2022 Six-State Insect Collecting Trip iReport

Welcome to the 12th “Collecting Trip iReport” covering a 22-day insect collecting trip (my longest in more than 20 years!) encompassing six states from May 15 to June 6, 2022. The trip started out with two days of setting traps in southern Illinois and across southern Missouri, continued with nine days of collecting in western Texas, three days of collecting in southeastern New Mexico, and five days of collecting in southeastern Arizona, and ended with a day of collecting in extreme northwestern Oklahoma at the halfway point during the long drive back to St. Louis. Along the way, I teamed up with six different people during different parts of the trip—Jason Hansen, Joshua Basham, and Tyler Hedlund in Texas and New Mexico and Norm Woodley, Steve Lingafelter, and Paul Kaufman in Arizona.

This report assembles the field notes generated during the trip, which are presented in “semi-rough” form—i.e., they have been lightly polished but not further modified based on subsequent examination of collected specimens. As with all previous “iReports” in this series, this report is illustrated exclusively with iPhone photographs (thus the term “iReport”). Previous iReports in this series include:
2013 Oklahoma
2013 Great Basin
2014 Great Plains
2015 Texas
2018 New Mexico/Texas
2018 Arizona
2019 Arkansas/Oklahoma
2019 Arizona/California
2021 West Texas
2021 Texas/New Mexico/Arizona
2022 Oklahoma

Packed and ready to go!

Day 1 – Setting out jug traps

Salt Lick Point Land & Water Preserve
Monroe Co., Illinois

I’m on my way to west Texas for a couple of weeks and southern Arizona for another week afterwards, but it’ll take a few days to get there while I hang insect traps here in southern Illinois and at a bunch of localities in southern Missouri and northern Arkansas. The traps are based on an idea from fellow cerambycid specialist Dan Heffern, who calls them “jug traps”. The traps utilize 8-oz bottles of 200 proof ethanol suspended inside a 1-G milk/water jug. A hole is drilled in the lid of the 8-oz bottle and a wick inserted to moderate release of the ethanol, which attracts the beetles. 500 mL of a 50:50 mixture of polypropylene glycol and water is placed in the bottom of the jug to act as a killing agent and preservative for beetles that are attracted to the trap and fall into the it. Early testing by Dan suggests the ethanol bait can last up to 4–6 weeks and the beetles that fall into the traps don’t decompose within that time period, enabling them to be placed at much more remote locations than the fermenting bait traps that I have been utilizing for many years now in the glades of Jefferson Co. just south of St. Louis but which need to be checked weekly. For my part, I am placing two traps at each location—one with pure 200 proof ethanol and another with a 50:50 mixture of ethanol and red wine (the latter is cheaper and works well as a beetle attractant on its own). If the mixture works as well as pure ethanol (remains to be seen), it would be a way to reduce cost.

Assembled jug trap with 50:50 red wine:ethanol in inner bait bottle and 500 mL 50:50 polypropylene glycol:water in the reservoir of jug.

I chose this location based on a visit last fall with the WGNSS Botany Group, during which we found Sideroxylon lanuginosum (gum bumelia)—host of the strikingly spectacular Plinthocoelium suaveolens (bumelia borer) and one tree in particular that showed evidence of active larval infestation by the telltale pile of frass at the base of the living tree. The trees are growing in dry hilltop forest adjacent to a renovated hilltop prairie remnant, and the beetle has not been formally reported from Illinois. I’ve had good luck trapping this beetle in the glades south of St. Louis, so I am hopeful these traps will also be effective and that I can document the occurrence of bumelia borer in Illinois.

Jug trap bait bottle hanging mechanism.

It was a tough hike—mostly uphill and I was trying to get in and out quickly. I had a bit of trouble locating the infested tree that we’d seen last fall (even with the location GPS recorded), but eventually I found it. At first, I had trouble throwing the carabiner and rope over a high branch—my slingshot idea with the rope tied to the carabiner did not work, so I ended up just throwing it and succeeded only after many attempts to develop my “technique.” Then, horrors… somehow the bottle carrying the mixture of red wine and ethanol broke and spilled much of the contents into my backpack. Fortunately, there was just enough remaining in the bottle to mostly fill the 8-oz bait bottle. From that point, the rest of the trap went together as planned, and I hoisted it high above eye level before assembling the ethanol-only trap and placing it about 100 feet from the first. The longer-than-I-remembered hike and problems with the first trap already had me close to an hour off schedule, so I hoofed it back to the car as quickly as I could, stopping only briefly to pick up a tiny Glaphyrocanthon viridis (one of our tiniest dung beetles) that I saw land on the trail. When I got back to the car and checked my recorded track, I noticed that I had made it about three-fourths of the way around the trail and could have saved time had I simply completed the circuit rather than doubling back! Ugh—an inauspicious start to a long trip!

Papilio glaucus (eastern tiger swallowtail).

St. Joe State Park
St. Francois Co., Missouri

This is another location that I visited recently with the WGNSS Botany Group and noted the occurrence of Sideroxylon lanuginosum (gum bumelia) in the dry post oak woodland bordering the bicycle trail south of the Harris Branch Trailhead. I hung a wine:ethanol trap not far from the parking lot on the west side of the trail and an ethanol-only trap about 300 feet further south on the east side of the trail.

Acmaeodera ornata (ornate yellow-marked buprestid) on flower of Coreopsis lanceolata (lance-leaved coreopsis).

When I returned to the car, somebody had used a sparkly wrist strap to tie a bicycle key chain to my door handle. I’m not sure if it was a gift from a fellow cyclist who recognized my “Share the Road” license plate or simply a random act of kindness—either way, I think I’ll hang the bicycle from my rear view mirror!

Gifts for me!

Hughes Mountain Natural Area
Iron Co., Missouri

This area features dry post oak woodlands surrounding xeric igneous prairie (glades). I’m not aware of the presence of Sideroxylon lanuginosum (gum bumelia) in these (or any other) igneous glades, but the post oak woodlands could support many other longhorned beetles that are attracted to ethanol and fermenting bait traps (especially species of Purpuricenus). I placed a wine:ethanol trap at the north end of the first glades and an ethanol-only trap just past the south end about 1/10 of a mile away.

Acmaeodera tubulus (family Buprestidae) on flower of Rubus sp. (wild blackberry) in dry-mesic upland deciduous forest.

Russell Mountain Trailhead
Iron Co., Missouri

This is another area with igneous glades surrounded by dry post oak woodland. I hung a wine:ethanol trap at the north end of the first glade and an ethanol-only trap at the south end about 400 feet away. While I was hanging the first trap, I saw a ground beetle (family Carabidae) crawling over the moss-covered rocks under the tree. I believe it is a species in the genus Dicaelus (notched-mouthed ground beetles).

Dicaelus sp. (notched-mouth ground beetle) in xeric rhyolite prairie.
Silene virginica (fire pink) in igneous post oak woodland.
Tradescantia longipes (wild crocus) in dry rhyolite forest.
Phemeranthus parviflorus (prairie fameflower) in xeric rhyolite prairie.

Peck Ranch Conservation Area
Stegall Mountain Natural Area
Carter Co., Missouri

This area has some of the most extensive igneous glades in southeastern Missouri, and it’s remote location makes it an attractive spot for placing traps. Unfortunately, the entire natural area —glades and surrounding woodlands—has been subjected to a recent prescribed burn. I’ve never had much luck collecting in recently-burned habitats, so I had little hope that placing traps here would be worth the effort. Nevertheless, I was there and figured if nothing else it would be a chance to gather some objective data comparing a recently-burned area with similar non-burned areas. I hung a wine:ethanol trap at the north end of the glade next to the fire tower and an ethanol-only trap in the dry post oak woodland about 400 feet to the southwest.

Recently-burned xeric igneous prairie (glade).
Fire tower.
I cannot claim those stairs.

Peck Ranch Conservation Area
Cater Co., Missouri

I had originally planned to hang these traps at Mule Hollow Glade Natural Area—also in Peck Ranch Conservation Area but distinct from Stegall Mountain by the fact that the glade substrate is limestone rather than rhyolite. This results in a calcareous versus acidic environment and a completely different (and richer) glade flora—including potentially Sideroxylon lanuginosum (gum bumelia) and its beetle associate Plinthocoelium suaveolens. Sadly, after driving from Stegall Mountain through this enormous conservation area for about half an hour, I encountered a gate that prevented access to the glades. It was late in the day, and rather than drop the location, I decided to just hang traps where I ended up—a dry-mesic upland deciduous forest apparently with a chert substrate. I hung a wine:ethanol trap near the car on the east side of the road leading south behind the gate and an ethanol-only trap also on the east side of the road about 230 feet further south.

Penstemon pallidus (pale beardtongue).

Day 2 – Setting out more jug traps!

Mark Twain National Forest
Bald Hill Glade Natural Area
Ripley Co., Missouri
I’d hoped to make it here yesterday, but the day just ran out and I ended up spending the night in Doniphan. A great little coffee shop in town put me in the right frame of mind this morning to make the trek into this—one of the most beautifully remote high-quality glades in all of Missouri. The Forest Service roads leading to the glade become increasingly rough the closer one gets, and the final 1-mile spur required a bit of log/branch removal to pass through and even bushwhacking around and under fallen trees before an impassable blockage about halfway down. The last half-mile has been abandoned for at least 10 years, and walking it by foot required a keen sense of reading the forest to discern the barely visible remnant path.

Ethanol-baited jug trap in Sideroxylon lanuginosum (gum bumelia) in xeric dolomite prairie remnant.

At last, I made it to the glade proper—a gorgeous tract of remnant xeric dolomite prairie (glade) with dry post oak woodlands interspersed within and surrounding the glades. I hung a wine:ethanol trap in a post oak on the east side of the main glade (past the first small glade) and an ethanol-only trap in a gum bumelia tree at the opposite end of the glade about 1/10 of a mile to the southwest. Along the way, I photographed and collected Nemognatha nemorensis (a blister beetle—family Meloidae) on a flower of Coreopsis lanceolata (lanceleaf coreopsis).

Nemognatha nemorensis (family Meloidae) on flower of Coreopsis lanceolata (lanceleaf coreopsis) in xeric dolomite prairie remnant.

As with the first stop yesterday, I underestimated the time needed to hike to the glade, hang the traps, and get back to the car, so by that time I was already off schedule. Nevertheless, considering the quality of the area, I decided to hang one of the two Lindgren funnel traps (baited with ethanol) near the car.

Shortly after turning off the spur onto the Forest Service road, I passed by a branch that looked suspiciously “pruned” (i.e., cut from the inside by a cerambycid larva). I stopped and walked back to the branch, which turned out to be Carya alba (mockernut hickory), and the size of the branch at the cut (~1” diameter) suggested it could be the work of Purpuricenus axillaris, a beautiful orange and black species that is very uncommonly encountered. I pruned off the excess twigs and collected the branch for rearing.


Caney Mountain Conservation Area
Long Bald Glade Natural Area
Ozark Co., Missouri

After finishing up at Bald Hill Glade, I blasted two hours west to Caney Mountain Conservation Area at the eastern edge of the White River Hills region in southwestern Missouri. The White River Hills is perhaps my favorite area in Missouri—I have collected insects at many spots here over the years, a number of which occur in Missouri only in this part of the state. Caney Mountain is only one of the sites I’ve selected for placing traps, but like the previous site it contains some of the highest quality and most beautifully remote xeric dolomite prairie remnants (glades) in the state—especially on the far west side of the area in and around Long Bald Glade Natural Area.

Acmaeodera neglecta (family Buprestidae) on flower of Coreopsis lanceolata (lanceleaf coreopsis) in xeric dolomite prairie remnant.

As before, I hung a wine:ethanol trap in a Sideroxylon lanuginosum (gum bumelia) tree on the west side of the first glade and an ethanol-only trap in another gum bumelia tree on the north side of the main glade about 1/8 of a mile west of the first trap. I also hung a second Lindgren funnel trap here—when I arrived at the site, I’d noticed a large area of post oak woodland on the other side of the road had recently been thinned (via chainsaw). With all the dead wood laying around (in a cool natural community), it almost screamed for a trap, so I baited it with ethanol and hung it right smack in the middle of the renovated area.

Ethanol-baited Lindgren funnel trap in dry post oak woodland.

By this time, the day was starting to get away and I still had four locations that I wanted to hang traps. With six hours of daylight left, it would be a stretch to get to all four, so I avoided the temptation to spend any more time poking around in this fantastic site and headed to the next location further west.

Mark Twain National Forest
“Blackjack Knob

Taney Co., Missouri
I’ve been to this knob several times and collected good numbers of Missouri’s disjunct population of the spectacular Cicindelidia obsoleta vulturina (prairie tiger beetle) as well as discovered the larva of Plinthocoelium suaveolens (bumelia borer) in the root of a living Sideroxylon lanuginosum (gum bumelia) tree. There is lots of gum bumelia at this spot, and since it is right along the highway and I could hang traps here quickly, I added the spot to my list of locations. I hung a wine:ethanol trap in a gum bumelia tree on the south side of the knob and an ethanol-only trap in another gum bumelia tree on the north side of the knob about 300 feet north of the first trap.

Sisyrinchium campestre (prairie blue-eyed grass) in xeric dolomite prairie remnant.

Mark Twain National Forest
Hercules Glades Wilderness
Taney Co., Missouri

Hercules Glades Wilderness contains some of the largest intact remnants of xeric dolomite prairie in the entire White River Hills region. Unlike those of other areas managed by state and federal conservation agencies, this designated wilderness has a “no management” mandate. As a result, there has been no effort to remove woody vegetation, either by chainsaw or by prescribed burning. While plenty of intact glade habitat remains, the margins and surrounding dry post oak woodlands are heavily colonized by Juniperus virginiana (eastern red-cedar)—a native tree that was historically restricted to bluffs and ledges but has since adapted to encroaching in glades and prairies as a result of fire suppression over the past one and a half centuries. I hung a wine:ethanol trap in a red-cedar near a Sideroxylon lanuginosum (gum bumelia) tree at the north end of the first glade and an ethanol-only trap in another red-cedar near gum bumelia about 450 feet to the southeast.

Oenothera macrocarpa (bigfruit evening primrose, Ozark sundrop, Missouri evening primrose) in xeric dolomite prairie remnant.

Along the trail in the dry oak-juniper woodland before reaching the glade, I found a Geotrupes splendidus (splendid earth-boring beetle) on its back waving its legs in the air. I flipped it over, took a photograph, and popped it in a vial.

Geotrupes splendidus (splendid earth-boring beetle) on trail in dry oak-juniper woodland.

Mincy Conservation Area
Taney Co., Missouri
Mincy Conservation Area is another area in Taney Co. with high quality remnant xeric dolomite prairie (glades). I have been here many times, and I couldn’t imagine placing traps in the White River Hills and not including this place. I hung a wine:ethanol trap in a Sideroxylon lanuginosum (gum bumelia) tree at the north end of the glade across the road from the campground area and an ethanol-only trap in another gum bumelia at the southwest end about 1/8 mile from the first trap.

A marvelously cryptic Dolomedes albineus (whitebanded fishing spider) on the trunk of Sideroxylon lanuginosum (gum bumelia) in xeric dolomite prairie.
Ethanol-baited jug trap in Sideroxylon lanuginosum (gum bumelia) in xeric dolomite prairie remnant during late evening.

Roaring River State Park
Chute Ridge Glade
Barry Co., Missouri

I got here right at sunset, so I knew I would have to work quickly to get two traps hung before I completely ran out of daylight. This high-quality xeric dolomite prairie remnant (glade) has undergone extensive renovation over the past 25 years since I first began coming here, and it’s character is now much improved compared to those early days. I hung a red wine:ethanol trap in a Quercus stellata (post oak) tree near some Sideroxylon lanuginosum (gum bumelia) in the treeline about halfway up the slope at mid-glade. It was at that time that I noticed the bait bottle was missing from the second trap, so I had to hustle back to the car in the waiting light to retrieve another trap. It was too dark by then to wander back up into the main glade, so I walked the 2-track near the road to the north end of the glade and then east up the slope until I encountered an area where gum bumelia was growing, hanging an ethanol-only trap in a nearby hickory tree about 1/6 mile north of the first trap.

Psellidotus cf. hieroglyphicus (family Stratiomyidae) on flower of Coreopsis lanceolata (lanceleaf coreopsis) in xeric dolomite prairie remnant. ID by Martin Hauser.

As I walked the 2-track, I heard the call of nighthawks flying overhead—a familiar sound during the day and early evening when in the glades. Once I started hanging the trap, the oncoming cloak of darkness was announced by the whip-poor-wills (a relative of the nighthawk, both species belonging to the “goatsucker” family). If that was not enough, a Chuck-wills-widow—yet another nighthawk relative—joined the chorus! It was a magical moment of pure natural history to celebrate the completion of my trap placing effort, after which I pointed the car towards west Texas (or at least Fort Smith, Arkansas) for the night before the long drive tomorrow.

Xeric dolomite prairie remnant at sunset!

Day 3 – Travel to West Texas

Fire southwest of Abilene, Texas.

Gold Mine Canyon
Val Verde Co., Texas

Today was a long, lonely, 11-hour drive from Ft. Smith, Arkansas to Dave Barker’s cabin above Gold Mine Canyon. I first came here last year a bit earlier in May with Rich Thoma to meet up with Dan Heffern, Brian Raber, and Ed Riley. Dave has been kind enough to make his cabin available to naturalists interested in studying the flora and fauna of the area around his cabin, and after getting a taste of the area last year I wanted to come back again and see it a tad bit later in the season. This time I met up with Jason Hansen, Joshua Basham, and Tyler Hedlund. The area got some rain in late April but has been dry since—and looked it. Nevertheless, insect activity at the lights (mercury-vapor and ultraviolet) was fairly high, although mostly moths and blister beetles. I did pick up one elaphidiine, a series of Hybosorus illigeri, and a few photogenic robber flies.

Efferia sp. male (family Asilidae) at mercury-vapor light in juniper chaparral.
Efferia sp. female (family Asilidae) at mercury-vapor light in juniper chaparral.
Brachynemurus sackeni (Sacken’s antlion) at mercury-vapor light in juniper chaparral.

Eventually the wind picked up to the point where I was worried about my light setup being blown down, so I took down the setup and searched (unsuccessfully) for Moneilema cactus beetles on the nearby Opuntia sp. (pricklypear cactus).


Day 4 – Devils River Area

Gold Mine Canyon
Val Verde Co., Texas

We stayed in the vicinity of the cabin to see what we could find. I wasn’t optimistic because of how dry everything looked but headed up the 2-track leading east of the cabin. Even though I was here earlier in May last year, things didn’t seem as far along—the Diospyros texanus (Texas persimmon) trees were just beginning to leaf out, and I didn’t see any of the plants in bloom that I’d seen last year such as Coreopsis or Opuntia. As I walked the 2-track, I noted a persimmon with emergence holes in the trunk that matched the size and shape expected for Spectralia robusta. The holes appeared fresh, suggesting there could still be some beetles inside, so I flagged the branch for later collection and rearing. Having flagged the tree, I decided I should beat the branches just in case, and a fresh adult landed on the sheet—sweet! I’ve collected this species before, but it was many years ago and I’d forgotten how pulverescent the adults are and how (unfortunately) the pulverescence rubs off when touched or placed in the kill vial, making it almost impossible to preserve. I focused on beating persimmons for the next hour, ending up with seven specimens, all of which were collected in the immediate vicinity of the tree that I’d originally flagged.

Hemiargus ceraunus (Ceraunus blue).

I had beaten a few other trees as well but wasn’t seeing anything, and by now temperatures were starting to soar, so I went back to the cabin to rehydrate and trade my aerial net for my sweep net so I could do some general sweeping. I worked my way back to the farthest point I’d gone before and shortly afterwards encountered Echinocereus enneacanthus intermedius (strawberry cactus) in bloom. To my surprise, I saw several Acmaeodera adults on the blossom, so I collected them with my aspirator and immediately thought of the larger clump of strawberry cactus blooms I’d checked earlier and not seen anything and then passed by this time. Again, there were quite a few Acmaeodera adults on the blossoms. As I collected the adults, others continued to fly in to the flowers, so I roamed back and forth between the two clumps collecting the adults until no more were seen. There were at least four species—the commonly encountered A. quadravittatoides and A. neoneglecta, the much rarer A. starrae (which I collected for the first time last year in Comstock), and a fourth species that I didn’t immediately recognize. In shuffling through the possibilities in my mind, A. riograndei came up based on my recollection of Nelson’s illustration of the species in the original description. I walked further east down the 2-track and encountered another cluster of plants in bloom, allowing me to increase my series of all four species. By then, temperatures were approaching 100°F and I was also hungry, so I returned to the cabin to rehydrate, eat, and rest to avoid pushing myself too hard. When Jason returned to the cabin later, he had also found the same four species on cactus flowers, and we both agreed the mystery species was A. riograndei—the first time either of us had seen this very rarely collected species.

Echinocereus enneacanthus intermedius (strawberry cactus) in juniper chaparral.
Echinocereus enneacanthus intermedius (strawberry cactus).
Acmaeodera starrae on flower of Echinocereus enneacantha intermedius (strawberry cactus).
Acmaeodera riograndei on flower of Echinocereus enneacantha intermedius (strawberry cactus).

Once I felt energetic enough, I braved one more trip even further down the 2-track to where it crosses the canyon, the latter in which Joshua had seen strawberry cactus flowers in bloom and collected all four species himself. I hoped to find a few more A. riograndei, as I had only a handful of specimens of that species. Joshua came along, and together were located and worked as many plants in bloom as we could find. It was hard work—the plants were very sparsely distributed and mostly on the steep-sloped portions of the canyon walls or up on top where footing was precarious. Most plants had the two common species, and I managed to collect several more A. starrae as well, but I never saw another A. riograndei until after we’d been out there for a couple of hours and I was almost ready to collapse from the heat (temps were by then ~105°F!). On the same flower that I finally found A. riograndei, I also found a fifth species—A. gillespiensis, a west Texas specialty. That was one of the last plants in bloom that I found before working my way back down into the canyon and heading back to the canyon. I really thought I was going to collapse from the heat before I got there, and I needed the rest of the afternoon to rehydrate and all evening to recover. It was too windy to blacklight, which was probably fortunate because I really needed to take it easy during the evening and let myself recover.

