the park administration. Our targets were several species of orchids and other rare plants that are known to occur in the area—showy orchis (Galearis spectabilis) being the one I hoped most to see.
There are no trails here—not even a place to park, as we squeezed our cars against the side of the road at a spot that appeared to provide good access. A bit of GPS-guided bushwhacking brought us to increasingly rough and sloping ground that ended up at the edge of a cliff overlooking one of the sandstone box canyons that this area is know for—down below was where we wanted to go. Wild azaleas lined the upper canyon edges with their stunning pink blossoms. We followed the canyon edge and found a way down, then circled back into the canyon to find a stunning waterfall, its sandstone walls dripping with mosses and ferns. We spent quite a bit of time here photographing the waterfall and surrounding area before eventually resuming the search for the orchids that we came to see.
Sadly, the orchids were not yet in bloom—not even close, another victim of the cold, late spring we’ve been experiencing. Casey, our group leader, did find some very small showy orchis leaves, and we saw some nice clumps of another native orchid, rattlesnake plantain (Goodyera pubescens), an evergreen orchid that blooms during late summer. We hiked up another drainage that led to another box canyon—lacking a waterfall but equally impressive, nonetheless—but found no orchids in bloom there, either. We did, however, see cinnamon ferns sending up their spike-like fertile fronds and aggregations of antlion larvae (a.k.a. “doodlebugs”) in the soil beneath the sandstone ledges.
You might think failure to find what we were looking for would result in a disappointing field trip—far from it! Time in the field with like-minded friends in a beautiful spot is always a pleasure, and when it comes to searching for rare plants (or insects, or whatever), failure is the norm—making success, when it does come, that much sweeter. There will be other chances to see showy orchis (perhaps in a couple of weeks).
Hot on the heels of the previous installment in this series, I present the sixth “Collecting Trip iReport”; this one covering a trip to Arizona during July/August 2018 with Art Evans and—like the previous installments in this series—illustrated exclusively with iPhone photographs (see previous installments for 2013 Oklahoma, 2013 Great Basin, 2014 Great Plains, 2015 Texas, and 2018 New Mexico/Texas).
This trip was a reunion of sorts—not only had it been 20 years since I’d collected in Arizona, it had also been 20 years since I’d spent time in the field with Art Evans—which just happened to be in southeast Arizona! For years I looked forward to our next opportunity, and when he told me of his plans for an extended trip to take photographs of his forthcoming Beetles of the Western United States, I couldn’t pass up the chance. Art had already been out west for five weeks by the time I landed in Phoenix on July 28th, and together we drove to Cave Creek Canyon in the Chiricahua Mountains and spent the night before beginning a 7-day adventure in and around the “Sky Islands” of southeastern Arizona.
As with the recent New Mexico/Texas post, the material collected still has not been completely processed and curated, so I don’t have final numbers of taxa collected, but there were a number of species—some highly desirable—that I managed to find and collect for the first time, e.g., the buprestids Acmaeodera yuccavora, Agrilus restrictus, Agr. arizonicus, Chrysobothris chiricauhua, Mastogenius puncticollis, and Lampetis webbii and the cerambycids Tetraopes discoideus and Stenaspis verticalis. Who knows what as-yet-unrecognized goodies await my discovery in the still unprocessed material?!
Day 1 – Chiricahua Mountains, Cave Creek Canyon
After arriving at Cave Creek Ranch late last night, we awoke to some stunning views right outside our room!
The first buprestid of the trip was a series of Pachyschelus secedens on Desmodium near Stewart Campground. We beat the oaks and acacia along the way to Sunny Flat Campground but didn’t find much. Once we got near Sunny Flat I did some sweeping in an area with new growth of Helianthus sp. and got a series of Agrilus huachucae, a few lycids, and one Leptinotarsa rubiginosa. I beat one Acmaeodera cazieri from Acacia greggii and found another on flower of prickly poppy (Argemone sp.). On the roadside at Sunny Flat I found several Acmaeodera spp. on a yellow-flowered composite – one A. rubronotata, one A. solitaria(?), and three A. cazieri. Also collected one A. cazieri on a rain gauge, Mecas rotundicollis and one as yet undetermined acanthocinine cerambycid on miscellaneous foliage, one tiger beetle (Cicindela sedecimpunctata?) on the roadside, and two orange lycids in flight.