Bee fly (subfamily Bombyliinae) in juniper chaparral.
Crotaphytus collaris (eastern collared lizard) in juniper chaparral.

Day 5 – Devils River area (cont.)

Devils River near Dry Devils River
Val Verde Co., Texas
We carpooled to a spot along the west side of Devils River that required fording the river and then traversing some of the roughest, rockiest roads I’ve ever traveled (my new Bronco Sport Badlands, chosen for just such roads, handled everything perfectly). There is a stand of mature Carya illinoiensis (pecan) along the river that Joshua and I headed straight for, suspecting they might harbor Anthaxia caryae. We spent a fair bit of time beating the lower reachable branches, and I spent even more time afterwards using the extensible net to sweep the upper branches. A single adult—on one of the first few branches that I beat—was all we got for our efforts. I also collected a little chlamisine chrysomelid by sweeping Salvia sp., but otherwise I saw little insect activity. Joshua had been beating the nearby oaks while I was working the pecans, and when I passed by he said he hadn’t gotten anything off the oaks either. I walked back up to the bluffs overlooking the river to see if I could find cacti in bloom but found no Opuntia (pricklypear cactus) and only the infrequent Echinocereus enneacantha intermedius (strawberry cactus) in bloom. Despite the generally poor condition of most of the flowers, I still managed to collect the same four Acmaeodera that we collected yesterday—including several A. starrae and two A. riograndei. While I looked for cacti, I also beat any Diospyros texana (Texas persimmon) that I encountered hoping to find more Spectralia robusta or the small Agrilus sp. that Tyler collected yesterday off the same at Gold Mine Canyon (I’m thinking it must be A. lautuellus), but all I collected was a single weevil and a single tenebrionid. By this time it was getting hotter than blazes, and we all returned to the car, drank some fluids and ate a bit, and decided the best way to spend the next two hours—the hottest part of the day—was by sitting in the river. We had no swim trunks, but underwear served the purpose just as well!

Zeltnera calycosa (Arizona or Buckley’s centaury) on limestone bluffs.
Zeltnera calycosa (Arizona or Buckley’s centaury).
Zeltnera calycosa (Arizona or Buckley’s centaury).

Gold Mine Canyon
Val Verde Co., Texas
After cooling off in the river and returning to the cabin, Jason and Joshua wanted to go check out the patches of Echinocereus enneacantha intermedius (strawberry cactus) from which we’d collected so many Acmaeodera to look for more A. riograndei, and Tyler and I decided to hike down to the canyon where he had collected a few Agrilus sp. on Diospyros texana (Texas persimmon). Along the way we beat some of the persimmon trees where Tyler had collected a few Spectralia robusta but found only a single epitragine tenebrionid. We also encountered a single strawberry cactus patch with a couple of closed blooms, and as I approached to see if any Acmaeodera were on the flowers I saw one approaching the flowers in flight. I instinctively swiped the net and caught it, and when I pulled it from the net I saw it was another A. riograndei. I must have around ten specimens of this species now—a nice series of a rare species for my collection. We had to pick a rough and precarious path to reach the canyon bottom, but once we did we started beating the persimmons on which he’d collected the Agrilus sp. (prob. A. lautuellus). Almost immediately he found another one and gave it to me (what a guy!), and we continued working the trees down the canyon. We did not see any more for awhile, but then suddenly I hit a hit spot where I collected one or a few off of successive plants, ending up with a total of nine specimens. Tyler never did find another one after that first specimen (but he’d collected a small series yesterday so he was fine). Once we worked all the persimmon that we could find, we worked our way up the canyon walls on the south side to look for more strawberry cactus flowers with Acmaeodera. We found a few plants here and there, but in all cases the flowers were closed and no Acmaeodera were seen—I suspect the flowers close and the Acmaeodera stop flying as a matter of routine at this time of day (now early evening). As we worked our way east above the canyon to a point where we could cross back over to the cabin, we beat persimmon, but I collected only a single anthribid. By this time we were hungry and thirsty and hoofed it back to the canyon to eat and get ready for blacklighting.

Late afternoon sun over Gold Mine Canyon.

Winds were not as bad as they were last night (and I was feeling much better than I did last night, having taken better care to keep myself fueled and hydrated), so we were anxious to put up the lights and see if we could collect cerambycids. I put up my mercury-vapor/ultraviolet light combination just east of the cabin, while Jason set up his mercury-vapor light on the west side. I picked up a few miscellaneous insects from each over the next hour after it got dark, but I hadn’t yet seen a single cerambycid and started exploring the surrounding area with Tyler. Not far from my lights I found an elaphidiine cerambycid on the 2-track—most likely it had been pulled to the area by the mercury-vapor lamp but landed in the area rather than coming all the way to the light. I tried (and failed) to photograph a mutillid female, so I collected it instead, and we found a cool Stenomorpha sp. (family Tenebrionidae) and some very impressive arachnids (two Hogna carolinensis wolf spiders—one juvenile and one adult female, a Centruroides vittatus scorpion, and a sun spider—Eremobates sp.)—all of which I did manage to photograph! Coming back to the lights, my generator had run out of gas and the lights died, so we brought the ultraviolet lights over to Jason’s setup and turned off his mercury-vapor lamp to encourage cerambycids that had been attracted to the area to come on in to the lights. A couple of Lagocheirus sp. turned up (Jason and Tyler got them), and I got a couple of Aneflomorpha sp. and one Elaphidionopsis fasciatipennis—a species I have not collected commonly. This would be the last beetle I collected on the evening, bringing to a close a second hot but relatively successful day of collecting. Tomorrow we will leave the cabin and start working our way west towards the Davis Mountains.

Sunset over Gold Mine Canyon.
Stenomorpha sp. (family Tenebrionidae).
Centruroides vittatus (striped bark scorpion).
Hogna carolinensis (Carolina wolf spider) juvenile.
Hogna carolinensis (Carolina wolf spider) adult female.
Hogna carolinensis (Carolina wolf spider) adult female.
Eremobates sp. (order Solifugae).
Eremobates sp. (order Solifugae).

Day 6 – To Comstock area

22 mi N Del Rio – Jct Hwys 277 & 377
Val Verde Co., Texas
We left Dave’s cabin in the morning, and on our way out to the Comstock area we stopped at this intersection where last May I collected Agrilus obtusus on Senna roemeriana (two-leaved senna). I got four more this time as well in the small patch of plants just inside the fence, but when I went up the north side where there used to be many more plants, I was disappointed to find that the highway department had dumped multiple loads of gravel over the area. Still, there were other plants in flower closer to the roadside, and when I went to look at them I noticed right away Batyle suturalis, Acmaeodera ornatoides, A. neoneglecta, and A. mixta on flowers of Thelosperma simplicifolium (slender greenthread). Over the next half hour or so, I collected more of the same plus a few additional species (including A. paradisjuncta) in smaller numbers. Another Batyle was taken off the flowers of Ratibida columnifera, but then I noticed Acmaeodera starrae on the small, low-growing flowers of Pinaropappus roseus (white rock-lettuce) and focused on those flowers, ending up with a fair series collected almost exclusively on the flowers of that plant save for single exceptions on the flowers of Sida abutifolia (spreading sida) and Stenaria nigricans (diamond-flowers). I found it interesting that no A. starrae were taken on the flowers of the much more abundant Thelosperma. To the contrary, I did find a few individuals of A. mixta and A. neoneglecta on the flowers of Pinaropappus.

Agrilus obtusus (family Buprestidae) mating pair on foliage of Senna roemeriana (twoleaf senna).
Thelesperma simplicifolium (slender greenthread).
Pinaropappus roseus (white rock-lettuce).
Sida abutifolia (spreading sida).
Stenaria nigricans (diamond-flowers).

11.5 mi SE Comstock on Hwy 90
Val Verde Co., Texas

Jason has collected Agrilus esperanzae and Acmaeodera opuntiae at this spot during previous visits—two species I’ve not yet collected myself, so we stopped here to try our luck. Sweeping along the mesquite/acacia fence line produced only one Agrilus—not A. esperanzae (probably A. addendus)—and a smattering of other beetles; however, we were successful in our quest for A. opuntia, which we found on the flowers of Tiquilia canescens (shrubby tiquilia). They were not common and required a lot of effort to see and capture—sweeping was ineffective because of the very low-growing nature of the plants, and since the beetles are among the smallest Acmaeodera there are I had to crouch over each flowering plant and inspect carefully (under overwhelming heat). Fortunately, I was able to successfully aspirate them once I did see them, and I ended up with a small handful along with similar numbers of A. neoneglecta and A. starrae. The adults of A. opuntiae are unlike those I have in my collection collected by Ed Riley further south—the vittae are more broken, giving them a linearly-spotted rather than vittate appearance. One cool find was the blister beetle Pleuropasta reticulata—one of the two I captured going to Tyler since he had actually targeted that species for the trip.

Tiquilia canescens (woody crinklemat, shrubby tiquilia).

After a hydration break, I went to the other side of the highway where a nice stand of Thelosperma filifolium (stiff greenthread) was hosting Acmaeodera and off which I collected a few A. miliaris and A. princeps amongst the more common A. mixta. There were also a few flowering Tiquilia plants on that side, and while I did collect a few more A. starrae and A. neoneglecta I did not see A. opuntiae. By then the heat had gotten to me and I worked my way back to the car—save for the efforts given to photograph a couple of robber flies and some neonate coreids.

Acmaeodera mixta (family Buprestidae) mating pair on flower of Thelosperma filiformis (stiff greenthread).
Efferia sp. (family Asilidae) female.
Ospriocerus aeacus (family Asilidae).
Newly hatched leaffooted bugs (family Coreidae) aggregating near egg shells.

Devils River at Bakers Crossing
Val Verde Co., Texas
After getting a hotel in town (I’m looking forward to a hot shower instead of a cold river, for once), we headed north on Hwy 163, along which Jason has had good collecting in the past. The first stop just north of Comstock was not productive despite the verdant plant growth, so we continued north to Bakers Crossing at the Devils River. It was now early evening, so the heat had broken, and immediately we started finding beetles by beating the various trees. I collected one Chrysobothris rossi, one Euderces reichei, and a weevil on Sapindus saponaria (soapberry), but it wasn’t until I started beating Prosopis glandulosa (mesquite) that the beetles started “raining” onto the sheet! Chrysobothris rossi was abundant—sometimes two or three falling into the sheet at once and scattering immediately despite the cooler temps. It took me a bit of time to perfect my technique to avoid losing as many as I was getting. Smaller numbers were also collected from dead branches of Vachellia constricta (white-thorn acacia), and just as dusk was falling I found numerous Stenosphenus dolosus bedded down on the thorny branches of Zizyphus obtusifolium. I also collected a couple more C. rossi on dead branches of Celtis sp. before calling it quits.

Danaus gilippus (queen butterfly) mating pair.
Stenosphenus dolosus (family Cerambycidae) mating pair on Zizyphus obtusifolia (lotebush).

Due to the lateness of the hour, and since it seemed to be such a good spot, we decided to stay out and put up the lights. Jason setup his mercury-vapor (MV) light closer to the river, while I put my MV/ultraviolet lights in the area where I had been beating. I collected a fair number of cerambycids (not a lot) and a variety of other beetles from the two lights while we waited for our coals to heat up so we could cook some dinner (a fiasco to recount on future trips).

Hogna carolinensis (Carolina wolf spider) adult female.
Hogna carolinensis (Carolina wolf spider) adult female.
Hogna carolinensis (Carolina wolf spider) adult female.
Corydalus luteus (family Corydalidae).

Day 7 – Pecos River area

The motel owner tooled around in his MAGA-mobile. Photo by Jason Hansen.
MAGA Power! Note Mike Pence’s name covered with “Impeach The Democrats” scrawled on duct tape. Photo by Jason Hansen.

Pecos River at Hwy 90
Val Verde Co., Texas
We awoke to much cooler temperatures (hallelujah!) thanks to a cold front that moved through the area last night, though without the 40% forecasted chance of rain (also good). Our plan today was to work the area around the Pecos River and then look for oaks in nearby Seminole Canyon State Park.

Pecos River at Hwy 90 bridge.
Pecos River at Hwy 90 bridge.

On top near where we pulled the vehicles off the road, I beat a Chrysobothris rossi off dead Vachellia rigidula (blackbrush acacia)—many of which had been killed in the great Texas freeze two winters ago. About that time, Jason and Tyler called me over to look at a buprestid larvae they had beaten from a dead branch of the same—it was not chrysobothrine or agriline, and based on the size of the many emergence holes observed in the branches of this tree we suspected either Xenorhipis osborni (which I have reared from this plant at this location in the past) or a small species of Acmaeodera. Jason collected the larva, and we both collected branches to bring back for rearing. Nearby I found another dead tree of the same but noted a complete absence of emergence holes, yet when I broke apart one of the branches I found a buprestid larvae just like the previous (as well as a C. rossi adult on the branch) and collected some branches for rearing from that plant as well.

Hemiargus ceraunus (Ceraunus blue).

I noted a few flowers of Ruellia parryi (Parry’s petunia)—in my experience other species of this genus are good attractors of Acmaeodera—but did not see any adults. The day was still cool and cloudy, so I hoped I might see some later after it warmed up and the sun came out.

Ruellia parryi (Parry’s ruellia).

I crossed over the highway about halfway down since the others had gone further down on the side I was on, wanting to avoid working trees they’d already worked, and found a fence crossover stand at the bottom under the bridge. I’d never been down that far before or noticed the crossover, which gave me access to the old road going all the way down to the river. I started beating Prosopis glandulosa (mesquite) along the way, eventually accumulating around 8–10 C. rossi and a couple of Agrilus sp. along with a few other insects. There was also dead Celtis pallida (spiny hackberry) near the bridge, but I only beat a single miscellaneous beetle off one of the plants.

Atlides halesus (great purple hairstreak).

Near the bridge I noticed a small purple flower of Justicia pilosella (Gregg’s tube tongue) that looked like it had been eaten by Acmaeodera. I did not see any adults on it, but as I started to walk away movement caught my eye—movement like that of an adult Acmaeodera dropping from the flower. I used my aspirator to pick through the soil underneath the flower and eventually found the little guy laying there playing dead—presumably A. neoneglecta, which I then aspirated into a vial. There were just a few other plants in flower around that one, none of which showed evidence of feeding or were hosting a beetle.

Justicia pilosella (Gregg’s tube tongue).

By that time, Tyler had also found his way across the crossover and down to where I was, so together we explored the vegetation on each side of the old road leading down to the river. At one point while I was beating Senegalia greggii (catclaw acacia)—from which I collected a single C. rossi, Tyler called my attention to lycids (netwinged beetles) on Karwinskia humboldtiana (coyotillo). I came over and told him we should look for Elytroleptus—cerambycid beetles that mimic lycids but that are much less frequently encountered. I noticed that the bush was abuzz with bees and Pepsis wasps, unlike many of the other plants of this species that I’d seen further up. Within a few minutes he called out that he’d found one, and within a minute or two I found one as well (I believe they are E. divisus). We searched the stand thoroughly but found no more and continued down the old road—our focus now on inspecting the patches of coyotillo along the way instead of beating the mesquites and acacias (by the way, I never got anything more off the dead acacias after beating the single C. rossi off the first one!). A little ways down the road, Tyler saw another Elytroleptus fly up from a coyotillo bush—also in flower and abuzz with bees and Pepsis wasps, and when he swung his net at it I saw another one fly up from the bush and netted it. This happened twice again on the way down, each of us seeing and netting an adult flying up from a plant and the other one doing the same immediately afterwards. As we neared the bottom of the road, we noticed the plants—more exposed than those further up—were now mostly past flower and were instead setting fruit with no beetles (or bees or Pepsis wasps) being seen.

Karwinskia humboldtiana (coyotillo).

After exploring the dry river bed for awhile, we headed back up the road and met Jason and Joshua looking at the very same plants from which we had collected the first Elytroleptus—although they had not yet seen that species. Joshua had just taken a swing at a Pepsis wasp on the bush when I saw another Elytroleptus fly up and away. Nobody else saw it, and I took off after it, successfully netting it to “win” the “Elytroleptus competition”! As we all walked up the road past the bridge, I noticed a R. parryi flower that was now hosting several A. neoneglecta, and we further noticed the Tiquilia canescens (woody crinklemat, shrubby tiquilia) flowers that were now open and collected a few A. neoneglecta and one A. starrae—a new western range extension.

Vachellia farnesiana (sweet acacia).

I didn’t find anything more the rest of the way up the old road, but once I got back near the vehicle I noticed Acmaeodera adults on flowers of Sida abutifolia (spreading sida)—most of which looked like A. neoneglecta but at least one possibly being A. opuntiae, collecting around half a dozen total. By then everybody was ready to go into town and look for something to eat, so I cut up and bundled the wood I’d collected and we drove back into Comstock. (Ironically, once back in town, we pulled up to the local eatery got out of our cars. As we approached the door, somebody inside turned the sign from “Open” to “Closed”! I joked that probably the motel owner had seen the #Libtardandproud sticker on my car window and called the restaurant to warn them. It’s a conspiracy, I tell you!)

Seminole Canyon State Park
Rio Grande & Canyon Rim Trails
Val Verde Co., Texas

After being denied service at the restaurant in Comstock, we drove to the state park and ate lunch at the picnic area (sardines and Triscuits for me) before divvying up the “oak-hunting duties”. We were hoping to see Spectralia roburella, an oak-associate that I have reared from Quercus fusiformis (plateau live oak) wood I collected at this site many years ago but which I still have yet to see in the field. Jason and Tyler took the Windmill Nature Trail, which has a stand of oaks, while Joshua and I took the Canyon Rim Trail (via the Rio Grande Trail), which has a couple of oak stands within a mile of the starting point. None of us had any luck with S. roburella, which I beat thoroughly as well as broke apart some of the dead branches, or anything on the trees for that matter. I did, however, collect a single Chrysobothris analis on Senegalia greggii (catclaw acacia) and a few clytrine chrysomelids on Vachellia vernicosa (viscid acacia) along the Rio Grande Trail, and shortly after starting down the Canyon Rim Trail I beat two Spectralia robusta from Diospyros texana (Texas persimmon). This latter capture renewed my enthusiasm for beating persimmon, which I did thoroughly whenever I was not beating oak, but I never saw another individual! At this point, my body was giving out (in spite of the much more tolerable temperatures today), and we regrouped to decide our next move for the trip (hint: we moved west!).

Vachellia vernicosa (viscid acacia).

We drove west to Sanderson and ended up in an RV park with tent sites for $7 and a few marvelous metal dinosaurs at the entrance (I was impressed with their selection of the rarely featured Allosaurus instead of the grossly overused Tyrannosaurus). It was not until around 2 am, however (and again at 5 am)—when the train rumbled by behind the campground—that we understood why the tent sites were so cheap!

Metal triceratops!
Metal allosaurus!
Ted MacRae, Jason Hansen, Joshua Basham & Tyler Hedlund.

Day 8 – To Monahans Sandhills

Despite its small size, the city of Sanderson offers a right nice cup of coffee to start the day by way of this retooled automobile dealership.

Ferguson Motor Co. retooled as a coffee shop.

17 mi N Sanderson, Jct US-285 & FR-2400
Terrell Co., Texas
Our plan had been to continue traveling west to Ft. Davis, but the weather forecast for that area called for rain and cool temperatures. We decided instead to travel northwest to Monahans Sandhills State Park where the forecast looked much better. The spot has been on my radar ever since the species Chrysobothris mescalero was described, and I’ve already looked for the species there twice without success. It was a good decision (more on that later). On the way, we saw a roadside area with lots of flowers in bloom, so we made a quick stop to see what might be visiting the flowers. I’m glad we did—I picked up a nice little series of Acmaeodera paradisjuncta along with a few A. mixta and some large bees (for Mike) on flowers of Wedelia hispida (Texas creeping-oxeye).

Acmaeodera paradisjuncta (family Buprestidae) on flower Wedelia acapulcensis (Acapulco wedelia).

I found it interesting that they were not on the much more abundant Coreopsis flowers and mentioned this to Tyler, who said he did see one on “this other yellow flower”—which turned out to be Senna roemeriana (two-leaved senna). I told him this was the host plant for Agrilus obtusus and that he should be on the lookout for the adults, which can be seen sitting on the leaves. I walked to another plant a short distance away, and there they were—two adult A. obtusus sitting on a senna plant, which I gave Tyler the chance to see before placing them in the bottle. Heading back towards the cars, we encountered a patch of Croton pottsii (leatherweed). I mentioned to him that this was the host of Agrilus lacustris, and almost immediately afterwards I saw two adults sitting on the foliage of one of the plants. It was a nice little stop that added one more species to my trip list.

Agrilus lacustris (family Buprestidae) on foliage of Croton pottsii (leatherweed).
Tyler examines Croton pottsii (leatherweed) looking for Agrilus lacustris.

Monahans Sandhills State Park
Jack Pump Picnic Area
Winkler Co., Texas
We arrived at the park early in the afternoon and, after checking in with the office, headed to the Jack Pump Picnic Area. Several Prosopis glandulosa (mesquite) were lining the parking lot, and with the first whack of a branch an Acmaeoderopsis sp. fell onto my sheet. I then spent a fair bit of time working the mesquites and collected not only a good series of Acmaeoderopsis but two different species of treehoppers and other miscellaneous types of beetles. I remembered collecting Acmaeoderopsis on mesquite a few years ago near Kermit, Texas and recalled their habit of dashing off the beating sheet when the day heated up and finding it easier at that point to net them as they flew to the tips of branches. Temperatures were still relatively moderate, so they were not yet doing that, but I started to look at the higher branches to see if I could see them flying to them anyway. I did not, but I did see small silhouettes of something buzzing around the flowers. I took a swipe with the net, and to my astonishment the net was filled with ghostly pale-yellow bees. I collected a few for Mike Arduser, feeling confident that he would find them of interest.

Me using a beating sheet to collect Buprestidae from Prosopis glandulosa (mesquite). Photo by Joshua Basham.

After working the mesquites around the parking lot (and having a hot dog with the very friendly family enjoying their holiday at the dunes), I moved out onto the dunes to see what might be going on. I had intended to look for stands of Quercus havardii (shin oak) to look for Chrysobothris mescalero but got distracted when I saw more Acmaeoderopsis—now flying to the branch tips of a line of mesquites. I spent a bit more time thus distracted but ended up with a nicer series of the species and then went back to the road to resume my search for stands of oaks. Along the way I collected an Acmaeodera immaculata and a Batyle suturalis on the flowers of Thelesperma megapotamicum (rayless greenthread) and a Lytta reticulata that was feeding on the flowers of Penstemon ambiguus (gilia bearstongue). At that point, Joshua came back with a Chrysobothris sp. that he’d collected on shin oak—it was definitely a member of the C. femorata species group but looked too large to be C. mescalero (could be C. caddo). Nevertheless, we were encouraged to focus our efforts at that point on searching the shin oak stands for the species. For this we decided to move over to the Shinnery Oak Picnic Area, but before doing this we again obliged the very friendly family, who were anxious to share with us more hot dogs!

Lytta reticulata feeding on flower of Penstemon ambiguus (gilia bearstongue) in sand hills.

Monahans Sandhills State Park
Shinnery Oak Picnic Area
Winkler Co., Texas
I searched this area twice last year looking for Chrysobothris mescalero, both times finding branches that looked “flagged” among the abundant stands of Quercus havardii (shin oak) in the area but failing to rear out the beetles. I had also swept the plants a bit each time but came up empty. This time, with four of us trying, we intended to give it a full effort—and it didn’t take long! Jason found the first one… and the second… and I joined him to see exactly where and how he was finding them. He had been sweeping the stands of plants in one of the depressions in the dune, while I was working the plants on the upper slopes and ridge tops. I decided to try working the depression instead and immediately came up with one myself. I worked the depression fully and got four for my efforts, then started working nearby plants that were not in depressions. I reasoned that being nearby they had a better chance of hosting beetles. Apparently this was poor reasoning, because I didn’t encounter any more beetles in the ensuing half-hour of sweeping.

Habitat for Chrysobothris mescalero—depression in sand dunes with stands of thigh-high Quercus havardii (shin oak).

Exhausted, I went back to the car to rehydrate and debate whether I wanted to continue, but the four of us motivated each other and back out we went. This time, I went west of the picnic area and found a depression similar to the first with mostly thigh-high plants… and got another four. I spotted another very nice-looking depression across the road and made my way over, again focusing on the knee-high plants. From this depression I collected three more adults. One final depression—and one more more beetle, and I was able to return to the car satisfied. I was excited to have figured out the secret to collecting these beetles and was anxious to share with the others. As it turned out, each of the others had also learned this secret, and collectively we had a very nice series of the beetle. We sat at the picnic bench and recounted what a good day of collecting it had been and, to celebrate, decided to head into town for pizza!

Jason Hansen, Tyler Hedlund, Ted MacRae & Joshua Basham. Photo by Jason Hansen.

Day 9 – Davis Mountains

Monahans Sandhills State Park
Shinnery Oak Picnic Area
Winkler Co., Texas
We had hoped to setup lights for insects at Monahans Sandhills State Park last night after returning from our celebratory pizza dinner, but extreme wind made not only that but even camping untenable. We tried to make it work, but the wind eventually blew my tent completely from its moorings, and I had no choice but to run into town and find a motel room.

This morning we set out for the Davis Mountains (our original plan yesterday), but on the way we stopped at this spot to see if we could find Agrilus cochisei on Ambrosia occidentalis (western ragweed). Jason, Tyler, and I had all collected the species in numbers near this spot over the past couple of years, and we wanted to give Joshua the chance to collect them as well. Unfortunately, the plants were not nearly as abundant or well developed as in previous years, and nary a beetle was to be found. While Joshua looked for the beetle, I swept Xanthisma spinulosum (spiny goldenweed), which was in bloom abundantly along the other side of the road. Two Acmaeodera mixta were swept from the flowers, and coming back I saw two more Acmaeodera—one A. neglecta/neoneglecta and one of a new species that Jason is describing—on the same. Back near the car there was a patch of Sphaeralcea angustifolia (narrow-leaved globemallow), which I swept hoping to find one of any number of buprestid/cerambycid species that could be on the plants but ended up only with a few weevils (Bob Anderson will be happy, however). There were a number of other flowers in bloom, but we avoided the temptation to look around further, as the Davis Mountains beckoned!

Xanthisma spinulosum (spiny goldenweed) along roadside.

Davis Mountains, 11 mi W Ft. Davis
Point of Rocks Picnic Area
Jeff Davis Co., Texas
We were a bit disappointed at how dry things looked as we climbed into the Davis Mountains and made our way to Ft. Davis. We noted flowers in bloom along the roadsides, but the grass along the roadsides and covering the hills was bright, crispy brown and the oaks were largely still without any new foliage. Nevertheless, we hoped collecting might still be good as there had been a little bit of rain in recent days. Point of Rocks Roadside Park is one of my staple collecting localities in the Davis Mountains—it’s where I first collected cerambycids of the genus Elytroleptus hiding amongst the much more numerous Lycus beetles—the latter poisonous and colored orange and black to advertise that fact, and the former completely harmless but similarly colored in an effort to fool would-be predators; and it’s also where I first reared what would become the holotype of Mastogenius texanus and later not only collected a good series of the adults but also discovered its larvae in branches of oak. This time, the oaks lining the picnic area showed no signs of new growth, so I didn’t even try beating on them. In addition, the wind was so extreme that trying to use the beating sheet would have been utterly futile. Instead, I walked the roadside inspecting the variety of flowers in bloom for beetle activity. Unfortunately, not a single beetle was seen despite the diversity of blooms, but I did pick up a couple of Lithurgopsis apicalis (orange-tipped woodborer bee) on a flower of Opuntia engelmannii (Engelmann’s prickly pear) for Mike. Having run out of flowers to check, and still thinking beating the oaks would be futile, I went over to the stand of Sapindus saponaria (soapberry), which were just beginning to produce foliage and where I’d collected Elytroleptus so many years ago (1994, I believe), to see if there might be any of the buprestid species associated with soapberry. I swept some of the lower branches, taking care to keep the still-extreme wind from catching my net like a sail, and saw a couple of Agrilus ornatulus in the net. I noticed when I got low and in certain positions around the grove of trees, I could minimize the wind, so I went back to the car and traded my sweep net for my beating sheet. Using the beating sheet in that kind of wind was a real challenge, but I still managed to collect seven adults of the species along with a number of clytrine chrysomelids. After having done this, I was less pessimistic about our prospects for collecting in the Davis Mountains—if only we could get out of the wind! We huddled and decided to go to Madera Canyon—all the way on the other side of the loop around the mountains, but higher in elevation and possibly more protected from the winds blasting up from the south.

Lithurgopsis apicalis (orange-tipped woodborer bee) in the flower of Opuntia engelmannii (Engelmann’s prickly pear).

Davis Mountains
Madera Canyon Preserve

Jeff Davis Co., Texas
Arriving at the trailhead parking lot, we were happy to see that the brutal winds that had harassed us for the past couple of days did not follow us up the mountain, and with temperatures not expected to exceed the high 80s it seemed a beautiful day was on tap. Hopefully the collecting would follow suit. We tapped on this plant and that as we entered the preserve, not seeing much (and not yet expecting to), and by the time the trail reached the creek bed crossing and began to ascend the mountain on the opposite side we began scattering in different directions. I continued following the creek bed and noticed that, while most of the oaks still were showing no signs of beginning to leaf out, the occasional tree was leafing out nicely. I beat the first such one that I encountered—Quercus grisea (gray oak) but collected only a few clytrine chrysomelids. Then I noticed a shrub in bloom—Fallugia paradoxa (Apache plum)—and beat a few miscellaneous beetles from it but still no buprestids.

Fallugia paradoxa (Apache plum) in oak-juniper woodland.

A bit further down the trail I encountered a large Quercus vaseyana (Vasey oak) along the creek bed that, unlike most of the species, had broken bud and was developing new foliage. I whacked a few branches and collected only miscellaneous beetles, but then I whacked a branch and saw a large Agrilus sp. (likely A. albocomus)! At last, not only a buprestid, but one that I had never collected before. I crossed back over the creek bed and noticed another gray oak with fresh foliage, and the first whack produced several beetles, including a strange, narrowly triangular-shaped beetle that I soon realized was Brachys querci. This was even more exciting, as I have only collected a scarce handful of western Brachys—and never this one, making the genus a big target of mine for this trip. Things stood as such for awhile, save for a few miscellaneous beetles that I beat from Senegalia greggii (catclaw acacia) and with only the occasional large, sparsely-leaved Vasey oak to beat as I went down the creek bed. I noticed another creek bed joining the one I was on and decided to explore up that valley, shortly encountering a small but well-leaved Vasey oak. I gave the tree a few whacks, and there on the sheet were three more Agrilus and one more Brachys! Okay, it seemed I was onto something by focusing only on oaks with new foliage. After relaying this information to the others, Tyler and I worked together up the valley, working each gray oak with fresh leaves that we could find (no more Vasey oaks were encountered). Over the course of the next hour, we added sparingly to our Brachys totals (with lots of clytrine chrysomelids and other miscellaneous beetles) until I gave a branch a whack and saw what I first thought was a much smaller species of Agrilus but then realized was an Agrilaxia. In this area, it could only be A. texana, a species I hadn’t seen since 1984 (before it was even described) until encountering them on my recent trip to northwestern Oklahoma. Shortly afterwards, Tyler and I came upon two groupings of large, freshly-foliated gray oaks—one bordering the dam of the now-dry Chico Tank and another further up on the western slope leading down to the former waterline. Tyler began working the near group, and I told him that I would go work the farther group because it looked like (and I hoped it would be) the “buprestid motherload”! That comment was prescient, as I just about doubled my series of the three species I’d already collected and added a couple of Chrysobothris axillaris! By the time we finished working the trees, we were exhausted but fully satisfied and began working our way back down the valley to the parking lot. We encountered Jason and Joshua near the trail at the base of a large gray oak—they’d also had great collecting and even found one larva and one adult of A. texana inside a dead branch of the tree. A beautiful mature male Aphonopelma hentzi (Texas brown tarantula) entertained us for a bit, and after spending four hours at the preserve we decided that yet another celebratory dinner at the Mexican restaurant in town was in order.

Aphonopelma hentzi (Texas brown tarantula) adult male in oak-juniper woodland.
Aphonopelma hentzi (Texas brown tarantula) adult male in oak-juniper woodland.

After dinner, we decided to head west for lodging, but to our surprise the lodge was closed—seemingly abandoned, and we had to double back all the to Alpine to find an open motel. I was biting my fingernails as I pulled into the first gas station I saw with my car’s miles-to-go indicator reading “1” mile! Our greeter at the motel seemed appropriate for the situation.

Our greeter at the motel in Alpine.

Day 10 – El Paso area

Rio Bosque Wetland Park
El Paso Co., Texas
We made the three-hour drive to El Paso for a couple of days of collecting in this area. I would have liked to have spent another day in the Davis Mountains, but Joshua had to fly out of El Paso this afternoon and we had no choice but to move on. It’s a good thing we did! Our first spot to explore in this area was selected based on the recent capture of Knowltonia atrifasciata—a very uncommonly encountered buprestid that none of us have seen belonging to a small genus that none of us have collected—a few years ago at this wetland park right on the Rio Grande River. Our GPS coordinates did not direct us straight to the park, but twice tried to direct us across the border into Mexico! We were too smart for that, and with a little online sleuthing and manual Google Map use we finally found the spot. Despite the name, the place was bone dry, and in what is becoming a daily ritual we had little optimism for how good the results would be. Knowltonia is associated with Atriplex, and we surmised that K. atrifasciata should be associated in this area with A. canescens, which we found occurring abundantly in some areas but not so much in others. I thought beating would be the best way to encounter the species—assuming it was present, so we all grabbed our beating sheets and started working through the area leading from the parking lot. The plants, which are wind-pollinated, were in full flower and released clouds of pollen with each whack of the stick, so in short order I was covered with pollen dust from head to toe. After beating for a while and not seeing anything, I became distracted by the Prosopis pubescens (screwbean mesquite) trees—a species I had never seen before and always wondered if I would be able to distinguish from the “normal” mesquite (P. glandulosa) with which they were interspersed, but which turned out not to be a problem due to their distinctively “corkscrewed” pods. I did get a few beetles of the species—one Chrysobothris rossi and a few miscellaneous beetles.

Prosopis pubescens (screwbean mesquite).

I turned my attention back to the Atriplex when I noticed emergence holes (which I presumed were the work of Knowltonia) and began breaking/cutting branches showing such holes to see if I could find at least a carcass in decent condition or—better yet—an unemerged adult. The frass was packed tightly in the galleries, which I took to be evidence that the galleries were the work of buprestid larvae and not cerambycids, the latter which I assumed always had open larval galleries kept free of frass by the larvae. For a while, in every stem I broke or cut into, the larval galleries appeared too old and I didn’t encounter either adults (live or dead) or larvae until I happened upon a cerambycid carcass in its pupal chamber in one of the branches. The head was missing, so I wasn’t sure at first if it was a cerambycid, but when I pulled it from the branch and got a better look at it I was convinced it was and placed it in a vial.

About that time, Joshua directed my attention to a nearby Salix gooddingii (Goodding’s willow) tree off which he and Jason had beat a fair series of an Agrilus sp. I assumed it was A. politus, but he said it wasn’t… than I assumed it was A. quadriguttatus, but it had complete and well-developed elytral vittae rather than spots. I couldn’t think of another species it might be and went to the tree to see if I could get some specimens as well, partly pessimistic since two people had already worked the tree but partly optimistic because I had one thing they did not—an extensible handle on my net! I extended the handle to its full length and swept the foliage of the upper branches thoroughly, getting four on that first round (and a Stenelytrana gigas to boot—my first time collecting that species)! I came back to the tree twice more, getting six more Agrilus on the first return and nine more on the second. At first glance, the adults look nothing like anything i was familiar with from Texas (or the U.S. for that matter), especially among willow associates—I’ll have to take a closer look to determine if it represents a species known from Mexico but not yet recorded from the U.S. or possibly even a new species! [We later determined the species to be A. fisherianus, which has not yet been recorded from Texas, so not a new or Mexican species but still a new state record.] By this time, I had given up on seeing Knowltonia and spent a fair bit of time beating both P. pubescens and P. glandulosa—the most significant capture being Acmaeodera delumbis on the former.

Stenelytrana gigas swept from foliage of Salix gooddingii (Goodding’s willow). Yes, it’s a crappy photo, but it was the best I could do with this very uncooperative subject!

Eventually the time came for Jason to take Joshua to the airport, so we said our goodbyes and made plans to meet up with Jason again at San Felipe Park near Fabens. As Jason and Joshua drove away, Tyler and I pondered our next move, and at that moment something out of the corner of my eye caught my attention. I looked at the Atriplex bush right next to me, and there near the tip of one of the branches was the unmistakable silhouette of a bright blue-green chrysobothrine beetle! I yelled out “That’s it!” and instinctively took a swing at it. Unfortunately, my net was in the wrong position, but I twirled it quickly and took an assertive, albeit one-handed, swipe just as the wary beetle quickly took flight—missing it by inches! There are times in the life of a field entomologist when a missed capture causes true heartbreak, and this was one of them. I was both thrilled we had seen the species after all but dejected beyond belief that I had missed it, perhaps without another chance to look forward to. Okay, time to get to work! We knew with certainly that the beetles were there and that they were active, and we also realized that an aerial net rather than a beating sheet would be the best way to go after them. I called Jason (en route to the airport with Joshua) to tell him to come back here instead, and then Tyler and I began slowly approaching each bush to look for that flash of blue-green near the branch tips, working the same plants we’d already tried beating earlier in the afternoon. It only took about 15 minutes before I spotted movement on a plant and saw the unmistakable silhouette of a chrysobothrine beetle on the back side of a branch near its tip. The beetle made another slight movement, triggering an instantaneous and assertive swipe of my net—this time already in proper position. My heart skipped a beat when I first looked into the net and did not see the beetle, but soon it appeared as it tried to fly up the net—it’s fantastically biramous antennae easily visible and confirming it to be a male. What earlier had been heartbreak turned to elation and vindication—we’d come here to find a very uncommon species, and we would be walking away successful once again. That said, the next hour and a half would further emphasize just how truly fortuitous a capture it was, as we never saw another adult despite combing the area thoroughly.

When I came back to the car, Jason—as yet unsuccessful in his more brief effort to look for the beetle—was tearing apart an Atriplex bush looking for evidence of larvae. This motivated me to give it another crack, but this time instead of working branches with emergence holes, I worked those without. I had noticed larval galleries in nearly every branch I looked at, and when I found fresh-looking frass in a gallery down the middle of an otherwise healthy, living branch I started carefully following the gallery, eventually finding a pupal cell with something in it and cutting away the wood to expose another longhorned beetle (this one teneral). Jason did the same and found two pupae, which he gave to me and which I will keep alive (along with the teneral adult) to let them emerge and harden up. The adult reminded of the genus Amannus, which I thought contained two species further west, but after consulting Larry Bezark’s photographic catalogue of Cerambyidae I realized they were Amannus atriplici—a Texas species that is yet another new one for me.

Amannus atriplici teneral adult in its pupal cell in a living branch of Atriplex canescens (fourwinged saltbush).
Amannus atriplici teneral adult (top) and pupa (bottom) in their pupal cells in living branches of Atriplex canescens (fourwinged saltbush).
Amannus atriplici teneral adult in its pupal cell in a living branch of Atriplex canescens (fourwinged saltbush).
Amannus atriplici teneral adult crawling out of its pupal cell in a living branch of Atriplex canescens (fourwinged saltbush).

By this time, we had been in the park for 5½ hours—a record for this trip, so we went into El Paso to have yet another celebratory dinner (Olive Garden this time) and plan our next move.

Sunset north of El Paso, Texas.

Day 11 – El Paso to Cloudcroft

Rio Bosque Wetland Park
El Paso Co., Texas
Last night was another lodging fiasco that had us driving from one closed campground to another closed campground before biting the bullet and securing a motel in Fabens. It was our intention to collect this morning at nearby San Felipe Park, where last year I’d done well with Gyascutus planicosta and several cerambycids. However, as soon as we arrived we heard a series of gunshots (this is Texas, after all!). If that wasn’t enough to give us pause, both Jason and I realized when we looked in the backs of our vehicles that we’d left our beating sheets on the trail at yesterday’s last locality (we’d ditched them in favor of having two hands on the aerial net while looking for Knowltonia atrifasciata). The collecting gods seemed completely against us collecting at our intended location, so we blasted back to Rio Bosque Wetland Park and were relieved to find our beating sheets on the trail right where we’d left them. None of us were keen on going back to the gunshots, so we decided to stay and take another look around for K. atrifasciata. It would also give me a chance to collect potentially-infested stems of Atriplex canescens and rear out a better series of Amannus atriplici. I inspected all the plants carefully in the areas around the two spots where I’d seen the beetles yesterday but did not see any adults. As I did this, I beat a few Prosopis pubescens (screwbean mesquite) and collected a few miscellaneous beetles, then went back to the spot where I’d collected Agrilus fisherianus on Salix gooddingii (Goodding’s willow) and used the extended handled net to sweep four more adults from the upper branches of the tree. Finally, I went back to the plant from which Jason and I had cut pupae and teneral adults of Amannus atriplici (the two pupae we collected yesterday had since emerged as adults inside their vials) and cut several whole branches from the plant to bring back for rearing. A suspicious character got us a little on edge, and we decided we’d seen enough and headed up north into New Mexico.

Stems of Atriplex canescens (fourwinged saltbush) infested with Amannus atriplici cut up and packed in a temporary emergence box.

Hwy 70 at Point of Sands
Otero Co., New Mexico
Our plan was to go to Cloudcroft and look for the recently-described Brachys rileyi. Before doing that, however, I wanted to stop at Point of Sands, a cool place where White Sands National Monument spills across the park border and down onto Hwy 70. Last July when I stopped here with Jeff Huether, I found a carcass of Sphaerobothris ulkei and figured they might be out at this earlier point in the season. It ended up being a good thing that we stopped here, because little did we know that that plan had already been thwarted by closure of the National Forest due to extreme fire danger (we would not find this out until we were ready to leave).

The author ready for netting buprestids. Photo by Jason Hansen.

I crossed the road to start checking the plant host for the species—Ephedra torreyana (Torrey’s jointfir) in this case. Conditions were again not hot with a light breeze—much more pleasant than the blazing hot conditions at the beginning of the trip or the brutal winds that followed. The ground, however, looked parched, and I was not optimistic about anything being out. I had gotten nearly to the end of the Ephedra stands on that side of the road without seeing anything (and was starting to think I never would) when I noticed a bee-like insect hovering around the tip of one of the Ephedra plants. I realized it was an Acmaeodera and quickly netted it. When I pulled it from the net, I was thrilled to see it was Acmeodera recticollis—an uncommonly encountered species that I’ve never collected before. I told Jason and Tyler what I had found, and the three of us spent the next hour working the ephedra plants in the area, collecting a pretty nice series of the species.

Jason and Tyler search for Acmaeodera recticollis on Ephedra torreyana (Torrey’s jointfir, Torrey’s Mormon tea).

I diverted my attention only a couple of times—once when I saw a striking robber fly perched on yucca that I just had to photograph, and then again when Tyler and I found the spectacular adult males and females of Tragidion armatum on flowers of Yucca elata (soaptree yucca). By then, we’d learned that the National Forest was closed but had managed to reserve a camping site at a commercial campground just outside the national forest boundary. Our plans to collect Brachys rileyi today might have been thwarted, but we got an unexpected species for the trip as a consolation, and I was really happy we would not have to worry about accommodations for this evening.