Desert flats east of Portal, Arizona
We came to this spot to look for Sphaerobothris ulkei on joint-fir (Ephedra trifurca), but after not finding any for awhile I got distracted by some big buprestids flying around. Caught several Hippomelas sphenicus, one Gyascutus caelatus, and two Acmaeodera gibbula on Acacia rigida, and the first and third were also on Prosopis glandulosa along with Plionoma suturalis. We finally found S. ulkei – searched the area for almost three hours, and Art and I each caught two and Margarethe caught one – also one each of P. suturalis and A. gibbula. I also got a mating pair of A. gibbula on Acacia greggii. After dinner, we went back and placed an ultraviolet light – checked it a couple hours later and got a nice series of Cylindera lemniscata and a few meloids (for Jeff).
Day 1 of the trip ended in typical monsoon fashion – heavy, thunderous rainstorms moved into the area during late afternoon, dimming prospects for blacklighting. Still, we set them up anyway at several spots and checked them later in the evening (flood waters preventing us from going to all the spots we wanted to). Not surprisingly, the one trap that yielded interesting specimens was in the lowest (warmest) area and received the least amount of rain. For me it was a nice series of Cylindera lemniscata.
Day 2 – Southwestern Research Station, Chiricahua Mountains, Arizona
There is a large stand of a narrow-leaved milkweed (Asclepias sp.) at the station, so we stopped by in our way up the mountain to check it for beetles. Got a nice little series of Tetraopes discoideus (tiny little guys!) on the stems as well as a few Rhopalophora meeskei, two Lycus spp., and one Pelonides humeralis on the flowers.
At the Southwestern Research Station with Barbara Roth, Art Evans, and Margarethe Brummermann.
Road from Southwestern Research Station to Ruster Park After leaving the SWRS on our way up to Rustler Park, we stopped to check a couple of bushes of New Mexico raspberry (Rubus neomexicanus). Margarethe thought there might be lepturines on the flowers, but instead we found a few Acmaeodera spp. and some Rhopalophora meeskei.
Further up the road we made another quick stop to check roadside flowers – just a single A. rubronotata on a yellow-flowered composite, but spectacular views of the valley below.
Gayle Nelson once told me about finding Chrysobothris chiricahuae on pine slash at Rustler Park, so I was pleased to see several fresh slash piles when we arrived. I saw a Chrysobothris (presumably this species) on the very first branch in the very first pile that I looked at, but I missed it (damn!) and didn’t see any more in that pile. However, in the next pile I visited I saw two and got them both. I looked at a third pile and didn’t see any, nor did I see any more on the two previous piles that I looked at. Still, two is better than none (assuming this is, indeed, what they are!).
Chiricahua National Monument
Not a bug collecting stop, but we wanted to drive into the monument and see the incredible rock formations which are best appreciated by driving through Bonita Canyon and then up to Massai Point. The unusual spires, columns, and balancing rocks are a result of erosion through vertical cracks in the compressed volcanic ash which was laid down in layers 25 million years ago and then uplifted. Tilting during uplift caused vertical fractures and slippage, into which water then worked its way to create today’s formations. One of the columns I saw is 143 feet tall and only 3 feet in diameter at one point near the base! Mexican jays were our constant, close companions as we hiked through the pinyon pine/oak/juniper woodland.
Vicinity Gleeson, Arizona
There is a wash across N Ghosttown Trail with stands of Baccharis sarothroides growing along the sides. Art previously collected a single Cotinis impia on one of the plants, so we came back to check them. We didn’t find any, but we did find two fine males and one female Trachyderes mandibularis on a couple of the plants. I also found a dead Polycestaaruensis.