Efferia sp. male (family Asilidae).

Sacramento Mountains, Mayhill
Otero Co., New Mexico
All we could do as we drove through the National Forest around Cloudcroft was stare longingly at the Quercus gambelii (Gambel’s oak) while signs posted at any potential pulloff flashed “Closed. No Entry.” Our commercial campground was, however, just outside the National Forest and looked to contain borderline-equivalent habitat, so we remained hopeful that we would be able to find Q. gambelii trees to beat for Brachys rileyi. While setting up camp, we saw a gorgeous red netwinged beetle (family Lycidae) that proved to be Lygistopterus rubripennis, and sweeping the lush vegetation along the nearby spring-fed creek produced several more along with a few other miscellaneous beetles. There was a trail leading from the campsite into the oak-juniper-pine woodland, and walking along it I found several small Q. gambelii—the tree we were looking for. Unfortunately, beating all the branches I could reach on all the trees I could find failed to produce B. rileyi (or anything else), so for the time being our quest for the species remains unfinished.

Quercus gambelii (Gambel’s oak).

Further down the trail under a powerline clearing I found stands of Q. havardii, which I swept in hopes of finding Brachys barberi but found only a couple of cryptocephaline chrysomelids and a clerid. I met up with Tyler and Jason on the other side of the creek and swept Cucurbita foetedissima (buffalo gourd) in hopes of finding Adetus brousi (another species I haven’t yet collected myself), but no cigar (and still haven’t collected it myself). There was a stand of Salix exigua (sandbar willow) nearby, and sweeping off of it produced only a series of galerucine chrysomelids.

Phlox nana (Santa Fe phlox).

As dusk approached, I setup the ultraviolet lights only (no mercur vapor lamp since I couldn’t fire up the generator), but I could tell the temperatures were cooling to the point that no beetles would be coming in. This proved to be the case, and with only a few lonely moths sitting on the sheet I turned to roaming the roadsides looking for night-active insects. Tyler and I found several Udeopsylla robusta (robust camel cricket), all barfing, defecating, and assuming hilariously defensive poses in response to our proddings.

Udeopsylla robusta (robust camel cricket).
Udeopsylla robusta (robust camel cricket).

Darkling beetles (family Tenebrionidae) were the main things collected, but at the top of the drive I found three Zopherus concolor, with their cool leg “racing stripes,” crawling slowly on the ground. By that time, I was exhausted from yet another long but successful day of collecting and turned in for the night.

Zopherus concolor (family Zopheridae).
Zopherus concolor (family Zopheridae).
Zopherus concolor (family Zopheridae).

Day 12 – Mescalero Sands

Mescalero Sands Recreation Area
North Dunes Picnic Area
Chaves Co., New Mexico
In the morning after breaking camp, we went back to Cloudcroft to see if we could park somewhere along the side of the road and access Quercus gambelii (Gambel’s oak) to look for Brachys rileyi, but all the places we could pull off were well marked “Stop. No Entry” due to the Lincoln National Forest closure. Our quest for B. rileyi would have to wait for another day. With that, we headed for Mescalero Sands, where we hoped to get another shot at collecting another Brachys species we were targeting—B. barberi. We’d hoped to see it at Monahans Sandhills State Park at the beginning of the week but found only the other main target—Chrysobothris mescalero. I was also happy to have another shot at collecting Agrilus hespenheide, a single specimen of which Jason collected at Monahans and still represented in my cabinet by only a single specimen collected many years ago at this very site. We also welcomed the opportunity to find C. mescalero at its type locality. Temperatures were already topping 100°F by the time we arrived, but the tough conditions would mean little to us since we immediately found B. barberi sweeping stands of Quercus havardii (shin oak) very near where we parked. As we were beginning our sweeping, Jason encountered a small white yogurt cup that had been discarded in the dunes and noticed a few Acmaeodera quadrivittatoides flying in and landing on the white rim of the cup. Jason collected them and then left, after which I emptied the sand and filled the cup with water to let it sit while I swept the stand of oaks next to it. When I returned, there were seven Acmaeodera trapped in the water—two clearly being A. quadrivittatoides but the other five not immediately recognizable (they look like very small A. starrae or A. riograndei—I will be anxious to look at them more closely). Over the next 2½ hours we would sweep the stands of shrubby oaks looking for B. barberi, often collecting none but sometimes getting as many as five individuals per sweeping pass. I worked each discrete stand systematically to avoid duplicative sweeping, as it was hard work in such heat, and in addition to B. barberi I collected two specimens of A. hespenheidei, four specimens of C. mescalero (two got away, as they moved fast in the heat!), two Acmaeodera neglecta, and another small Acmaeodera that looks very much like A. riograndei, which we collected much further south at Gold Mine Canyon in Val Verde Co., Texas. This latter find is significant , as the species had not been recorded before our trip from outside of the Big Bend region—if the ID is correct, the capture represents a new state record and a significant extension of the known range. In addition to the buprestids, treehoppers of the genus Cyrtolobus or near (two species) were abundant on the oaks, along with a gorgeous species of cassidine leaf beetle. I collected nice series of each and a smattering of many different species of beetles in other families. I was having so much success sweeping that eventually I had to force myself to stop and take a break to rest and rehydrate (I’m not 26 anymore!). I thought I wanted to go out for more A. hespenheidei after the break, but after the first sweeping pass I realized my body was done!

Sand dunes at Mescalero Sands Recreation Area.

Mescalero Sands Recreation Area
Vic. North Dunes entrance
Chaves Co., New Mexico
After finishing with the oaks in the recreation area, we headed out to the highway near the entrance to work the nearby stands of Sapindus saponaria (soapberry). I had good success during my last visit here in 2018 with Agrilus sapindi in these stands, a species I have found very sparingly in other locations and which were best collected here by sweeping the small saplings rather than beating the branches of larger trees. We encountered the species almost immediately after we arrived, collecting them exactly as I’d done before (despite Jason’s skepticism). This also produced a few Agrilus ornatulus, and beating the branches produced one or two of each species as well. Jason also collected one individual of Agrilus limpiae, a species I collected on soapberry during my recent trip to western Oklahoma, but I did not encounter it here. I wonder what it is about this soapberry stand that A. sapindi likes so much?

Mescalero Sands Recreation Area.

We were all exhausted after working the sweep nets and beating sheets all day at Mescalero in the +100°F heat, so we ran into town to eat dinner (mine including a large milkshake!) before making the hour+ drive to our campsite near Carlsbad. Before leaving town, however, we had to stop at the alien memorial for photos and to leave our own contributions on the pedestal.

Ted MacRae, Jason Hansen & Tyler Hedlund.
My contribution to the alien memorial.

Capitan Reef Rd at Pecos River
Chaves Co., New Mexico
Jason knew a spot not far from the campground where we could setup the lights, and despite my exhaustion I agreed to go along since it seemed like it could be a good night. It wasn’t a good night—though it wasn’t a bad night either. Only two cerambycids were collected at the mercury vapor/ultraviolet lights—one tiny elaphidiine, and a conversely large Aneflus sp. (maybe A. prolixus). However, there was a smattering of other interesting insects that made the night not a bust—a few clerids of several different species, the same with bostrichids, lots of tiny bruchids (I don’t typically see these at lights—or maybe I just haven’t noticed them), a couple of small melolonthine scarabs, and one darnine treehopper. I had hoped for a greater diversity of beetles, especially longhorned beetles, but considering the success I’d had today I couldn’t complain.


Day 13 – Going solo!

Capitan Reef Rd at Pecos River
Chaves Co., New Mexico
We came back to the spot where we blacklighted last night to look for Gyascutus planicosta, which Jason had collected a few years ago on Atriplex (saltbush). Gyascutus planicosta adults are among the largest in the family in North America and are powerful fliers, so catching them can be a challenge even for the experienced entomologist. Sadly, we did not see any this time—I suspect we were a bit on the early side for the likes of such. The area was supremely uninteresting—choked with clumps of Prosopis glandulosa (mesquite) on raised mounds with ORV paths (apparently a very popular pastime in this area) winding amongst them and a few large, invasive Tamarix (tamarisk, salt cedar) trees. Nevertheless, I managed to beat a few centrotine and one darnine treehoppers and some miscellaneous beetles from the mesquite, and at the furthest point I walked before I’d had enough I encountered a small series of Acmaeoderopsis hulli flying to the tips of mesquite. At this point, it was time to bid Jason and Tyler adieu, as they started heading back to south Texas while I start making my way to Arizona. Jason and I each ended up with 43 species of Buprestidae for the trip, though he will likely get more on his way back to south Texas and I almost certainly will get many more (hopefully a few dozen!) over the next week in Arizona.

Leptotes marina (marine blue).
Checkered whiptail (Aspidoscelis tesselatus).

Sacramento Mountains, Mayhill
Otero Co., New Mexico
My path to Arizona took me right by the camp we stayed at two nights ago, where Jason had managed to collect a couple of Taphrocerus chevrolati (but I had not) by sweeping grasses along the side of the road. I did not see any sedges (typically the hosts of Taphrocerus species) mixed in the grasses and thought it was odd that these two individuals would be found on rather dry grasses. Nevertheless, since I had another opportunity I decided to see if I could come up one or two this time. I came up with 10 and don’t quite know what to think about this many individuals in patches of grasses that surely cannot be a larval development host. At any rate, these specimens will come in handy (the first I’ve collected in New Mexico) as I continue with my revision of the North American (north of Mexico) Taphrocerus fauna.

Roadside grasses from which I swept a series of Taphrocerus chevrolati.

1.8 mi W Cloudcroft on Hwy 82
Otero Co., New Mexico

Just west of Cloudcroft, I noticed a lot of Quercus gambelii (Gambel’s oak)—host for Brachys rileyi—along the edge of the road and a large turnout without any “Stop. No entry.” signs posted where I could seemingly park the car. I hoped maybe staying along the roadside would not be a problem with the National Forest closure and began beating the trees looking for the beetle. I’d worked the trees less than 15 minutes, collecting a few miscellaneous beetles and treehoppers, when a Forest Service vehicle pulled up alongside me and, in a friendly voice that didn’t belie whether they thought I was doing something wrong, asked what I was doing. I replied that I was collecting beetles, to which they smiled and explained that the National Forest is closed and I was in violation of the order. Clear enough! I thought of asking if I could just move along or did I need to go to jail, but I decided not to tempt fate and promised to leave at once. I was kind of surprised they left without sticking around to make sure I actually did leave at once—I suppose I must have a trustworthy face.

Bronco Sport in its native habitat.

Hwy 70 at Point of Sands
Otero Co., New Mexico

My route to (eventually) Arizona also took me by this spot that Jason and Tyler and I had visited two days ago, where we ended up finding Acmaeodera recticollis on Ephedra torreyana (Torreys’ jointfir). I’d gotten a modest series and wanted to see if I could get a few more, but what I really wanted was to find Sphaerobothris ulkei, which utilizes the same plant as a larval host. Tyler had seen two adults when we were here before but had not manage to capture them, both flying beyond the fence when he saw them. Winds were as severe as I’ve ever experienced, and at one point I had to use the extended handle of my net to prevent my hat from escaping on the other side of the barbed-wire fence lining the roadside. The A. recticollis adults were not nearly so abundant today as they were two days ago, but I managed another modest series with which I can be satisfied. I never did see a S. ulkei adult, although three times a similarly shaped/sized insect fooled me into thinking that I had one. I also checked the Yucca elata (soaptree yucca) for more Tragidion armatum but struck out.

White Sands National Monument spills out onto the highway.

One interesting story—as I was checking the Ephedra, a Border Patrol vehicle pulled up to check out my vehicle, then circled back around to check out me (there is a Border Patrol station just up the highway). The officer explained that they had been notified about a person “walking along the fence” and came to check up on it. It ended up being a very pleasant conversation as I explained what I was doing, each answer bringing up another question out of seemingly genuine interest. You know you’ve exceeded expectations when a Border Patrol officer extends their hand to shake yours when they’re ready to leave.

Her eggs all laid, life has ended for this Pepsis sp. (a tarantula hawk wasp). Meanwhile, her victims are now “unable to scream!”

Organ Mountains—Desert Peaks National Monument
Pine Tree National Recreational Trail

Doña Ana Co., New Mexico
For my final stop of the day, I’d planned to hike a trail on the other side of the mountains where Chrysobothris culbersoniana—a species I’ve not yet collected—has been recorded. However, as at the previous site the winds were so severe that using a beating sheet would have been impossible. I decided to instead take one more shot at Brachys rileyi. BugGuide shows a photo of a specimen collected at “Aguirre Springs” (presumably Aguirre Spring Campground), and iNaturalist shows another specimen with geo-coordinates very close to that spot along the Pine Tree Trail.

Approaching Organ Mountains—Desert Peaks National Monument.

Since both records state the beetles were collected on Quercus gambelii (Gambel’s oak), I kept an eye out for such. I never saw any on the parts of the trail that I hiked, but there was Q. grisea (gray oak) and the occasional Q. arizonica (Arizona white oak), and off the former I beat a few miscellaneous beetles but no Brachys querci (which has also been recorded here and which does utilize gray oak). At a few places I encountered Celtis reticulata (netveined hackberry), off which I beat a few more miscellaneous beetles including a single Agrilus lecontei celticola. In the meantime, I began to wonder if the geo-coordinates were accurate and left a comment on the iNaturalist record asking about it, hoping I might hear back before I left the area.

On Pine Tree National Recreational Trail at Organ Mountains—Desert Peaks National Monument.

Eventually I made it to the spot indicated by the geo coordinates on iNaturalist, but still the only oaks present were gray oaks. Unfortunately, by then I still had not heard back from the iNaturalist user who posted the record, so I surmised it was more likely that they had collected the species on gray oak and misidentified the host (I would not be surprised if this species turns out to use other oaks as hosts, as this is common among oak-associated Brachys) than the geo coordinates being incorrect. By then, I’d beaten enough oaks to conclude that neither Brachys species was active at the moment and headed back to the car.

On Pine Tree National Recreational Trail at Organ Mountains—Desert Peaks National Monument.

After I got into town later that evening, I saw a response from the iNaturalist user, who stated that the beetle was actually collected on the upper part of the Pine Tree Loop—not where indicated by the geo coordinates. Unbeknownst to me, Gambel’s oak does occur on the upper part of the loop, and in the user’s opinion the host ID rather than the geo coordinates should have been given weight for locating the plants. This is debatable—both are important and should be reported accurately, and the episode illustrates the importance of being cautious about relying on crowd-sourced data.


Day 14 – Travel Day (Las Cruces to Hereford)

Sunday was a day off from collecting while I drove west to the home of Norm Woodley and Steve Lingafelter in Hereford, Arizona. Norm and Steve are experts in Buprestidae and Cerambycidae, respectively, and were kind enough to host me during the third week of my trip for some pre-monsoon collecting in southeast Arizona. Insect numbers and diversity at this time of year may not compare with those seen once the monsoons start (usually in early to mid-July); however, there are a number of buprestid species (especially in the genera Agrilus and Brachys) that are generally seen earlier in the season and disappear by the time the monsoons arrive. I’ve collected a number of times in southeast Arizona during and after the monsoons; however, this would be my first attempt at collecting in the area before the monsoons. Thus, it was my hope that I would encounter many species that I haven’t seen before, especially some of those that are completely lacking in my collection. I didn’t arrive at Norm and Steve’s until late afternoon, so instead of collecting we enjoyed grilled burgers and fine spirits.


Day 15 – Dragoon & Huachuca Mountains

Before leaving for the Dragoon Mountains, Steve and I set out two jug traps in the wash on the south side of their house—one on the south side of wash baited with 50:50 red wine:ethanol and another on the north side baited with pure ethanol.

W side Dragoon Mountains
3.0 mi NE Rd 687 on N Middlemarch Rd

Cochise Co., Arizona
A few years ago, Norm discovered one of the rarest Acmaeodera species in North America—Acmaeodera horni—on flowers of Fallugia paradoxa (Apache plume) at a spot in the Dragoon Mountains. That was undoubtedly the biggest of the several priorities I had this week as I spend the last part of this three-week trip collecting in southeastern Arizona with Norm and Steve. The “horni spot” was first on our agenda for this first day of collecting with them, since Norm has seen them as late as June 1st but mostly in mid-May. When we arrived, we noticed the plants were a bit past peak bloom, as only a few sporadic flowers were present on the plants compared to the much more numerous fruiting structures. The plants are primarily along Clifford Wash, so we walked down the wash, looking at any flower we could find. At first I got distracted by the abundance of Acmaeodera quadrivitttatoides on the flowers and quickly collected my share. When Norm and I met up again about 15 minutes later, he had found two but I had yet to see one. Knowing they were still out, however, renewed my motivation to continue searching. After a while, I encountered Steve, and he too had caught one, while I still had yet to see one. I continued searching, and eventually I heard Norm call out my name. I came to where he was, and there it was—sitting on a flower that I had looked at not five minutes earlier! I easily netted the beetle, happy (and relieved) to have caught one but still wishing somewhat I could have found one on my own. I will just have to do that sometime in the future (with a mid-May trip). While I was looking for the beetle, I also collected a modest series of what may be Acmaeodera variegata to go along with the many A. quadravittatoides and three A. horni (Norm gave me his two). I also collected a couple of A. variegata and A. quadrivittatoides on flowers of Verbesina encelioides (cowpen daisy), as well as a very small lycid (the smallest I’ve ever seen) and a couple of chrysomelids on the white flowers of Mimosa aculeaticarpa.

Acmaeodera quadrivittatoides on flowers of Fallugia paradoxa (Apache plum).
Acmaeodera horni, taken from flowers of Fallugia paradoxa (Apache plum).

After scouring the patches of Fallugia one more time to ensure we had not left any A. horni behind, we drove 0.4 miles back down the road to another spot where the wash crossed the road and where Fallugia is again fairly abundant. We checked the plants thoroughly, and while A. variegata and A. quadrivittatoides were present, A. horni was not. With that, we said goodbye to the “horni spot” and proceeded to another spot where we suspected Tragidion armatum could be found on Yucca elata (soaptree yucca).

Hadoa simplex mating pair on Fallugia paradoxa (Apache plume).

W side Dragoon Mountains
0.4 mi N of N Middlemarch Rd on Rd 687

Cochise Co., Arizona
This location was very different from the previous, with mesquites dotting a dry grass plain and the occasional Yucca elata (soaptree yucca)—many sending up flowering stalks. I checked a few for Tragidion armatum but didn’t see any beetles on the stalks, and then I got the idea to use my extended net handle to pry the rosette of stiff, sharp leaves away from the base of the stalk to see if any adults were hiding there. Success! I found one female hiding in the rosette of the first plant I checked, another female in the third plant I checked, a male and a female in the fourth plant I checked (but then no more in any of the many plants I checked after that).

Tragidion armatum taken from stalk base of Yucca elata (soaptree yucca).

3.0 mi NE Hwy 80 on N Middlemarch Rd
Cochise Co., Arizona
As a final stop for the day (not considering blacklighting for insects at nightfall), we stopped at a spot where there is a stand of Sapindus saponaria (soapberry), off which Norm has collected a new species of Agrilus that he is describing. I let him sweep the plants to try to get more for the species description, while I headed towards an old gnarled Prosopis glandulosa (mesquite) off which Norm said he has routinely collected a variety of buprestids. I didn’t have quite the luck with it myself, collecting only a single Chrysobothris rossi and a few other miscellaneous beetles, but a neighboring mesquite with lusher foliage did produce a new species for the trip—Acmaeoderopsis junki and a second C. rossi. A large, partially fallen Vachellia constricta (whitethorn acacia) in full bloom looked particularly inviting for buprestids, but I beat only a few miscellaneous beetles from it. It turned out that Norm had already worked it and gotten a mating pair of Chrysobothris merkelii from it. I swept the soapberry after Norm had finished with it, but as he’d only gotten a single specimen of the Agrilus after working it, I didn’t have much expectation of finding another one (and I didn’t). By this time we were hot and tired, and a quick stop in nearby Tombstone with the hope of ice cream and drinks was singing her siren song.

E side Huachuca Mountains
Lower Hunter Canyon

Cochise Co., Arizona
After dinner, Steven and I went to nearby Hunter Canyon for some pre-monsoon blacklighting. Hunter Canyon is not nearly as well known as Miller Canyon, its much more famous neighbor to the north, but has similar habitat and is right in the heart of one of the coolest natural history spectacles of the insect world—the mass emergence of the cerambycid species Megapurpuricenus magnificus (formerly Crioprosopis magnificus)! This species develops as larvae in the trunks of living oaks and emerge once every three years in synchrony right after the first rains of the summer monsoons. For a brief few days afterwards, one can easily see a hundred adults flying high in the canopy—a true spectacle given their enormous size and fantastic red/black coloration. The spectacle last for only a few days, so seeing it requires careful planning and a bit of luck. The last emergence here was in 2021, so I am making plans to visit in 2024 in hopes of seeing this incredible phenomenon with my own eyes. For this evening, however, I would have to content myself with whatever pre-monsoon species might happen to be out and about. We set up our sheets just a bit out of sight from each other (to avoid competing for the same insects), each of us using one 175-w mercury-vapor (MV) bulb and two (me) or four (Steve) 15-w ultraviolet (UV) light bulbs. Since getting my MV bulb last year I have started using the setup/technique recommended by Steve and other cerambycid specialists—a vertical sheet with ground cloth on both sides, the MV on a tripod or stand raised to a level above the top of the sheet, and at least one UV light hanging on each side of the sheet. The MV light will attract beetles from a distance, but due to its brightness the beetles may land in the vicinity rather than on the sheet itself. For this reason, the MV light is then turned off after about an hour and all the nearby vegetation shaken or beaten to disturb the beetles, which are then attracted to the sheet itself by the UV lights. After a while, the MV light is turned back in and the cycle repeated. On this night, however, such technique would not be terribly important, as the number of beetles flying was rather small. The first cerambycid—Anelaphus simile—came to Steve’s sheet, and later an Anelaphus brevipes would also appear. A couple of the former also came to my sheet, and from both sheets I picked a smattering of miscellaneous beetles in other families. One of the more exciting finds of the night was a very late-occurring female of the spring species Knulliana sonorensis (I do not accept the current placement of this taxon as a subspecies of K. cincta), which was crawling on the ground in the parking lot and which may have been attracted to a pile of recently cut oaks that were near Steve’s lights. This prompted a search for other individuals that may have been so attracted, but the only thing we found was a headless carcass of Acmaeodera sp. (perhaps A. decipiens). When the expected 9:30 p.m. flush of cerambycids did not materialize, we called it a night, took down the lights, and headed back the house for some French Open replay action.

Choristostigma roseopennalis at ultraviolet/mercury-vapor light in mountain oak woodland.

Day 16 – Huachuca Mountains (cont.)

W side Huachuca Mountains
Lower Copper Canyon

Cochise Co., Arizona
I knew it would be difficult to top yesterday’s Acmaeodera horni experience, but I still had hopes of collecting at least a few species that I’ve never seen in the field before, especially if they represented species not present in my collection. Copper Canyon is a famous locality for collecting, and Steve and Norm like to visit this spot regularly due to the variety of interesting species they have seen over the years here. We walked the lower mile or so of the trail up the canyon, beating primarily the different oak species looking for mostly Agrilus and Brachys. Many of the Arizona species of Agrilus and most of the Brachys occur prior to the monsoons, and since I’ve only visited Arizona during or after the monsoons I’ve not collected many of the Arizona species in these genera.

Lower Copper Canyon Trail.

It took a while for the collecting to pick up, but eventually we started focusing on the occasional trees with newly flushed foliage and had good success. Quercus arizonicus (Arizona white oak) was the most productive, yielding good series of Agrilus quercus and A. chiricahuae as well as a couple of Brachys cephalicus—all species I’ve never collected before. Quercus emoryi (Emory oak) was less productive (owing to the fact that only a few trees were flush with fresh leaves), yielding two more B. cephalicus and an A. chiricahuae. Quercus hypoleucoides (silverleaf oak) was the least productive—no buprestids were collected off these trees, defying Norm’s expectations. Other than buprestids, a variety of other insects—mostly leaf beetles, weevils, and a good variety of treehoppers—were collected from all three oak species. About a mile up, the trail crossed a wash and got steeper and narrower, and I found it difficult to beat off of anything except trees right along the trail, so I turned around and joined Norm in slowly working our way back down the trail, beating/sweeping the trees along the way and adding slightly to our series. Once back at the bottom, I hung a jug trap baited with ethanol near the wash next to a large silverleaf oak, which Steve will check during the rest of the season and which hopefully will produce some nice species of longhorned beetles.

Lower Copper Canyon Trail.

Before moving to our next spot, we made a quick visit to the nearby U.S./Mexico border. The Nazi-esque barbed wire atop steel cross bars was a far cry from the promised “big, beautiful wall” as it slashed across the landscape and up over the mountains. I joked with Norm that we should stick our nets over the fence and sweep for insects so we could label them as having been collected in Mexico!

U.S./Mexican border.
On this side is the U.S., and on that side is Mexico.

W side Huachuca Mountains
Lower Ida Canyon

Cochise Co., Arizona
After a solid three hours at Copper Canyon, we were beat but wanted to check nearby Ida Canyon, the lower reaches of which has a stand of sedges from which Norm has collected several species of Taphrocerus. The sedge patch was completely dry and no Taphrocerus were found, so we beat/swept a bit on the oaks—again focusing on those occasional trees with a fresh flush of foliage. I got one Agrilus quercus on Quercus arizonica (Arizona white oak) and two Brachys cephalicus off of a single Quercus emoryi (Emory oak) along with a smattering of miscellaneous beetles off each. Once again, Quercus hypoleucoides (silverleaf oak) produced a smattering of miscellaneous insects but no buprestids. Beetle activity didn’t seem to be as high here as at Copper Canyon, so we didn’t spend too much time here and headed home for dinner before another night of blacklighting.

Lower Ida Canyon.

Huachuca Mountains
Montezuma Pass

Cochise Co., Arizona
On the way back to Norm’s and Steve’s home, we stopped at the ever-scenic Montezuma Pass, which offers spectacular views of the U.S./Mexico border to both the east and the west. A large Cylindropuntia imbricata (tree cholla) in full bloom added a splash of color to the tawny-colored view.

View west from Montezuma Pass.
View east from Montezuma Pass.
Cylindropuntia imbricata (tree cholla).

E side Huachuca Mountains
Upper Miller Canyon Rd

Cochise Co., Arizona
This spot near the top of Miller Canyon Rd is another of Steve’s favorite blacklighting spots, so we set up our lights here in the same manner as last night—just out of sight of each other along the trail and both setups using both ultraviolet and mercury-vapor lamps. Both light setups produced a smattering of miscellaneous insects but few longhorned beetles, a few Anelaphus (probably A. simile and A. brevipes) coming to my light but none coming to Steve’s. Who knows why this happens?

Menkeleon bellulus (family Myrmeleontidae) adult at ultraviolet/mercury-vapor light in riparian montane oak woodland.
Agrius cingulata (pink-spotted hawkmoth) at ultraviolet/mercury-vapor light in riparian montane oak woodland.
Cypherotylus californicus (blue fungus beetle) on trunk of juniper in riparian montane oak woodland at night.

Day 17 – Chiricahua Mountains

E side Chiricahua Mountains
Herb Martyr Campground

Cochise Co., Arizona
The Chiricahuae Mountains are about a two-hour drive from Norm’s and Steve’s place, so our plan for the day was for Norm and I to spend the day collecting in Cave Creek Canyon and then Steve meet up with me to blacklight. Our target for this first spot of the day was Agrilus howdeni, which Norm has collected on the fresh leaves of Platanus wrightii (western sycamore) during June. This spot is very near the type locality for the species (Southwestern Research Station), and though it was the first day of June we hoped it would already be out. The stop had an inauspicious start—as soon as we got out of the car we saw a large Chrysobothris land on the sheet metal eave of the campground outhouse. It escaped Norm’s grasp but returned, only to then escape my net as well. Not a good start, but I did at least beat a longhorned beetle (probably Sternidius decorus) from Quercus hypoleucoides (silverleaf oak) before we headed down to the creek bed where the sycamores could be found. Once in the creek bed, I beat a few miscellaneous beetles off of Quercus arizonica (Arizona white oak) before turning my attention to the sycamores. We quickly realized that A. howdeni was not only present, but occurred in the biggest numbers Norm had ever seen—apparently the species is an earlier spring species than realized and the individuals seen later in June were probably hangers on. Collecting for me was slow at first, as I was limited by my beating sheet and 6-ft extensible handled net to the lower branches of the trees, while Norm was able to reach the higher branches with his much longer net handle. Eventually, however, after hiking up the creek bed a fair distance I encountered several trees with plenty of low growing branches from which I was able to collect a good series of adults and then make my way back. Back at the car, Norm and I both checked the outhouse again hoping that the Chrysobothris had returned, but no such luck. I did notice, however, a few small Robinia neomexicana (New Mexican locust), from which I swept a single Agrilus egenus, and then Norm saw a large Buprestis (probably B. laeviventris) land on the parking lot sign and nabbed it. Having gotten our fill of A. howdeni, we decided to move up to a higher elevation spot where Gayle Nelson had once collected the very rare Brachys apachei on Quercus hypoleucoides (silverleaf oak).

Leptotes marina (marine blue) on flowers of Amorpha fruticosa (leadplant) in montane woak woodland.

E side Chiricahua Mountains
Rd 42 at East Turkey Creek

Cochise Co., Arizona

East Turkey Creek.

The forest type was different at this higher-elevation spot, with pine and fir sharing the canopy with oaks. I had intended to focus on the oaks in hopes of finding the rare Brachys apachei, but I was immediately distracted by large flowering shrubs that turned out to be Ceanothus integerrimus (deerbrush ceanothus) and from which I beat a diversity of miscellaneous beetles and one treehopper (but no buprestids).

Ceanothus integerrimus (deerbrush ceanothus) in montane oak-pine woodland.

I then turned my attention to beating the abundant stands of Quercus hypoleucoides (silverleaf oak) in earnest. Norm got one Brachys floccosus—a very good find, but I collected only a clerid, a few treehoppers, and one very large and very gravid Judolia instabilis. Occasional lycids turned up in sweeps and in flight, and a single clerid was collected off of one of the few Quercus arizonica (Arizona white oak) that I beat. We had hoped to go even higher to Rustler Park, but the time was getting away from us so we headed back down the canyon to eat something before meeting up with Steve for blacklighting.

Judolia instabilis gravid female.

E side Chiricahua Mountains
1.2 mi NW Jct Rd 42 & 42A

Cochise Co., Arizona
We saw this area along Cave Creek with lush-looking areas near the water’s edge that looked promising for Taphrocerus. Norm swept the areas upstream from where we parked, and I swept the areas downstream—Norm finding a single Taphrocerus (either T. chevrolati or T. sulcifrons) but me sweeping only a single hispine leaf beetle.

Sceloporus jarrovii (Yarrow’s spiny lizard) in montane oak woodland.

On the way back down Cave Creek Canyon to Portal, we passed the property of the Cazier family—originally occupied by Mont Cazier, first director of the American Museum of Natural History’s nearby Southwestern Research Station.

The family of Mont Cazier, first Director of the nearby Southwestern Research Station, still owns his former residence in Cave Creek Canyon.

E side Chiricahua Mountains
Herb Martyr Campground

Cochise Co., Arizona
Steve and I agreed that blacklighting at lower elevations would be more productive and decided to try a campground that Steve has done a few times near the Southwestern Research Station. Unfortunately, when we arrived it was already occupied (unusual, according to Steve), so we took the road the rest of the way to Herb Martyr Campground where Norm and I had collected earlier in the day. Fortunately it was vacant, so I set up my UV/MV lights in the spot nearest the parking lot while Steve set his up about 220 feet down the trail. Things looked promising when very early in the evening a male Prionus heroicus came to my lights, but that would be the only longhorned beetle we would see that night! Nevertheless, I picked up a diversity of other insects—especially cryptocephaline leaf beetles (including the smartly-dressed Griburius montezuma) and cyrtolobine treehoppers—to avoid having to consider the evening a waste. When the hoped for “9:30 cerambycid flush” did not materialize, we took down the lights and made the 2-hour drive back to Hereford.

Blacklighting with Steve Lingafelter.
Griburius montezuma at ultraviolet/mercury vapor lights in montane oak-pine woodland.

Day 18 – Huachuca Mountains (again!)

E side Huachuca Mountains
Lower Carr Canyon

Cochise Co., Arizona
This is another of Norm’s regular spots, where he has collected a variety of Agrilus and Brachys from oaks over the years. After seeing the success that he has had during the past two days by using a very long-handled, large-rim aerial net to sweep the foliage in the higher canopy of the trees, I decided to give the method a try myself (fortunately, I already have and had brought with me such a net) and see how it compared with my standard approach of beating (which reaches the lower branches only). It was not a good day to make the comparison, as there were very few beetles to be found. I focused on sweeping Quercus arizonica (Arizona white oak) while Norm preferentially swept Quercus hypoleucoides (silverleaf oak), and both of us collected but a single buprestid—mine being the not uncommon Agrilaxia arizonae but Norm’s turning out to be the very rare Mastogenius puncticollis! Otherwise on the oak I collected only a smattering of leaf beetles. I also swept Platanus wrightii (Arizona sycamore) and collected another A. arizonae and a small eumolpine leaf beetle. With our luck running dry in the lower elevations of the canyon, we decided to go up to a higher elevation site for hopefully better luck.

“Beta-testing” the long-handled, large-rim sweep net method for sampling higher in the canopy.

E side Huachuca Mountains
Reef Townsite Campgeound

Cochise Co., Arizona
I first came to this spot during last year’s trip, and while I didn’t collect many insects I did bring a lot of infested wood back home to put in the emergence boxes. That wood (both oak and pine) has been kicking out sawdust ever since, and I am hopeful that I will end up rearing series of some nice species as a result. Again, there is a lot of pine at this higher elevation, but Norm and I focused on Quercus hypoleucoides (silverleaf oak) and Q. arizonica (Arizona white oak), respectively, in an effort to find Agrilus and Brachys. Pickings were slim, but I managed to sweep single specimens of Brachys floccosus and Brachys cephalicus from the latter tree, along with a smattering of other insects (mostly leaf beetles and weevils) while Norm collected a few Agrilus from the former. There was a large, recently wind-thrown silverleaf oak in the campground that we looked at starting out hoping to see Chrysobothris running on the trunk, but none were seen. Norm went back to check periodically, however, and got a Chrysobothris costifrons on the trunk (which he gave to me). I was happy to receive the specimen, although I would have liked to have seen the beetle come to the tree since I have yet to collect the species myself. Despite the few insects collected, I was quite happy with the day, as Brachys floccosus is a very uncommon species that I have not collected previously, and I no doubt would not have collected it had I not been using the long-handled, large-rim aerial net to gain access to the higher branches in the canopy. The technique certainly warrants far more use than I have been giving it.

Quercus arizonica (Arizona white oak).

E side Huachuca Mountains
Upper Miller Canyon Rd

Cochise Co., Arizona
On the way back to Hereford, we stopped off at Norm’s “Taphrocerus spot” near the upper end of Miller Canyon Rd. Norm has collected not only T. chevrolati and T. sulcifrons by sweeping the small patch of sedges at this spot (species I have previously collected at a spot lower down in the canyon during my first visit to Arizona way back in 1987), but also T. leoni—a Mexican species heretofore not formally recorded from the U.S. I have tried, without success, to collect Taphrocerus from the lower spot on several subsequent visits, so I was hopeful that being here earlier in the season would result in better success. It happened quickly! We each got a few specimens, including T. leoni (distinctive in the field due to its shiny appearance, larger size, and distinct pubescent maculations), by sweeping the isolated plants on the north side of the creek bed and more specimens by sweeping the patch of mixed sedges, rushes, and grasses on the south side of the creek bed. Later examination of the specimens under the microscope revealed that all three species were represented—success! Nearby, there were a few Quercus arizonica (Arizona white oak) that had been trimmed a few years ago and were generating vigorous resprouts, off which Norm swept a few Agrilus abditus. This is another species that I have not encountered previously, and Norm generously gave the specimens to me. As this was my last day staying with Norm and Steve and we’d planned to go out for a nice dinner, we called it a day and headed back to the house. Still, despite the few number of specimens collected on the day, I could not be disappointed considering they represented eight species of buprestids—four of which I’d never collected before!

Miller Peak from upper Miller Canyon Rd.

We closed out my week’s visit with Norm and Steve with a tasty dinner at Pizzeria Mimosa, a glass of fine cognac, and the persistent affections of their dog Noxy!

Me, Norm Woodley, and Steve Lingafelter at “Pizzeria Mimosa.”
Celebrating a successful week of collecting with a fine cognac!
Enduring the affections of “Noxy”!

Day 19 – Apache Junction area

Superstition Mountains
Needle Vista Viewpoint

Maricopa Co., Arizona
After bidding adieu to Norm and Steve, I drove north to the Superstition Mountains to meet up with Paul Kaufman for a day of collecting and reconnecting. I first met Paul many years ago when he lived in Missouri and contacted me after collecting Saperda fayi—a very uncommon longhorned beetle that, at the time, had not yet been collected in Missouri. Paul and I spent time in the field a couple of times after that—once in southeast Missouri and another time after he moved to Farmington, New Mexico, and for many years afterwards Paul continued to send me beetles that he’d collected for identification. It was good to see him again after so many years, and I enjoyed chatting as we roamed the mountainous desert looking for beetles.

Paul had arrived first and, upon seeing how dry it was (and had been for a very long time), was not optimistic about our chances of success on the day. Nevertheless, I got out the long-handled net to see what we might find on the upper branches of the mesquites and acacias that dotted the landscape. We quickly became a little more optimistic when a Chrysobothris octocola ended up in my net with the very first tree that I swept, and over the next couple of hours I swept a variety of buprestids from Prosopis glandulosa (mesquite) and Vachellia constricta (whitethorn acacia)—including five species of Chrysobothris (C. knulli and C. merkelii both new species for the trip) and a nice series of Acmaeoderopsis sp.

For his part, Paul was surprised at the number of specimens that I managed to collect compared to what his expectations were starting out, and I have to give the credit to my use of the long-handled net, which I now firmly believe is a superior collecting method for buprestids compared to the beating sheet (at least in certain situations). The net bag essentially takes the place of the beating sheet, but since it is deeper the beetles are much less likely to escape like they can when they land on the sheet. Rather than hitting the branch from the top, the net bag is placed over the whole branch tip and gently shaken or placed under the branch and the rim tapped against the branch from below. There is less disturbance to neighboring branches if done carefully, and as a result the entire tree can be sampled in the same amount of time that is required to sample only the lower branches using a beating sheet, making it much more efficient. I estimate that on average I collected about twice as many specimens with this technique compared to beating, plus the ability to get into the upper canopy allowed me to capture some species that I would not have encountered by beating only. I am looking forward to making greater use of this technique in other areas and habitats.

Superstition Mountains
1 mi NE Tortilla Flat

Maricopa Co., Arizona
Having worked the area at Needle Vista Viewpoint sufficiently, we went to another nearby spot in the Superstition Mountains recommended by Norm and Steve.

Late afternoon sun over Mesquite Creek near Tortilla Flat.

The trees were quite a bit smaller at this location, and temps were starting to drop as we were later in the day, so I opted for the beating sheet instead of the long-handled net. Almost immediately I beat a single Chrysobothris knulli off of Vachellia constricta (whitethorn acacia), but further beating produced only a series of clytrine leaf beetles. As I was working the tree, I noticed an herbaceous plant clump below that showed evidence of feeding on the leaves, and unfurling the damaged leaves revealed numerous Microrhopala rubrolineata on what turned out to be Ambrosia ambrosioides (canyon ragweed). Interestingly, on many of the leaves with M. rubrolineata, there was also a small chlamasine leaf beetle (possibly Exema sp.) cohabiting the leaf (see photo).

Ambrosia ambrosioides (canyon ragweed).
Microrhopala rubrolineata (and a single Exema? sp.) on Ambrosia ambrosioides (canyon ragweed).

In a small area, we encountered a “hot spot” of buprestids—first I beat Acmaeodera pubiventris lanata from a dead branch of Prosopis glandulosa (mesquite). After I put the specimen in the vial, I noticed an Acmaeoderopsis junki sitting on the sheet that I’d overlooked—it was so pulverescent that I almost didn’t see it! Over the next half hour or so, I/we beat several buprestids from the mesquites in that small area, including Chrysobothris merkelii, C. octocola, and C. rossi along with a few other miscellaneous beetles. Paul also collected a couple of Agrilus (possibly A. felix) from Senegalia greggii (catclaw acacia), but my beatings from the plants produced only a few miscellaneous beetles. Finally, I found Quercus turbinella (turbinella oak, shrub live oak, gray oak—sometimes considered a subspecies of Q. dumosa), one of the few oaks that occur in lower elevation desert chaparral habitats, but beating its dense branches yielded only a single leaf beetle.

Acmaeodera pubiventris lanata, beaten from dead branch of Prosopis glandulosa (mesquite).
Acmaeoderopsis junki, beaten from dead branch of Prosopis glandulosa (mesquite).

By this time, it was getting late in the day and I wanted to setup blacklights back at Needle Vista Viewpoint, so Paul and I said our goodbyes before he headed back home and I headed for Needle Vista. On the way there, I stopped at a couple of scenic turnouts and enjoyed spectacular evening and sunset views to the west.

Late evening vista from Apache Trail above Needle Vista Viewpoint.
Late evening vista from Apache Trail above Needle Vista Viewpoint.
Late evening vista from Apache Trail above Needle Vista Viewpoint.
Sunset vista from Apache Trail above Needle Vista Viewpoint.
Sunset vista from Apache Trail above Needle Vista Viewpoint.
Sunset vista from Apache Trail above Needle Vista Viewpoint.
Me enjoying a sunset vista from Apache Trail above Needle Vista Viewpoint.

Superstition Mountains
Needle Vista Viewpoint

Maricopa Co., Arizona
I returned to Needle Vista Viewpoint with just enough time to set up the UV/MV lights and enjoy a nice brew while darkness settled. High temps and low wind held the promise of a good night, and I was encouraged by the number and diversity of insects that began to flock to the lights as the last vestiges of sunlight silhouetted the mountains behind me and a stunning crescent moon blazed over them. Cryptocephaline leaf beetles came to the lights in numbers, and eventually the longhorned beetles started coming—sporadically at first, and then regularly once the “9:30 ‘bycid flight” began. Most of the longhorned beetles were elaphidiines, presumably species of Anelaphus, and around 10:00, just as suddenly as it had begun, the ‘bycid flight ended. It was an appropriately successful last night of blacklighting for my last night in Arizona and the beginning of the long trip back home starting the next morning—with a planned detour into western Oklahoma before finally heading home.

Enjoying a brew while the blacklights hum.
A crescent moon blazes above a mountain silhouette at nightall.
Creeping nightfall.
Sinking moon.
Insara elegans (elegant bush katydid) at ultraviolet/mercury vapor light.

Day 20 – Travel Day (Phoenix to Boise City)

Today’s plan was to drive from Phoenix, Arizona to the area around Black Mesa, Oklahoma in hopes that I would arrive in time to setup blacklights in the area. That plan did not work out, as the distance was just a bit too far (going from Arizona Standard Time to Central Daylight Time also robbed me of an additional two hours!). My route took me through northeastern Arizona and northern New Mexico on roads that I’ve not previously traveled, so I at least got a good look at some parts of the country that I haven’t seen before. It was well after dark by the time I reached the western panhandle of Oklahoma, so blacklighting was not an option. Instead, I headed straight for the motel and hoped for success in the morning.

“I did that!”