Vicinity Tombstone, Arizona
Art saw Gyascutus caelatus here previously, so we came back and found them abundantly in sweet acacia (Acacia rigidula), which was in full bloom. They were extremely flighty and hard to catch, so we each got only four. I also collected one Stenaspis solitaria on the same and a Trachyderes mandibularis female in flight.
At another spot nearby, we stopped to look for Lampetus webbii, which Art had seen but not been able to collect when he was here a couple of weeks ago. We did not see any (but read on…), and I saw but did not collect a Trachyderes mandibularis and two Stenaspis solitaria. I also saw and photographed some giant mesquite bugs (Thasus neocalifornicus).
The day ended enjoying steaks, Malbec, and Jameson with two of the best hosts ever!
Day 3 – Box Canyon, Santa Rita Mountains, Arizona
Our first stop of the day was Box Canyon, a gorgeous, rugged canyon on the east side of the range. Mimosa dysocarpa was in bloom, off which I beat two Agrilus aeneocephalus, several Hippomelas planicauda, and one Stenaspis solitaria. Norm gave me an Acmaeodera cazieri that he’d collected on an unidentified yellow-flowered composite, and right afterwards I found some small, low-growing plants with purple flowers and sticky leaves (eventually ID’d as Allionia incarnata, or trailing four o’clock) to which Acmaeodera yuccavora and A. cazieri were flying in numbers. After that I crawled up top and beat the mesquites, getting one Chrysobothris sp., a mating pair of S. solitaria, and a couple of large clytrine leaf beetles.
Vicinity Duquesne, Arizona
We came here to look for Tetraopes skillmani (this is the type locality). We found the host plant (Sarcostemma sp.), but there were no beetles to be seen anywhere. Maybe another location nearby…
Patagonia Pass, Patagonia Mountains, Arizona We went up higher into the mountains to get into the oak woodland, where I hoped to find some of the harder-to-collect oak-associated Agrilus spp. Right away I beat one Agrilus restrictus off of Emory oak (Quercus emoryi), but no amount of beating produced anything more than a single Enoclerus sp.. I also beat the Arizona oak (Q. arizonica) and got only a single Macrosaigon sp. On Desmodium sp. I collected not only Pachyschelus secedens but a nice series of Agrilus arizonicus. For me it is the first time I’ve collected either A. restrictus and A. arizonicus, the former being quite uncommon as well, so all-in-all not a bad stop.
Sycamore Canyon, Santa Cruz Mountains, Arizona
We came here for night lighting, but while we still had light I did some sweeping in the low vegetation and collected a mixed series of Agrilus arizonicus (on Desmodium sp.) and Agrilus pulchellus – the latter a first for me, along with two small cerambyids that could be Anopliomorpha rinconia. Conditions were perfect (warm, humid, and no moon), and we had lots of lights (Art’s five LED units, Steve’s MV/UV combo setup, and my UV setup), but longhorned beetles were scarce – just one Prionus heroicus and one Lepturges sp. for me, and Steve got a few others including a nice Aegomorphus sp. I did also collect a few scarabs – Chrysina gloriosa and Strategus alous – because they’re just so irresistible!
Day 4 – Prologue One of the downsides (if you can call it that) of having great collecting is the need to take periodic “breaks” to process all the specimens and make my field containers available for even more specimens. Thanks to Steve and Norm for making their place available to Art and I so we can do this before heading out to our next set of localities.
Copper Canyon, Huachuca Mountains, Arizona
Copper Canyon is the classic spot for finding the charismatic Agrilus cavatus (see photo), but first we did some sweeping in the low vegetation near the parking area, where Norm got one Agrilus arizonicus and two Agrilus latifrons – and gave them to me! (Thanks, Norm!) I did some beating of the oaks, and after much work I ended up with a single Agrilaxia sp. and pogonocherine cerambycid on Emory oak (Quercus emoryi) and a couple of giant clytrines on the Arizona oak (Q. arizonicus). I then started sweeping the low-growing Acaciella angustissima – right away I got two A. cavatus. They were in the area past the cattle guard on the right where lots of dead stems were sticking up, and although I continued to sweep the plants more broadly in the area I never saw another one. Finally, Norm called me up to a small Mimosa dysocarpa near the car off which he collected three Agrilus elenorae – and gave them to me! (Thanks, Norm!) I gave the tree a tap and got one more, and in my last round of sweeping I came up with a Taphrocerus sp. (must be some sedges growing amongst the grasses).
Bear Canyon Crossing, Huachuca Mountains, Arizona
There was quite a bit of Mimosa dysocarpa in bloom along the roadsides on the west side of the Bear Canyon crossing, which I beat hoping to find some more Agrilus elenorae. I didn’t find any, but I did get several more Hippomelas planicauda, which is a nice consolation prize – and a great photo of the last one! Other than that I did a lot of sweeping and found only a single Acmaeodera cazieri.
Appleton-Whittell Research Ranch of the National Audubon Society, Elgin, Arizona
Cool temperatures and a blustery wind discouraged most insects from finding our blacklights. However, our blacklight did find some other interesting local residents. These two individuals could be the stripe-tailed scorpion, Paravaejovis (Hoffmannius) spinigerus, a common species in Arizona and southwestern New Mexico.
Day 5 – Miller Canyon Recreation Area, Huachuca Mountains, Arizona
There was a lot of Baccharis sarothroides growing in the lower canyon near the parking area, so I checked it all out hoping to find Tragidion annulatum. None were seen, and in fact there was very little insect life in general. I did pick up a couple of Acmaeodera solitaria by sweeping – not anything significant but the 15th species buprestid of the trip and found a dead Cotinis mutabilis, and Art got a nice series of Chalcolepidius click beetles on B. sarothroides and Prosopis glandulosa. Puzzling the lack of insect activity, given how green all the plants were and how fresh the growth looked. I guess we’ll have to look elsewhere.
Vicinity Naco, Arizona
We decided to try some desert thorn-scrub habitat so headed east towards Bisbee. Just north of Naco we saw some habitat where it had rained recently – everything was green with the sweet acacia (Acacia rigidula) and creosote (Larrea tridentata) in full bloom. Immediately out of the car I found a Dendrobias mandibularis on Baccharis sarothroides (and when I came back to it later I found a big, major male on it – see photos). On the sweet acacia we found a handful of Gyascutus caelatus (one of which I got a nice photo of), a mating pair of Sphaenothecus bivittatus, and a Cymatodera sp. Finally, out along the roadsides a riot of different yellow composites were in full bloom, including Heliomeris longifolia off which Art got a couple of Acmaeodera solitaria and I got two specimens of a large Acmaeodera sp. (blue-black with numerous small irregular yellow spots).
Vicinity Tombstone, Arizona
We decided to go back to the spot north of Tombstone where Art had earlier seen Lampetis webbii and give that species another shot. We looked at the Rhus sp. tree that he’d seen them on, and then we each followed the wash in opposite directions looking at the Rhus trees along them, which growing above the banks but never further away than about 25 feet. Along the way I collected several more Gyascutus caelatus on sweet acacia (Acacia rigida), which were more abundant this time than last and also easier to catch. After walking about 1/4-mile down the wash I saw something fly from a Rhus tree and land low on the bushes nearby. I quickly netted it, pulled it out, and was elated to see that it was, indeed, Lampetis webbii! I searched the Rhus in the area more carefully but didn’t find any more, then found some Rhus growing up along the road. At one point, I saw a large buprestid fly and land high in the top of another Rhus tree. I couldn’t tell for sure if it was L. webbii, but I extended my net as far as I could, positioned it beneath the beetle, and tapped the branch hoping it would fall in. Unfortunately, it flew away instead of dropping, so I can’t say for sure whether it was L. webbii or just a wayward G. caelatus. At any rate, L. webbii is yet another species that I have not collected before now and the 17th buprestid species of the trip.