Day 21 – Oklahoma Panhandle

Black Mesa State Park
Cimarron Co., Oklahoma
I’ve been to this place twice before, and both times my efforts to collect were thwarted—first by dry conditions during another early June trip in 2013, and then again just four weeks ago by cold, rainy conditions. Still, I can’t help but feel that this area has a lot of potential—if I can just get the timing right. The rainout during my last trip made me think now would be the time, as it was super dry but receiving plenty of moisture while I was there. My only doubt was whether four weeks afterwards would be soon enough or if everything would have already happened and the place had dried out again before I got there. As it turns out, I believe that I may have still been too early (more on my reasoning for that later). I wanted to access the small canyon on the north side of the park, where soapberries and hackberries in the craggy rocks promised to yield a diversity of buprestids associated with those plants, and parked at the Scenic Overlook to hike down into the canyon. Puzzlingly, I did not get anything on either plant, save for a single clytrine leaf beetle on Celtis reticulata (net-veined hackberry), many of which were still pushing out new foliage. Sweeping the soapberrys, all still pre-bloom, was also fruitless, and even the few flowers that I found failed to produce any buprestids. At least other people will be happy—I collected a few dasytine beetles for Matt Gimmel and a couple of bees for Mike Arduser on flowers of Berlandiera lyrata (lyreleaf greeneyes). Hiking back up above of the canyon, I noticed a lone Prosopis glandulosa (mesquite) in full leaf and flower—a last chance for buprestids—but collected only a few more clytrine leaf beetles. It was a rare “buprestid strike out” for the trip, and I was beginning to think once again I would fail in my effort to unlock the potential of this area in the extreme northwestern corner of extreme northwestern Oklahoma. The day was young, however, and I decided to try my luck at another spot outside the park near Black Mesa Preserve where rocky outcroppings feature stands of oak—unusual in this area and always a potentially good host for buprestids.

Sapindus drummondii (soapberry) and Celtis reticulata (net-veined hackberry) dominate the rocky bluffs at Black Mesa State Park.

1.6 mi E Kenton on Hwy 325
Cimarron Co., Oklahoma
I’ve also been to this spot a couple of times before—first during the aforementioned dry-as-a-bone June visit, and again the following year during late June. On that latter visit, I caught a number of Prionus heroicus (then a new state record for Oklahoma) and found buprestid-infested oak branches that I brought back for rearing (sadly, nothing emerged). Otherwise, I did not find much else going on, giving the visit a “too late” feel. This time started out much the same, as I started beating the oaks (ID’d on iNaturalist as Quercus × undulata, or wavyleaf oak). No insects were found at first, but I noticed a small tree that looked recently dead and encountered a chrysobothrine larva shortly after I began splitting branches. I went back to the car to get the hand saw and cut up the branches to bring back for rearing—hopefully I will have better luck this time rearing adults from the wood. I worked a number of oaks on the outcropping without finding anything, noting that many of them were just beginning to push new leaves, before crossing the highway and beating a single leaf beetle from a lone tree in full leaf.

Oak-colonized rock outcropping.

By now I was losing interest in the spot, but I’d seen a few mesquites further down the highway and thought I should at least give them a try. Again, nothing but a few leaf beetles and lots of leaffooted bugs (probably Mozena obtusa), so I finally accepted defeat—it seemed that I was again “too early”—and began working my way back to the car. At least I had the promise of buprestids from the wood I was bringing back for rearing. Rather than retracing my steps, however, I decided to walk the 2-track around the back side of the outcropping back to the car. About halfway up the road I saw an impressive Efferia sp. robber fly and, with not much left to do, occupied myself with trying to photograph it despite its repeated loping flits away from me.

Efferia sp. male (family Asilidae).
Efferia sp. male (family Asilidae).
Efferia sp. male (family Asilidae).

Resuming my trek back to the car, I then noticed Thelesperma megapotamicum (rayless greethread) flowers—with an Acmaeodera mixta adult sitting on one of them! Not that this species is at all uncommon, but if this species was at flowers then perhaps other species were as well. I’d seen precious few flowers to this point, so I began looking intently to make sure I wasn’t walking by any (Thelesperma can be easy to overlook due to its lack of ray flowers). I didn’t see any more flowers until I got back to the car, and there, not five feet from the car, were a few small Xanthisma spinulosum (spiny goldenweed)—each with one or two Acmaeodera sp. prob. neglecta. Finally, buprestids!

Thelesperma megapotamicum (rayless greethread).
Xanthisma spinulosum (spiny goldenweed).

I put my plans to leave on hold, set out some white bowl traps along the 2-track, and began searching for other flowers there and in areas adjacent to the highway. I would end up spending several more hours at the spot, finding additional Thelesperma and Xanthisma plants with the aforementioned buprestids on the flowers and also some small black individuals that could either be immaculate forms of the A. neglecta-like species or a different species altogether. Also in the same area, I found a single flowering plant of Calylophus lavandulifolius (lavender leaf sundrops) that, after visiting several times, produced several Acmaeodera (including one individual that I don’t recognize—larger and broader than A. neglecta and with uniform vittae rather than irregular spots)—and a single flowering individual of Senecio flaccidus (threadleaf groundsel) with a few A. mixta.

Calylophus lavandulifolius (lavender leaf sundrops).
Senecio flaccidus (threadleaf groundsel).

In the flats above the outcroppings, I noticed Echinocereus v. viridiflorus (green-flower hedgehog cactus) now in bloom (in contrast to earlier in the day at Black Mesa State Park). Most of the flowers on the plants were swarming with dasytine beetles, but a lone plant with a single flower that lacked dasytines had one A. neglecta-like buprestid in the flower. At some point while making the rounds between the flowers, I noticed an A. mixta in flight and successfully netted it.

Echinocereus v. viridiflorus (green-flower hedgehog cactus).

Once I’d monitored all the flowers in the area to my heart’s content, I picked up the bowl traps—all of which contained numerous Acmaeodera of multiple species (so happy that I saw this method work, first from Mike during last month’s Oklahoma trip, then earlier on this trip by accident in Mescalero Sands).

White bowl trap with several Acmaeodera spp.

My impression now, seeing how many of the trees were still pushing out leaves and that buprestid activity was limited to species of Acmaeodera, is that early June is still early season for this area (at least this year), and the timing of the season probably depends much more on the timing of rains—more similar to the western U.S.—than on the calendar and temperatures as in the eastern U.S. As a final effort to maximize my haul, I cut up some recently-cut branches of Juniperus monosperma (one-seed juniper) that I’d noticed earlier, finding a small cerambycid larva (probably Callidium sp.) under the bark when I cut into it. With both batches of wood cut up, bundled, and loaded into the car, it was finally time to make the final push home—or so I thought…

6 mi E Harmon
Ellis Co., Oklahoma
As I was leaving the Black Mesa area, I remembered a spot in Ellis Co. where I’d looked for Brachys barberi during last month’s western Oklahoma trip with Mike Arduser. Even though I was not successful in finding the species during that trip, I did collect a series of Agrilaxia texana—represented in my cabinet at the time by just two specimens collected decades ago—and one Elytroleptus floridanus—represented previously in my cabinet by just a single specimen and with this most recent collection representing a significant northwestern range extension and new host record for the species. The spot would not be too far out of the way, and since I would need to spend another night on the road anyway another attempt after allowing the season to progress a bit more might be worthwhile. Still, it would be a three-and-a-half-hour drive, which would get me there less than an hour before dusk. When I arrived (around 8:00 p.m.), I was happy to see the stands of Quercus havardii (shin oak) we’re putting out fresh foliage—something they were not yet doing on my previous visit (fresh oak foliage = Brachys!). I began sweeping the stands nearest the car and quickly came up with more A. texana along with a variety of miscellaneous beetles (mostly leaf beetles and weevils), but it wasn’t until I started sweeping stands with larger plants and the copse of tree-like plants that I finally found a couple of B. barberi. There wasn’t much time left to sweep other stands, as by then it was getting too dark to see into the net. I was, however, able to see several longhorned beetles that had bedded down on flowers of Helianthus petiolaris (prairie sunflower) growing near the car and along the road, including Batyle ignicollis, B. suturalis, and Strangalia sexnotata. It wasn’t long before dusk had turned to darkness, and it truly was time to call it a day and officially bring to an end the collecting activities for the trip after 21 days. With nearly 600 miles still to go, tomorrow will be strictly travel with no detours… for a change.

Helianthus petiolaris (prairie sunflower) with Strangalia sexnotata and a mating pair of Batyle suturalis bedded down for the night.
Everything’s bigger in Texas… and western Oklahoma!

Day 22 – Travel Day (Woodward to St. Louis)

There is nothing sadder than the final travel day home after a long collecting trip—especially one as successful as this one. Nevertheless, I looked forward to starting the day with coffee from my favorite “creationist” coffee shop (which Mike and I discovered on our prior trip to western Oklahoma just a few weeks earlier). I’m not normally one to patronize such overtly evangelical businesses, but I had to admit they served a good cup of Joe. Sadly, a sign on the door read “Permanently Closed”—I suppose proselytizing and sipping Joe just don’t mix.

Sad to find my favorite “creationist” coffee shop has closed permanently!

Arriving home later that evening closed out my longest ever driving trip—5,181 miles! Had Norm and Steve not been kind enough to do the driving while I stayed with them during the final week of my trip, the number of miles would have been even higher.

Final stats for the trip—5,181 miles with nearly 100 hours of drive time (= 53.5 mph average).

Following is a preliminary checklist of the Buprestidae collected during the trip—66 species in all! This number surely will increase once I mount and examine all the specimens, since only the species that I recognize with some degree of confidence are listed, but it already exceeds the number of species collected on any other trip I’ve made, at least in the U.S. (trips to South Africa and Mexico probably well exceed this number). Interestingly, of the 66 species collected, 17 are species I’ve never collected (five also being completely new to my collection), despite having already made several trips to many of the areas I visited. This speaks to the importance of repeated visits to the field, especially at different times of the season and in different years, continual refinement of collecting techniques and strategies, and taking advantage of opportunities to learn from others. I’m already planning next year’s trip out west, which will surely involve different timing and multiple field companions.


Preliminary List of Buprestidae Collected 17 May to 6 June 2022 in Illinois, Missouri, Texas, New Mexico, Arizona, and Oklahoma
*species not previously collected
**species new to my collection

  • Acmaeodera
    • A. bowditchi
    • A. decipiens
    • A. immaculata
    • A. gibbula
    • A. gillespiensis
    • A. horni**
    • A. immaculata
    • A. miliaris
    • A. mixta
    • A. neglecta
    • A. neoneglecta
    • A. obtusa
    • A. opuntiae*
    • A. ornata
    • A. ornatoides
    • A. paradisjuncta
    • A. princeps
    • A. pubiventris lanata
    • A. quadrivittatoides
    • A. recticollis*
    • A. riograndei*
    • A. starrae
    • A. tubulus
    • A. variegata*
    • A. n. sp.
  • Acmaeoderopsis
    • A. hulli
    • A. junki
  • Actenodes
    • A. mendax
  • Agrilus
    • A. abditus*
    • A. addendus*
    • A. albocomus
    • A. chiricahuae
    • A. egenus
    • A. lautuellus
    • A. fisherianus*
    • A. howdeni*
    • A. lacustris
    • A. lecontei celticola
    • A. obtusus
    • A. ornatulus
    • A. pectoralis
    • A. quercus*
    • A. sapindi
  • Agrilus (Engyaulus)
    • A. (E.) hespenheidei
  • Agrilaxia
    • A. arizonae
    • A. texana
  • Anthaxia (Haplanthaxia)
    • A. (H.) caryae*
  • Aphanisticus
    • A. cochinchinae seminulum
  • Brachys
    • B. barberi
    • B. cephalicus**
    • B. floccosus*
    • B. querci
  • Chrysobothris
    • C. analis
    • C. axillaris
    • C. costifrons*
    • C. knulli
    • C. lateralis
    • C. merkelii
    • C. mescalero**
    • C. octocola
  • Knowltonia
    • K. atrifasciata**
  • Spectralia
    • S. robusta
  • Taphrocerus
    • T. chevrolati
    • T. leoni**
    • T. sulcifrons
  • Texania
    • T. campestris

©️ Ted C. MacRae 2022

2022 Oklahoma Insect Collecting Trip iReport

Welcome to the 11th “Collecting Trip iReport”; this one covering a very short (4 days) trip to northwestern Oklahoma on May 3–7, 2022. My collecting partner for this trip was long-time friend and hymenopterist Mike Arduser. Mike is one of the best natural historians that I know and, like me, has a special love for the often overlooked beauty of western Oklahoma and its fascinating insect fauna. It had been 13 years, however—too long, in my opinion, since our last joint field trip when we sampled the bee (Mike) and beetle (me) fauna at The Nature Conservancy’s Four Canyon Preserve in Ellis Co. Thus, I was happy for the chance to once again spend some time in the field with such a knowledgeable naturalist in an area we that both know and love.

As with all previous “iReports” in this series, this report is illustrated exclusively with iPhone photographs (thus the term “iReport”). Previous iReports in this series include:
2013 Oklahoma
2013 Great Basin
2014 Great Plains
2015 Texas
2018 New Mexico/Texas
2018 Arizona
2019 Arkansas/Oklahoma
2019 Arizona/California
2021 West Texas
2021 Texas/New Mexico/Arizona


Day 1 – Gloss Mountain State Park (Major Co.)
It took most of the day to get here—Tulsa threw us a couple of obstacles in the form of a construction-mediated wrong turn and a motorcycle engulfed in flames. I’ve been to Gloss Mountain a number of times, but never this early in the season. Skies were sunny (unlike St. Louis when we left this morning), but temps didn’t get much above 60°F and even dropped down into the upper 50s before we finished up at sunset.

Gloss Mountain State Park – early evening view from atop the mesa.

Surprisingly, despite the earliness of the season and cool temps, beating was quite productive. Working the low areas around the parking lot, I beat a fair number and diversity of beetles and hemipterans—mostly chrysomelids—but only a single Agrilus sp. off of Prosopis glandulosa.

This must be Oenothera macrocarpa (bigfruit evening primrose, Ozark sundrop, Missouri evening primrose), though it looks very different from populations south of St. Louis.

I knew there were other trees, principally Celtis reticulata (net-veined hackberry) and Sapindus drummondii (soapberry), on top of the mesa and wanted to see if anything was on them. Bingo! Even before reaching the top, I beat a few Agrilus (several spp.) from the Celtis, and up on top I beat quite a few more off the same. There were also additional mesquite trees up top, off which I again beat a single Agrilus sp. along with a few other things, notably a series of ceresine treehoppers. The Sapindus was just starting to leaf out, and I found nothing by beating them other than a single ceresine. A notable find was the pile of larval frass of Plinthocoelium suaveolens (bumelia borer) at the base of a living Sideroxylon lanuginosum (gum bumelia) tree—a sure sign of active infestation by a beetle I have yet to formally record from this place.

Frass at the base of a small Sideroxylon lanuginosum (gum bumelia)—evidence of an active larval infestation by Plinthocoelium suaveolens (bumelia borer).

On the way back down from the top, we hit the sunset perfectly as it “touched” a peak in the foreground! Despite my success here this evening, Mike saw no bees of interest on the few flowers that were found due to the cold temps and chilling winds, so tomorrow we will continue west hoping for warmer conditions on the western edge of Oklahoma.

Dramatic sunset “landing” on a small foreground peak.
The opportunity lasted for only a minute!

Back in town, we searched for an open sit-down restaurant—fruitlessly because of the late hour—and ended up with a mediocre breakfast burrito from a fast food shop I’ve never been to before. The local Buick dealership, however, with its 1950s neon lights shining brightly in the night sky, was a taste of Americana that makes these trips so enjoyable. Life on the road!

Jensen’s Buick, Fairview, Oklahoma.

Day 2 – Black Mesa State Park (Cimmaron Co.)
Welp! We awoke this morning to cold temps (low 60s), thick fog, and low hanging clouds, and the forecast for the area showed essentially no improvement through at least the day. Our plan had been to hit a spot about an hour southwest before heading back north to Beaver Dunes State Park, but the forecast for both those areas also was cold and wet. It was not until we looked at the forecast for Black Mesa—our last planned stop of the trip and a 4½-hour-drive to the west—that the forecast seemed to be in our favor, so we decided to blast on out there. We figured we would get there at about 2:00 pm and could spend the rest of the day there collecting, camp there tonight, and start heading back east tomorrow (assuming the forecast improved for the areas we missed).

Pronghorn antelope (Antilocapra americana).

Wrong! When we got there, it was not only cloudy and cold, but dry as a bone! Even if it had been sunny with warmer temps, there still would not have been any insect activity to speak of. The leaves of oaks and hackberries in the area were just barely starting to break bud, and the only flowers we saw at the park were a large willow in full bloom—but not a single insect visiting them. Knowing that there was no other place where conditions were better that we could drive to within the next couple of hours and collect for at least a short time, we instead decided to make it a hiking day and hike the High Point Trail at nearby Black Mesa Nature Preserve.

Black Mesa Nature Preserve (Cimmaron Co.)
When we arrived and looked at the signage, we learned that the hike to the oracle at the official high point would be a more than 8-mile hike! Just reaching the top of the mesa itself would be a more than 3-mile-hike, with the high point another mile on top. Not knowing if we had the appetite for such a distance (or time to do it before sunset) and with the wind cold and biting, we started out anyway and gave ourselves permission to turn around at any point if we felt like it.

View along High Point Trail.
View along High Point Trail.
Berlandiera lyrata (lyreleaf greeneyes).

Nevertheless, we persevered. We checked the cholla (Cylindropuntia imbricata) along the way hoping to see Coenopoeus palmeri (one of the cactus longhorns, which I’m not sure has been recorded from Oklahoma) or at least one of the more widespread Moneilema species, but none were seen (nor really expected). The trail up the side of the mesa was steep and spectacular, and the trail atop the mesa was surreal—especially given the cold winds and low-hanging clouds. Eventually, we made it to the official high point and enjoyed the fun facts carved into each side of the granite obolisk marking the spot.

View along High Point Trail about halfway up the climb.
Juniperus monosperma (one-seed juniper).
Oklahoma High Point obolisk.
Oklahoma High Point marker.
Mike (right) and me at Oklahoma High Point.

Coming back down was not much easier than going up, the steepness of the trail jamming my toes into the toe box of my new hiking boots (which performed admirably!), but I did find an insect—a largish black weevil torpidly crawling on the trail. Even on the relatively level lower portion of the trail once we got there was difficult, our legs really starting to feel the miles now. As we hiked the last mile back to the car, the temperature continued to plummet as it started to sprinkle, turning to rain soon after we reached the car and then heavy rain as we headed down the highway back to the east. The irony of the situation—rain coming to a parched landscape just when we are ready to leave—did not escape us. We’ll spend the night in Boise City and hope for a better forecast tomorrow!

Descending the upper slope on the High Point Trail.

Day 3 – Beaver Dunes State Park (Beaver Co.)
Temps were down in the mid-40s when we awoke this morning, but skies were sunny and we were heartened by a promising forecast of continued sun and highs in the low to mid-60s. Our first destination—Beaver Dunes—was a relatively short 2-hour drive further east, and when we arrived sunny skies still prevailed. Unfortunately, temps still hovered in the mid-50s with a biting wind that made using the beating sheet difficult to impossible.

Dunes at Beaver Dunes State Park.

That said, I managed to beat a fair series of Agrilus spp. (probably mostly one species) and a few other beetles off living Celtis reticulata (net-veined hackberry) dotting the roadside along the entrance to the Picnic Area. Under the main group of hackberries I noticed new growth of Cucurbita foetedissima (buffalo gourd) along with last year’s dead stems. I’ve never collected Dorcasta cinerea (a longhorn beetle that utilizes buffalo gourd as a larval host), so I began splitting open the old stems to see if I could find unemerged adults. I didn’t, but what I did note inside the stems was evidence of boring by some insects and, eventually, the tiniest little scolytine bark beetles that I’ve ever seen. They were always found right at the node, usually in pairs (perhaps male and female?), and I ended up collecting a series of about a dozen specimens from two different stems.

Tradescantia occidentalis (prairie spiderwort, western spiderwort).

Also in the main group of hackberries, I noticed a dead branch hanging from the tree, which had fallen but gotten snagged on a lower branch to remain off the ground. The branch was obviously infested and showed a few emergence holes indicative of both buprestids and cerambycids, and when I broke into it I found two unemerged adult Agrilus (different species), which caused me to cut and bundle the branch to being back for rearing. At the entrance, I went to examine the stand of yellow flowers that greeted our arrival, determining them to be Pyrrhopappus pauciflorus (smallflower desert-chicory, Texas false dandelion). While I was on the ground photographing the flowers, I noticed a red and black hister beetle that proved to be Margarinotus bipustulatus—aptly named considering the two red maculations on the elytra. I also noticed a couple of tiger beetle larval burrows in the hard-packed sandy soil and found a long, thin plant stem to “fish” the larvae out. I managed to snag the larva in one of the burrows, which I believe is Tetracha carolina (Carolina metallic tiger beetle) by virtue of the thin white margin around the prothorax and the open habitat in which the larval burrow occurred. If this is true, then it is a second instar because it is slightly smaller than a typical Cicindela sp. third-instar larva.

Pyrrhopappus pauciflorus (smallflower desert-chicory, Texas false dandelion).
Pyrrhopappus pauciflorus (smallflower desert-chicory, Texas false dandelion).
Margarinotus bipustulatus (family Histeridae).

Afterwards, I went over to the dunes to see if Mike had found anything, but temps were still too cold to see anything flying. He did, however, show me an interesting stand of Penstemon that he’d found and that we determined to be P. fendleri (Fendler’s penstemon). The plants were all on the north side of the dune in apparently protected spots, and I noted that on iNaturalist our observation was the northernmost record for the species (save one suspicious, disjunct Colorado record).

Penstemon fendleri (Fendler’s penstemon).
Penstemon fendleri (Fendler’s penstemon).