Ramsey Canyon, Huachuca Mountains, Arizona
After returning from Tombstone, we visited Pat & Lisa Sullivan at their home at the end of Ramsey Canyon. Pat is a scarab collector who runs lights at his home nightly, and after a delicious dinner we spent the rest of the evening checking the lights. I was hoping to collect Prionus heroicus, and I got my wish. Also got Prionus californicus and several other non-cerambycid beetles such as Chrysina beyeri, C. gloriosa, Lucanus mazama, and Parabyrsopolis chihuahuae (the latter a first for me). I also placed a prionic acid lure (thanks Steve!) and got three more male P. heroicus. We also hunted around the rocks and roadsides hoping to find Amblycheila baroni but didn’t find any. Art did, however, find a female P. californicus and gave it to me (thanks!).
Day 6 – Vicinity Sonoita, Arizona
Unsuccessful attempt to collect Hippomelas martini, only recently described (Bellamy & Nelson, 1998) and part of the type series taken somewhere near this spot (“20 mi NE Patagonia, Hwy 82”) by “sweeping roadside vegetation”. At other locations it had been recorded on Calliandra sp., and I found patches of the plant here along and on top of the road cuts. This gives me confidence that I found the right spot, but I didn’t encounter this or any other beetles by sweeping the patches or visually inspecting them.
Box Canyon, Santa Rita Mountains, Arizona
We decided to come back to Box Canyon since we’d had such good luck last time. I started at the spot above the dry falls where I collected so many Acmaeodera cazieri and A. yuccavora on flowers of Allionia incarnata. This time it was hotter, drier, and windier, and the flowers were semi-closed. Still I found a few of each. I then started walking down the road towards the lower canyon crossing where I would meet up with Art. Things were really hopping on the Mimosa dysocarpa, with Hippomelas planicauda abundant (finally collected my fill) and several other Buprestidae also beaten from the plants: Agrilus aeneocepahlus, Acmaeodera scalaris, Acmaeodera cazieri, Chrysobothris sp., and a species of Spectralia! (seven species of Buprestidae at one location I think is the high for the trip.) I checked other plants and flowers along the way down but didn’t find much.
Lower Madera Canyon, Santa Rita Mountains, Arizona
Madera Canyon is perhaps the most famous insect collecting locality in Arizona – maybe in the country, and it is hard to make a visit to Arizona without stopping by here. We elected to work the lower canyon first in an area where Chrysobothris chalcophoroides has been taken on Arizona oaks (Quercus arizonicus). Hiking towards the oaks I found some Stenaspis solitaria in a Baccharis sarothroides and marveled at the variety of other insects active on the plants (see photos) – later I would also collect an elaphidiine cerambycid on the plant. Next I started working the oaks, beating every branch I could reach with my net handle. With one whack of the stick a single Paratyndaris sp. and a single Brachys sp. landed on my sheet – those would be the only buprestids I would collect off the oaks! Other than that I collected one Hippomelas planicauda on Mimosa dysocarpa for the record. While I was working the oaks up in the knoll, the weather started turning with blustery winds, and I could see the rain coming in the distance. By the time I got down from the knoll the rain had arrived, and I walked back to the car in a sunny downpour using my beating sheet as an umbrella!
Montosa Canyon, Santa Rita Mountains, Arizona
Just to try something different, we went to Montosa Canyon – the next canyon south of Madera Canyon – for tonight’s blacklighting. We set my sheet up just E of the crossing and Arts ground units back to the west along a gravel road on the south side of the crossing. Moths came in numbers, but the beetles were light – I collected only blister beetles (Epicauta sp.) and a Cymatodera sp. checkered beetle at the sheet, a series of tiger beetles and a female Strategus cessus at the second ground unit, and a male Strategus aloeus and two Stenelaphus alienus at the third ground unit.