On the way back to the car, I beat a few more beetles off living Celtis reticulata. By now, we’d seen all we needed to see here and decided to head southeast to one of the Brachys barberi locations (that were the reason for this trip in the first place).

5 mi E of Harmon (Ellis Co.)
This

Recently, another coleopterist collected Brachys barberi—more typically a southwestern species—on Quercus harvardii (shinnery oak) at this spot. I’ve not managed to find the species myself yet, and as it was collected on May 3rd last year I hoped the timing would be right. Quercus havardii dominated the landscape at this spot, mostly as thick stands of low-growing shrubs but also as a copse of small trees.

Quercus havardii (shinnery oak, shin oak, Havard oak).

At first, I swept the lowest-growing plants, collecting a variety of mostly chrysomelids and curculionids and even one Agrilus sp., before moving to beating along the sunny edges of the patches of taller shrubs and collecting similar species (but no Agrilus sp.). Just to the north, I noticed a stand of individuals tall enough to be considered trees (presumably a clonal stand) and began beating them. Immediately I began collecting not only the chrysomelids and curculionds that I was collecting before, but also several Agrilus spp. and what must be Agrilaxia texana—a species represented in my cabinet by just two specimens that I collected in northeastern Texas way back in 1984.

I worked nearly the full perimeter of the copse, noticing that most of the beetles were being collected only on the south-facing sunny (and leeward) side. When I was just about ready to call it quits, a much larger black and yellow beetle landed on the sheet. For an instant I thought it was a lycid, but it moved characteristically like a longhorned beetle, and I quickly realized that I had collected Elytroleptus floridanus—a quite rare southeastern U.S. species that I have only seen once before when I reared a single individual from dead oak that I collected in the Missouri bootheel (and representing a new state record). I wasn’t sure the species had ever been recorded from Oklahoma, so I found Gryzmala’s revision of the genus online and saw that it had been previously recorded from the state—but all the way over on the east side near the border of Arkansas. All records from Texas as well are from the eastern side of the state, so today’s capture appears to represent a significant northwestern extension of the species’ known geographic range by about 300 miles!

Sadly, I never saw Brachys barberi, but collecting Elytroleptus floridanus (in Oklahoma!) was a pretty good consolation prize.😊


Day 4 – Prologue (“Good to Go” coffee shop)
We awoke to bright sunny skies, and though a tad chilly it was still warmer than the previous mornings and with a good forecast to boot! It would take about an hour to drive to the day’s collecting spot—the one and only Gloss Mountain State Park (where we visited briefly a few days ago to start the trip), but not until after an unexpected and hilariously bizarre experience at a coffee shop in town called “Good to Go”.

“Good to Go” coffee shop lounge.

Mike was the first to notice the velociraptor in the lounge—saddled up for a ride! Okay, that’s cute. Then he noticed the sign on the outdoor display that read “Stegosaurs roamed the Earth about 5,000 years ago.” At first I thought, okay, they’re a little confused on the timeline, but what they’re trying to say is that dinosaurs lived a long time ago.

“Stegosaurs roamed the Earth about 5,000 years ago.”

Then I noticed a granite plaque in the background that clearly read “The Holy Bible”, and it dawned on me that we had entered a creationist’s den! Had we not already ordered our coffee, I might have surreptitiously tiptoed my long-haired hippy butt out of there before somebody pointed at me and began slowly chanting “Lucifer!”

Apparently this is overwhelming evidence that humans saw living dinosaurs.

Once we were outside the shop, our coffee secured and the need for hushed tones no longer muffling our reactions, we took a quick walk with the dinosaurs to admire their seeming scientific accuracy. I was impressed with the T. rex in particular, it’s body axis realistically horizontal with the tail straight and strong—not the lumbering, upright, tail-dragging version that I learned about as a kid. At least they were accepting some of the current body of scientific evidence on dinosaurs and ignoring only that dealing with their age—or so I thought…

A remarkably scientifically accurate rendition of T. rex with the more recently advocated horizontal posture.

The stegosaur as well appeared to be fairly accurately rendered, its tail also straight and strong and a youngster trailing closely behind, until I noticed something atop the adult—an angel riding it! ‘God’s creatures big and small’, I guess.

Note the angel riding the stegosaur!

The coup de grace was the information plaque behind the stegosaur. Rather than providing information on dinosaurs, I was instead treated to a barrage of hilariously unsupported claims advocating the idea that humans and dinosaurs once lived together. Each “factoid” on the plaque was more bizarre and quotable than the one before. Did you know that the adult stegosaur probably died 4,000 years ago in the Great Flood, but that the baby—happily—likely survived by getting a ride on the Ark with Noah! And all that scientific evidence that pinpoints the Cretaceous extinction to 65 million years ago? Apparently it has merely been fabricated as part of a global conspiracy because scientists just don’t want to agree with the Bible. I just about lost it, however, when I reached “It is uncertain if humans ever rode Dinosaurs, but there is overwhelming evidence that humans saw living dinosaurs.” I mean—What?!

I don’t even know where to begin!

Our unplanned morning entertainment now done, we hit the road for our next—and final—collecting spot for the trip.

Gloss Mountain State Park (Major Co.)
We arrived at about 10 am with a plan to spend the rest of the day there—whether the collecting was good or bad, this would be our final stand. We hiked up to the mesa, stopping at an accessible spot about halfway up to work the trees (me) or set out pan traps (Mike). Beating the Celtis reticulata (net-veined hackberry) yielded a similar assortment of beetles as last time—a couple of Agrilus spp. along with the occasional chrysomelid or curculionoid and a few other beetles, and the same was true with Prosopis glandulosa (mesquite), with the exception that I did not find any Agrilus this time.

Gloss Mountain State Park – view west from atop gypsum-capped red clay mesa.

Atop the mesa, I decided to do an entire perimeter hike—something I’ve always wanted to do but never actually accomplished. The idea was to beat all of the C. reticulata, P. glandulosa, and Sapindus drummondii (soapberry) that I could find in an effort to “leave no stone unturned” in my quest for beetles. Soon after starting out, I saw a nice Pasimachus elongatus ground beetle running across the mesa top and “forced” it to cooperate for photos by pinning a hind tarsus to the ground with my finger tip (barely visible in the upper left side of the photo). I collected it, as well as another that I saw a short distance away, and then proceeded with the beatings! Beating the C. reticulata was quite productive, with perhaps three Agrilus spp. and numerous other beetles being collected off of nearly every tree that I beat. Beating P. glandulosa also was productive for various beetles, though again no Agrilus were encountered. The biggest surprise came when I started beating S. drummondii, most of which were still in the earliest stages of leafing out. I got nothing from most of the trees (the majority of which were clustered in a small copse near the front of the mesa), but in the back part of the cluster were a couple of trees with noticeably more foliage—beating them yielded perhaps a dozen Agrilus limpiae, a soapberry specialist that I haven’t seen in numbers since 1986 when I collected a series on soapberry in south-central Kansas.

Gloss Mountain State Park – view west from atop gypsum-capped red clay mesa.
Pasimachus elongatus (family Carabidae) atop gypsum-capped red clay mesa.

I rarely get anything beating Sideroxylon lanuginosum (gum bumelia), but I beat most of the trees that I saw anyway and collected one cryptocephaline chrysomelid and two curculionoids. A single Eleodes hispilabris (apparently on its last leg) was seen near the north end of the mesa, which I photographed and collected, and on the way back I encountered a small patch of Sphaeralcea coccinea (scarlet globemallow) in bloom, from the flowers of which I collected a few small melyrid-type beetles and a small halictid bee for Mike. Also on the north part of the mesa I saw a young eastern collared lizard (Crotaphytus collaris), who posed just long enough for me to get off a shot before blasting away from my approaching lens.

Eleodes hispilabris (family Tenebrioindae) atop gypsum-capped red clay mesa.
Sphaeralcea coccinea (scarlet globemallow) atop gypsum-capped red clay mesa.
Crotaphytus collaris (eastern collared lizard) atop gypsum-capped red clay mesa.

Throughout the hike atop the mesa I kept my eye out for “new-to-me” plants (of which there are many), finding for the first time Toxicodendron rydbergii (western poison ivy) and blooming individuals of Chaetopappa ericoides (rose heath). Physaria gordonii (Gordon’s bladderpod)—a relative of the federally threatened P. filiformis (Missouri bladderpod)—was blooming abundantly atop the mesa. At this point, Mike and I rejoined and relayed to each other our more notable findings. For Mike’s part, he had seen a couple of cacti that I had missed—Escobaria missouriensis (Missouri foxtail cactus) and Echinocereus reichenbachii perbellus (black lace cactus)—and took me to the spots where he had seen them. While retracing our steps, we also found Gaillardia suavis (pincushion daisy, perfumeballs) and the strikingly beautiful Penstemon cobaea (cobaea beardtongue, prairie beardtongue, foxglove penstemon).

Toxicodendron rydbergii (western poison ivy) atop gypsum-capped red clay mesa.
Chaetopappa ericoides (rose heath) atop gypsum-capped red clay mesa.
Physaria gordonii (Gordon’s bladderpod) atop gypsum-capped red clay mesa.
Escobaria missouriensis (Missouri foxtail cactus) atop gypsum-capped red clay mesa.
Escobaria missouriensis (Missouri foxtail cactus) atop gypsum-capped red clay mesa.
Echinocereus reichenbachii perbellus (black lace cactus) atop gypsum-capped red clay mesa.
Echinocereus reichenbachii perbellus (black lace cactus) atop gypsum-capped red clay mesa.
Gaillardia suavis (pincushion daisy, perfumeballs) atop gypsum-capped red clay mesa.
Penstemon cobaea (cobaea beardtongue, prairie beardtongue, foxglove penstemon) atop gypsum-capped red clay mesa.
Penstemon cobaea (cobaea beardtongue, prairie beardtongue, foxglove penstemon) atop gypsum-capped red clay mesa.

By this time, I had been on the mesa top for five hours, and even though temperatures were mild (mid-70s) I desperately needed food and water. Mike, for his part, had also had a wildly successful day with bees, capturing many at the flowers and many more in the various pan traps (both in top and halfway up the slope). I descended the steep slope with its mixture of metal steps, cut rock, and wooden planks and enjoyed a quick feast of sardines and Triscuits (a decades-long bug-collecting-trip staple) washed down with Gatorade before getting back to work on the mesquite around the parking lot. I was committed to trying to find Agrilus on the plants—a single individual of which I’d beaten from the plants three days earlier, and after beating several plants and seeing none (but collecting a great number of clytrine and cryptocephaline chrysomelids along with other insects) I finally found one! I continued to work the trees and collect primarily chrysomelids, but no more Agrilus were seen. I am hopeful that it will be a southwestern species not currently known from Oklahoma—a situation I have found with several other Prosopis-associated beetles in this part of northwestern Oklahoma.

Gloss Mountain State Park – view north from atop gypsum-capped red clay mesa.
Gloss Mountain State Park – history of the name.

I hadn’t intended to work any additional Prosopis beyond the road into the parking lot, but there were a few particularly large trees along the front of the park next to the highway rest stop. The first one I beat yielded a very large cryptocephaline that I had not seen on any of the other Prosopis, so I continued beating them and collected a nice series along with a few other clytrines, pachybrachines, and curculionoids. At the furthest point west, I recalled having seen during a previous visit a western diamondback rattlesnake a bit further to the west, so I continued to the spot hoping to see another. No such luck, so I tiptoed through the tall grass back to safety and made my way back to the car to wrap up seven and a half hours of collecting on a spectacular day—sadly, the last of the trip!


Epilogue
This trip was just a warm-up. In just over one week, I will head out again—this time to western Texas and southern Arizona for sure, and maybe elsewhere depending on how things go. At three weeks, it will be the longest collecting trip I’ve done since I went to South Africa in 1999 and Ecuador 10 years before that. I’m also looking forward to meeting up with a number of other coleopterists at various points during the trip—Jason Hansen, Joshua Basham, and Tyler Hedlund in Texas, and Norm Woodley and Steve Lingafelter in Arizona. If there is time, I may stop off at a place or two in northeastern New Mexico and at Black Mesa on the way back. Look for an iReport on that trip sometime in early-mid June!

©️ Ted C. MacRae 2022

2019 Arkansas/Oklahoma Insect Collecting Trip iReport

Alternative title: Rich and Ted’s Excellent Adventure

This is the seventh “Collecting Trip iReport”; this one covering a 5-day trip to Arkansas and Oklahoma from June 7–11, 2019 with my friend and local collecting buddy Richard Thoma. As with all previous “iReports” in this series, this one too is illustrated exclusively with iPhone photographs.

Previous iReports include the following
2013 Oklahoma
2013 Great Basin
2014 Great Plains
2015 Texas
2018 New Mexico/Texas
2018 Arizona

Image may contain: 2 people, including Ted MacRae, people smiling, people standing, beard, tree and outdoor
…and so it begins!

Day 1 – Ozark National Forest, vic. Calico Rock, Arkansas
It’s been many years since I’ve visited these sandstone glades overlooking the White River near Calico Rock. Conditions were partly sunny when we arrived, but water on the ground suggested rain earlier in the day. We had only a short time to start exploring before the wind started blowing up and the smell of rain filled the air. I did manage to beat one Amniscus sexguttata from a branch of living Pinus echinata and collect a couple of Strigoderma sp. from Coreopsis lanceolata flowers before steady rain forced us to retreat.

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White River near Calico Rock, Arkansas—before the rain.
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White River near Calico Rock, Arkansas—rain’s a comin’!

Day 2 – Ouachita National Forest, Winding Stair Campground & Ouachita Trail, Oklahoma
We walked the trail from the campground S about 2½ miles and back. I started off with Acmaeodera tubulus on Krigia sp. flowers, eventually finding a lot of them on this plant at higher elevations along with a single Acmaeodera ornata, and I beat a few Agrilus cepahlicus off of Cornus drummondii. This had me thinking it would be a good buprestid day, but it wasn’t, the only other species collected being some Chrysobothris cribraria off of small dead Pinus echinata saplings and Pachyschelus laevigatus on Desmodium sp. Also beat a few miscellaneous insects off of Cercis canadensis and Vaccineum arborea and swept some from grasses and other herbaceous plants. Back at the campground I collected Chrysobothris dentipes on the sunny trunks of large, live Pinus echinata trees.

Emerald Vista, along the Talihema Scenic Drive.
Acmaeodera tubulus on flower of Rosa carolina.
Acmaeodera tubulus on flower of Rosa carolina.
Canthon sp.
Apheloria virginiensis reducta (ID by Derek Hennen).
The biggest cairn I’ve ever seen.

Ouachita National Forest, Talimena Scenic Dr at Big Cedar Vista, Oklahoma
There were lots of native wildflowers like Coreopsis tinctoria and Ratibida columnifera in bloom, so we stopped to check them out. There were lots of butterflies, however, I found only a single Typocerus zebra on Coreopsis lanceolata.

View south from Talimena Scenic Drive at Big Cedar Vista.
Coreopsis tinctoria (plains coreopsis).
Ratibida columnifera (upright prairie coneflower).

Ouachita National Forest, Winding Stair Campground, Oklahoma
We returned to the campground in the evening to do some blacklighting. I had high hopes, but only five cerambycids came to the lights, all represented by a single individual: Monochamus carolinensis, Acanthocinus obsoletus, Amniscus sexguttatus, Eutrichillus biguttatus, and Leptostylus tranversus (the first four are pine-associates). I also picked up a few other miscellaneous insects.

Rich processes the day’s catch as the blacklight hums in the background.

Day 3 – Medicine Park Primitive Campground, Oklahoma
There wasn’t much insect activity going on in eastern Oklahoma, so we drove out west to the Wichita Mountains for hopefully better luck. We found a small park with a primitive campground in the city of Medicine Park—my first thought was to beat the post oaks dotting the campground, but when I went into the native prairie between the campground and the creek I never came out! Right away I found what must be Acmaeodera ornatoides on flowers of Opuntia sp., then I found more on flowers of Gallardia pulchella along with Acmaeodera mixta. The latter were also on flowers of Thelesperma filifolium along with Acmaeodera neglecta—took a nice series of each, and I also got a few of the latter on flowers of Coreopsis grandiflora. Strangalia sexnotata were on flowers of C. tinctoria and Torilis arvensis, and then on the latter plant I saw a male Strangalia virilis—a Texas/Oklahoma specialty that I’ve never collected before! I spent the next hour looking for these guys and ended up with 3 males and 2 females along with a few Trichiotinus texanus—another Texas/Oklahoma specialty—and a single Agrilaxia sp. nr. flavimana (could be A. texana). One single Typocerus octonotatus was on flowers of Achillea millefolium. I think we may come back here tomorrow—I’d like to look for more S. virilis and beat the post oaks (the reason we stopped here to begin with).

A cacophony of native wildflowers!
An orgy of Euphoria kernii (Kern’s flower scarab) in Opuntia sp. flower. Multiple color forms exist for this species.
At first I thought this was a type of hover fly (family Syrphidae), but eventually I determined it to be Esenbeckia incisuralis, a horse fly (family Tabanidae)—incredible emerald green eyes!
Papilio polyxenes asterius (black swallowtail) caterpillar.
Echinocereus reichenbachii baileyi (lace hedgehog cactus).

Lake Lawtonka nr. Ma Ballou Point, Oklahoma
We stumbled into this area while looking for stands of Sapindus drummondii (soapberry)—found a small stand along the road, but it was too inaccessible. The same diversity of blooms were present as at the previous spot, so I picked a few longhorns off flowers of Coreopsis grandiflora and Gaillardia pulchella. Super windy, so we didn’t stay long.

View across Lake Lawtonka from Ma Ballou Point.
Neochlamisus sp. (case-bearing leaf beetle) larvae inside their “casas de caca” on Monarda fistulosa (bee balm).
I believe this is Harrisina coracina, a leaf skeletonizer in the family Zygaenidae. Both BugGuide and the Moth Photographers Group show records only from Texas.

Day 4 – Wichita Mountains National Wildlife Refuge, Oklahoma
Before starting the day’s collecting, we wanted to go into the Wichita Mountains National Wildlife Refuge to have a look around. On the way into the refuge we some American bison near the road and had to stop, take photos, and simply admire these massive, majestic beasts. We then went to the Cedar Plantation, where I had visited before back in 2012 and photographed black individuals of Cicindelidia obsoleta vulturina (prairie tiger beetle). No tiger beetles were out now (they come out in the fall), but I’d hoped to maybe see Cylindera celeripes (swift tiger beetle) along the 2-tracks in the area. No such luck—nevertheless, we saw a myriad of interesting insects, including several more Esenbeckia incisuralis (green-eyed horse flies) and a beautiful Trichodes bibalteatus (checkered beetle), the latter of which I photographed on flowers of Ratibida columnifera and Achillea millefolium with the big camera. Afterwards we visited the “prairie dog town” and got marvelous views and photographs of black-tailed prairie dogs.

Native American wildlife on a native American landscape.
American bison (Bison bison bison).
Wichita Mountains from Cedar Plantation.
Acmaeodera mixta on flower of Rudbeckia hirta (black-eyed Susan).
Typocerus octonotatus on inflorescence of Achillea millefolium (yarrow).
Strangalia sexnotatus on flower of Ratibida pinnata (gray coneflower)
Trichodes bibalteatus on flower of Rudbeckia hirta (black-eyed Susan).
Trichodes bibalteatus on flower of Ratibida pinnata (gray coneflower)
The author walks a bison trail through the Cedar Plantation.
Black-tailed prairie dog at its burrow entrance.
“Watch you lookin’ at, Willis?!”
Black-tailed prairie dog (Cynomys ludovicianus).

Medicine Park, Jack Laughter Park, Oklahoma
We’d noticed this spot yesterday because of the old post oaks and wealth of wildflowers blooming up the mountainside. There wasn’t much going on today, however—just a few Acmaeodera mixta on flowers of Gaillardia pulchella. I did find an Anthaxia (Haplanthaxia) sp. on my arm! Otherwise I spent some time photographing the landscape and some geometrid larvae on flowers of Gaillardia pulchella.

Opuntia sp. (prickly-pear cactus) amidst rocky exposures.
Small Oncoptus fasciatus (large milkweed bugs) nymphs colonize seed pods of Asclepias asperula (antelope-horns).

Medicine Park Primitive Campground, Oklahoma
We returned to this spot since we had so much luck yesterday. I was hoping to collect more Acmaeodera ornatoides and Strangalia virilis, but there was much less going on today than yesterday—basically didn’t see anything for the first hour and a half. I didn’t give up, however, and kept checking the area where we saw most of the S. virilis yesterday, and eventually I saw another male in the same area as yesterday on the same stand of Torilis arvensis. I found two more males in the same area over the next hour, so three males on the day was a good reward for the time spent looking for them. I also collected Trichodes apivorus and Trichiotinus texanus on flowers of Allium sp. Interestingly, beating the post oaks—the reason why I originally wanted to stop here—produced nothing. So, not very many specimens on the day, but happy with those I did get.

Thelesperma filifolium (stiff greenthread).
Coreopsis grandiflora (large-flowered tickseed).
Gaillardia pulchella (firewheel).
Coreopsis tinctoria (plains coreopsis).
Torilis arvensis (erect hedge parsley), introduced.
Opuntia sp. (prickly-pear cactus).
Opuntia sp. (prickly-pear cactus).
Allium sp. (wild onion).

Medicine Park, Jack Laughter Park, Oklahoma
I was pretty much done for the day after spending all morning at the refuge and most of the afternoon at the previous spot, but Rich wanted to take another look at Jack Laughter Park because he’d found some interesting grasshoppers there. As with earlier in the day there were few beetles of interest to me, but I did collect a couple of Trichiotinus texanus on flowers of Cirsium undulatum. I checked out some large post oaks with large dead branches thinking that might be what Strangalia virilis was breeding in but never saw any, and eventually I turned my attention to photographing a few interesting native plants that I found along the way.

Krameria lanceolata (trailing krameria).
Cirsium undulatum (wavyleaf thistle or gray thistle) inflorescence.