Day 7 (last day) – Vicinity Continental, Arizona
There was a photo posted on BugGuide of Stenaspis verticalis taken last week, so we decided to give it a shot and see if we could get lucky and find it ourselves. We checked all the Baccharis sarothroides within ½-mile if the spot but didn’t find it. I did, however, collect four Euphoria leucographa, two Chalcolepidius smaragdula, two Aneflus spp., and singletons of Stenaspis solitaria and Dendrobias mandibularis. I also took a couple of Hippomelas planicauda on Mimosadysocarpa – just for the record!
Lower Madera Canyon, Santa Rita Mountains, Arizona
We returned to work the lower canyon area. I’d heard that the tiger beetle Cicindelidia obsoleta santaclarae has been taken in the area last week so was hoping to run into it. While Art worked the east side of the road I worked the west, initially following FR-781 into what looked like grassland areas where the tiger beetle might occur. I didn’t see any but took Acmaeodera scalaris on Heterotheca sp. flowers and Acmaeodera solitaria on Argemone mexicana flowers. There was also a fresh wind-thrown mesquite (Prosopis glandulosa) with a bunch of Chrysobothris octocola and one Chrysobothris rossi on it. Still the area looked abused from grazing and was uninteresting, so I looked for another area to explore.
Northwest of the parking lot I spotted another grassy area that was dotted with Baccharis sarothroides, so I decided to give that area a look. After clambering several times through barbed wire fence, I reached the area and began to give it a look. Still no tiger beetles, but every time I passed a B. sarothroides I inspected it closely. I’d looked at several plants when I came upon one with a Stenaspis solitaria sitting in the foliage, and when I looked down on one of the stems and saw a big male Tragidion sp. on the underside of the stem. After securing it, I looked closer at the plant and saw a pair of annulated antennae crawling up another stem – I knew right away it was a mating pair of Stenaspis verticalis! After carefully moving to the other side to confirm, I dared to take a few photos in situ (see below) and then secured the couple. Of course, this gave me newfound motivation to work the entire area to look for more. It was very hot by then, and I was already quite thirsty, but I summoned up all the stamina that I could and worked as many plants as I could, ending up with six Tragidion spp. and three Stenaspis verticalis. The latter was one of my top priority targets for this trips, and the only thing more satisfying than getting it is doing so on my last day on the field.
Montosa Canyon, Santa Rita Mountains, Arizona
We returned to Montosa Canyon and stopped at the Astronomy Vista partway up. It was hotter than bejeebuz! There was not an insect to be seen except giant cactus bugs and a single Euphoria leucographa that Art found on a sapping Baccharis sarothroides. Temp was 103°F even at this elevation!
We needed to escape the heat, and I wanted to see oaks for one more crack at Mastogenius, so we drove up to the 13-km marker and I collected on the way back down to below the 12-km marker. Conditions were much more agreeable (temps in the 80s), and near the top there was a Ceanothus sp. bush in bloom, off which I collected Rhopalophora meeskei and Stenosphenus sp. – both genera represented by individuals with black versus red pronotum. Then I started beating the (Mexican blue, I believe) oaks, and right away I got a Mastogenius sp.! Kinda small, so I’m thinking not M. robusta and, thus, probably M. puncticollis (another species new to my collection). I also beat a largish Agrilus sp. that I don’t recognize, a few clerids, two R. meeskei, one Stenosphenus sp., and a couple of leaf beetles. There was also another type of oak there – Arizona white, I believe, which I beat as well but only got one clerid.
We stopped by the Astronomy Vista again on our way back down the canyon, and I found a pair of Moneilema gigas on cholla (Opuntia imbricata).
It was a fantastic seven days in the field with Arthur, and it was a great pleasure to (in some cases, finally) meet Margarethe, Barbara, Steven, Norm, and Pat. I appreciate the warmth, generosity, and hospitality that all of them displayed to me and look forward to our next encounter, hopefully in the near future. Now, for some light reading during the plane ride home!