Cirsium undulatum (wavyleaf thistle or gray thistle) can be distinguished by its wavy leaves that are gray-green on both upper and lower surfaces.

Day 5 – Epilogue
We were tempted to do one last little bit of collecting on the way back to St. Louis, but since had pretty good luck during the last couple of days and the drive alone would take more than nine hours we decided to leave well enough alone and get home at a reasonable hour. A walk with Beauregard when I got home to stretch the post-drive legs was the perfect way to end the mini-vacation.


©️ Ted C. MacRae 2019

A flower visiting jewel beetle that is not an Acmaeodera

Agrilus muticus

Agrilus muticus LeConte, 1858 | Alabaster Caverns State Park, Woodward Co., Oklahoma

In North America, jewel beetles (family Buprestidae) routinely associated with flowers usually belong to the genus Acmaeodera. Often black with yellow and/or red elytral markings (e.g., A. immaculata, A. macra, A. ornataA. neglecta, A. tubulus, etc.), these beetles are especially diverse in western North America and bear a striking resemblance to stinging bees and wasps (maintaining the charade even during flight by holding their fused elytra together above the abdomen). A few other less speciose genera (e.g., Anthaxia and Agrilaxia) also regularly visit flowers, but for the most part outside of these genera encounters on flowers are at best incidental.

Agrilus muticus

Adults are associated almost exclusively with flowers of winecup (Callirhoe involucrata).

Of course, exceptions are the rule in biology, and in the genus Agrilus there are few species that are found almost exclusively on flowers. One of these, Agrilus muticus occurs in Texas, Oklahoma, and Kansas and has been found only in the flowers of winecup, Callirhoe involucrata (Fisher 1928). The species is fairly routinely encountered wherever these plants are found in bloom, and I have observed adults in these flowers on numerous occasions feeding on the petals and mating (and see this BugGuide photo of group sex!). While winecup can be presumed to be the larval host, it has never been reared from this (or any other) plant, and in fact I am not aware of anything beyond descriptions and catalogue listings that have been published for the species. I suspect the larvae tunnel within the stem base or main roots of living winecup plants (and, thus, making them more difficult to rear than species associated more typically with dead wood).

Only one other North American Agrilus has been documented routinely visiting flowers. Agrilus blandus is frequently encountered on flowers of Eriogonum, and these plants have also been confirmed as larval hosts (Nelson & Westcott 1976). None of the remaining species of Agrilus in North American are known to routinely visit flowers, and in fact the vast majority of them are associated with woody rather than herbaceous plants. The latter include A. concinnus, which breeds in the base of living Hibiscus (MacRae 2006), and A. malvastri, which is usually found on Sphaeralcea (Fisher 1928), although whether that plant also serves as the larval host remains unconfirmed. It is interesting that most of these non-woody plant hosts (CallirhoeHibiscus and Sphaeralcea, but not Eriogonum) all belong to the same plant family—Malvaceae. Plants in this family also serve as known or suspected larval hosts for several species of jewel beetles in the genus Paragrilus—a close relative of the genus Agrilus. Again, these beetles are normally found on the foliage and not on the flowers.

REFERENCE:

MacRae, T. C. 2006. Distributional and biological notes on North American Buprestidae (Coleoptera), with comments on variation in Anthaxia (Haplanthaxia) viridicornis (Say) and A. (H.) viridfrons Gory. The Pan-Pacific Entomologist 82(2):166–199 [pdf].

Nelson, G. H. & R. L. Westcott. 1976. Notes on the distribution, synonymy, and biology of Buprestidae (Coleoptera) of North America. The Coleopterists Bulletin 30(3):273–284 [pdf].

Fisher, W. S.  1928. A revision of the North American species of buprestid beetles belonging to the genus AgrilusU. S. National Museum 145, 347 pp. [Smithsonian Libraries].

© Ted C. MacRae 2015

Crypsis? Mimicry? Crypsimicry?

Continuing with the previous post’s theme on crypsis, here is an interesting insect that I photographed in north-central Oklahoma in late June 2014. I was checking standing and fallen trunks of large, dead eastern cottonwood (Populus deltoides) trees in Woods Co. near the Cimarron River, where just a few days earlier I had found a jewel beetle (family Buprestidae) that had eluded me for more than 30 years—Buprestis confluens. I had found only a single individual and returned to the spot in the hopes of finding more. As I searched the trunk of one particularly large, fallen tree—its trunk still covered with bark, I noticed movement but couldn’t make out right away what I was seeing. A closer look revealed the movement to be from a wasp-like insect, its antennae curiously quivering in a manner that reminded me of an ensign wasp (family Evaniidae). More careful looking, however, revealed the insect to be not a wasp, but a longhorned beetle (family Cerambycidae), which I then recognized to be the species Physocnemum brevilineum.

Physocnemum brevilineum

Physocnemum brevilineum (Say, 1824) on fallen cottonwood (Populus deltoides) | Woods Co., Oklahoma

This beetle is commonly referred to as the elm bark borer, a reference to the larval habit of mining within the bark of living elm trees, but as far as I can tell this beetle is anything but common. Like the Buprestis confluens that I had found a few days earlier, this was a species known to me only by pinned museum specimens (I’m always amazed when a woodboring beetle species is apparently common enough to warrant a common name and is said to reach pest status in some cases, yet eludes my net for decades!). At any rate, my impression based on these pinned specimens and published images was that the species is another of the many longhorned beetles that seem to mimic ants (Cyrtinus, CyrtophorusEudercesMolorchus, and Tilloclytus being among the others). Like many of these other mimics, the species is dark with small amounts of red and bears polished, ivory-colored ridges at mid-elytra to give the illusion of a narrow waist. After seeing a living individual, however, and especially its behavior—in particular the very wasp-like manner in which it moved its antennae, I’m not so sure that ant mimicry alone explains the appearance and behavior of the species.

Physocnemum brevilineum

A lateral view reveals the beetle, but is it trying to mimic an ant, or a wasp, or both?

Of course, there is no reason why it must be ant mimicry or wasp mimicry (or crypsis, for that matter). Evolution has no rule stating that only one survival strategy can be employed at a time, and if, as it seems to me, the beetle is utilizing both crypsis and mimicry—the first to avoid detection and, failing that, the second to give the potential predator pause, then there is no reason why the mimicry portion of its defense couldn’t be modeling both ants and wasps as a way to maximize an overall “nasty hymenopteran” appearance.

© Ted C. MacRae 2015

Just how widespread is Prionus heroicus?

On our insect collecting trip to the western Great Plains last June, Jeff Huether and I encountered several species of beetles in the genus Prionus—longhorned beetles (family Cerambycidae) known commonly as “prionid root borers”. I’ve already recounted our experiences with two of them—P. integer in southeastern Colorado and P. fissicornis in northeastern New Mexico, both of which we encountered unexpectedly and by the “bucketload” using prionic acid baited traps in shortgrass prairie habitats. The third species we encountered was also rather unexpected, so much so that we were not even sure about its identity until we examined the collected specimens more closely. Unlike the two previously mentioned species, however, the behavior of this third species—P. heroicus, especially regarding its attraction to prionic acid, was quite different, and one of the localities where we found it raises some interesting questions regarding its recorded distribution.

Prionus heroicus male

Prionus heroicus male | Mills Canyon, Harding Co., New Mexico.

Prionus heroicus is among the largest species in the genus, and we first encountered it in northeastern New Mexico while searching for suitable habitat around the small town of Mills (Harding Co.) where several grassland-associated species in the genus had been found by other collectors. While we were searching we were intrigued by the USDA Forest Service road sign pointing to “Mills Canyon” and decided to head down the gravel road to see what it looked like. A few miles down the road the endless shortgrass prairie quickly changed to sandstone canyonland with oak/pine/juniper woodland. By now it was late afternoon, and the rich, pulsing buzz of cicadas filling the air as we drove slowly through the area suggested the potential for good insect activity. We stopped to take a closer look at things, and within minutes an enormous prionid beetle came flying up to the car—apparently attracted to the prionic acid lures inside!  We netted the beetle, and shortly afterwards another beetle flew into the vicinity… and another! We hurriedly set out some lure-baited traps, and over the next hour or so we collected close to a dozen beetles. Our first thought, based on their large size and the presence of 12 antennal segments, was that they might be P. californicus. This was a bit of a letdown, as our goal was to find little known and uncommonly encountered Great Plains species, not common and widespread western species like P. californicus. However, the blackish rather than reddish color and relatively broad pronotum eventually convinced us that we had actually found P. heroicus—not a rare species, but also not something we expected to see at this relatively northeastern location compared to its usual haunts in southeastern Arizona.

Prionus heroicus male

Males generally landed in the vicinity of the prionic acid lures but seemed unable to locate its precise whereabouts.

Interestingly, none of the beetles came directly to the lure-baited traps but, rather, flew to the vicinity and then either circled broadly in the air or landed some distance from the trap and ran rapidly on the ground as if searching. We missed a few that we tried to net in mid-air, which responded by immediate straight line flight far away into the distance. Those that we watched while on the ground searched for up to several minutes but seemed unable to locate the lure, eventually taking flight and flying away. This suggests that prionic acid is more important as a component of the female’s calling pheromones for long distance detection, but that other components might be necessary to allow the males to locate females in the immediate vicinity. We returned to the site the following day around noon and were surprised to see males flying to our lures immediately after our arrival, since we had observed P. integer and P. fissicornis activity to occur almost exclusively during the morning and evening hours. Linsley et al. (1961), however, also note that adults of this species fly during the day, as well as at night. Again, none of the beetles seemed capable of finding the precise location of the lure.

Prionus heroicus male

Males rapidly searched the ground after landing but seemed unable to locate the lures and eventually flew off.

The larval habits of P. heroicus are not known, but the closely related P. californicus feeds on roots of living oaks or occasionally in dead oak logs or stumps (Linsley 1962). Prionus heroicus may have similar habits, since adult females have been found in leaf litter at the bases of living oak trees and stumps (Hovore & Giesbert 1976, Skiles 1978). Presumably the Mills Canyon population is associated with Gambel oak (Quercus gambelii), which was the only oak species noted at this location. An interesting defensive behavior was observed for an adult beetle that I kept alive and photographed later in the hotel room. I had put the beetle on a light-colored tray, and while I was getting the camera ready to take some shots the beetle began crawling away. I picked it up and put it back into position, and for a brief period it assumed the rather comical-looking “standing-on-its-butt” pose shown in the accompanying photograph. It maintained this pose for some seconds and then gradually lowered itself in the front before beginning to crawl again. If I poked it before it finished lowering itself it immediately pushed its front all the way back up again and held the pose briefly before beginning to lower itself again. I did not encounter this behavior in the field—beetles on the ground immediately tried to flee when I handled them and took flight if allowed to.

Prionus heroicus male

Defensive posturing as a result of being disturbed.

The story does not end here, however, as this would not be our last encounter with the species. While we were traveling back towards the east, we decided to visit the area around Black Mesa in the extreme northwestern corner of the Oklahoma panhandle. I had been to this site the previous summer, and although conditions were very dry at the time the presence of Gambel oak in the area—perhaps the easternmost outpost of this western species—had me thinking about the possibility of the area serving as an unknown eastern outpost for western species of jewel beetles (family Buprestidae). Conditions were once again dry, and no jewel beetles were found, but this quickly became a moot point as not long after our arrival a large prionid beetle came flying up to me—again in the same manner as P. heroicus at Mills Canyon and in the middle of the afternoon. We decided these, too, represented P. heroicus and remained in the area for the rest of the day, counting at least a dozen males attracted to the vicinity of the lures that we’d set out. As with the males we observed in New Mexico, none seemed able to find the precise location of the lure, and some circling only briefly and then flew off without even landing. Unlike New Mexico, however, we did succeed in trapping a few males in lure-bated traps that we set out overnight and checked the next morning.

Prionus heroicus male

This Prionus heroicus male  near Black Mesa, Oklahoma confirms the presence of this species in the state.

Our capture of P. heroicus at Mills Canyon in northeastern New Mexico and near Black Mesa in northwestern Oklahoma not only seems to suggest an association of this species with Gambel oak, but also confirms the previously questionable occurrence of the species in Oklahoma. Alexander (1958) reported the species (under one of its synonyms, P. tetricus) from Payne Co. in north-central Oklahoma. This record is dubious, since Payne Co. is well east of the 100th meridian and has a decidedly eastern flora, including its oaks. That record has not been mentioned by any subsequent authors addressing the overall distribution of P. heroicus (Linsley 1962, Chemsak 1996). The presence of P. heroicus in the Black Mesa area, with its decidedly western flora, does not necessarily give credence to the Payne Co. record, but it does bring up the question of just how broadly distributed P. heroicus really is. Most records of this species are from mountainous areas of southern Arizona and New Mexico, but more recently it has been recorded from Texas (Big Bend National Park) by Van Pelt (1999, 2007) and extreme northwestern Colorado (Moffat Co.) by Heffern (1998). The latter record is particularly interesting, as this represents a considerable range extension from any previously recorded locality and begs the question regarding its occurrence in other parts of Colorado.

REFERENCES:

Alexander, D. R. 1958. A preliminary survey of the cerambycids (Insecta: Coleoptera) of Oklahoma. Proceedings of the Oklahoma Academy of Science 38:43–52 [pdf].

Chemsak, J. A. 1996. Illustrated Revision of the Cerambycidae of North America. Volume I. Subfamilies Parandrinae, Spondylidinae, Aseminae, Prioninae. Wolfsgarden  Books, Burbank, California, x + 149 pp., plates I–X [description].

Heffern, D. J. 1998. Insects of Western North America 1A Survey of the Cerambycidae (Coleoptera) or Longhorn Beetles of Colorado. Gillette Museum of Arthropod Diversity, Department of Bioagricultural Sciences and Pest Management, Colorado State University, Fort Collins, i + 32 pp., 1 color plate [summary].

Hovore, F. T. & E. F. Giesbert. 1976. Notes on the ecology and distribution of western Cerambycidae (Coleoptera). The Coleopterists Bulletin 30:350 [JSTOR].

Linsley, E. G. 1962. The Cerambycidae of North America. Part II. Taxonomy and classification of the Parandrinae, Prioninae, Spndylinae, and Aseminae. University of California Publications in Entomology 19:1–102, 1 plate [OCLC WorldCat].

Linsely, E. G., J. N. Knull & M. Statham. 1961. A List of Cerambycidae from the Chiricahua Mountain area, Cochise County, Arizona (Coleoptera). American Museum Novitates 2050:1–34 [pdf].

Skiles, D. D. 1976. Taxonomy and description of two prionine Cerambycidae from southern Arizona: a new species of Stenodontes and new status for Neomallodon arizonicus (Coleoptera). Proceedings of the Entomological Society of Washington 80:407–423 [Biodiversity Heritage Library].

Van Pelt, A. F. (ed.). 1999. Inventory of insects of Big Bend National Park, Texas. Big Bend Natural History Association, Big Bend National Park, and Texas.

Van Pelt, A. F. (ed.). 2007. Inventory of insects of Big Bend National Park, Texas. Report to Big Bend National Park, 204 pp.

© Ted C. MacRae 2015

A Buprestis hat-trick!

In North America, few beetles can rival jewel beetles in the family Buprestidae for sheer beauty, and within the family this beauty is perhaps best exemplified by species in the family’s namesake genus Buprestis. Often flashing yellow, green, coppery, or red, species in this genus combine brilliant colors, moderately large size, and relative rarity to make them desirable additions to the collections of casual and serious collectors alike.

Buprestis (Knulliobuprestis) confluenta | Woods Co., Oklahoma

Buprestis (Knulliobuprestis) confluenta | Woods Co., Oklahoma

Over the years I’ve had some success collecting certain species in this genus, but even after 30 years of collecting there still remain several species that have eluded my net. A few weeks ago I posted photos of and wrote about my experience collecting not one, but two of these species that had so far eluded me—B. striata and B. salisburyensis—both as single specimens from a single tree in the mountains of Tennessee. Three weeks after collecting these species, I began a collecting trip west into the heart of the Great Plains with the primary purpose of using recently developed lures to collect some uncommonly encountered prionid longhorned beetles. Before reaching the first locality, however, field mate Jeff Huether and I stopped off in Hardtner, Kansas to visit with fellow beetle enthusiast “Beetle Bill” Smith and do a little collecting with him before continuing on. Bill took us to a spot just south of the nearby state line into Oklahoma to a spot where he had collected, among other things, yet another of the Buprestis spp. that had so far eluded me—B. confluenta.

Buprestis (Knulliobuprestis) confluenta

If B. striata and B. salisburyensis are beautiful, then B. confluenta is downright stunning! Brilliant green, perhaps with a slight coppery brown to purplish blue hue and with more or less confluent (thus the species name) fine yellow flecks densely scattered over the elytra, it is one of the easiest to identify of any species in the genus. Moreover, unlike the previous two species, I had already made several attempts to collect B. confluenta during several previous trips to the Great Plains, stopping wherever I saw a nice stand of the large eastern cottonwood trees (Populus deltoides) in which larvae of the species are known to develop. Frustratingly, as if to taunt me, I have even seen specimens on more than one occasion in one of the many 4H and student collections that I have examined over the years in Missouri and Kansas.

Buprestis (Knulliobuprestis) confluenta
Today was different, however—I could just feel it. Pulling up to the spot, my attention was immediately drawn to a grove of large cottonwood trees, some dead with bark-less trunks still standing. I thought to myself, “Those look like Buprestis trees!” While Bill and Jeff swept nearby herbaceous plants looking for a blister beetle that Jeff was interested in, I picked my way through the cottonwood grove, carefully approaching each tree—especially the dead ones—and scanning the trunk for any sign of movement or flash of color. It took a long time—well over an hour, by which time I had almost given up hope and begun thinking that this would be yet another unsuccessful attempt to collect the species. Suddenly, there it was sitting on the trunk of a recently fallen tree in all of its unmistakable glory! I froze at first, afraid of spooking it by too excited an approach, then remembered that species of Buprestis in general are not very skittish or as quick to take flight as many other members of the family (e.g., species in the genus Chrysobothris). I resumed my approach, albeit still cautiously just in case, and easily secured the specimen in a vial. My season now included a Buprestis hat-trick! The beetle was not only calm but actually seemed disinclined to flee, prompting me to release it back onto the tree trunk for a few quick field photos before placing it back in the vial for better photos later in the security of my hotel room that evening.

Buprestis spp. love large, dead, barkless cottonwood trunks.

Buprestis spp. love large, dead, bark-less cottonwood trunks.

Among North American jewel beetles, B. confluenta has one of the stranger geographic distributions, having been recorded from most of the central and western states and provinces as far east as Quebec and Indiana (Nelson et al. 2008). This is no doubt a consequence of the distribution of its primary larval hosts—cottonwood and quaking aspen (Populus tremuloides) (Burke 1917, Nicolay & Weiss 1918). In my Missouri checklist (MacRae 1991) I saw only two specimens from the state, both in counties along its western edge near Iowa and Kansas. Considering how abundant cottonwood is in Missouri, it is a mystery to me why this species should be so rare in the state. But then, I’ve not found it easy to come by in Kansas or Oklahoma either, where more records seem to exist than in any other state. Even on this occasion, when I would finally find the species after 30 years of searching, only this one, single specimen was seen. Interestingly, I would also collect about a half dozen specimens of the closely related species, B. rufipes (more commonly associated with oaks), off of the same trees that I searched so intensively for B. confluenta.

The requisite frontal portrait!

The requisite frontal portrait!

REFERENCES:

Burke, H. E. 1917. Flat-headed borers affecting forest trees in the United States. U.S. Department of Agriculture Bulletin 437, 9 pp. [Biodiversity Heritage Library].

MacRae, T. C. 1991. The Buprestidae (Coleoptera) of Missouri. Insecta Mundi 5(2):101–126 [pdf].

Nelson, G. H., G. C. Walters, Jr., R. D. Haines & C. L. Bellamy.  2008.  A Catalogue and Bibliography of the Buprestoidea of American North of Mexico.  Coleopterists Society Special Publication No. 4, The Coleopterists Society, North Potomac, Maryland, 274 pp. [description].

Nicolay, A. S. & H. B. Weiss. 1918. A review of the genus Buprestis in North America. Journal of the New York Entomological Society 26(2):75–109 [pdf].

© Ted C. MacRae 2014

2014 Great Plains Collecting Trip iReport

During the past year or so I’ve followed up my longer (one week or more) insect collecting trips with a synoptic “iReport”—so named because they are illustrated exclusively with iPhone photographs. It may come as a surprise to some, but iPhones actually take pretty good pictures (especially if you pay attention to their strengths and weaknesses), and their small, compact size makes it easy to take lots of photos while trying to use time in the field wisely. I find the iPhone to be a great tool for documenting the general flavor of a trip and for taking quick photos of subjects before getting out the big rig. I will, of course, feature photographs taken with the ‘real’ camera in future posts.

For this trip, I teamed up with Jeff Huether for the third time since 2012. Our quarry for this trip was longhorned beetles (family Cerambycidae) in the genus Prionus. Larvae of these beetles are subterranean, with some species feeding on roots of woody plants and others on roots of grasses and other herbaceous plants. Among the latter are an array of species occurring in the Great Plains, many of which have been very uncommonly collected. However, in recent years lures have been produced that are impregnated with prionic acid—the principal component of sex pheromones emitted by females in the genus. Originally produced for use in commercial orchards (which are sometimes attacked by P. laticollis in the east and P. californicus in the west), these lures are proving themselves to be useful for us taxonomist-types who wish to augment the limited amount of available material of other, non-economic species in the genus. While Prionus was our main goal, rest assured that I did not pass on the opportunity to find and photograph other beetles of interest.

I began the trip by driving from St. Louis to Wichita, Kansas to meet up with Jeff, who had flown there from his home in upstate New York. Our plan was to visit sites in southeastern Colorado and northeastern New Mexico, where several of the Prionus spp. that we were looking for were known to occur. Before doing this, however, we stopped in Hardtner, Kansas to see “Beetle Bill” Smith and tour his amazing natural history tribute, Bill and Janet’s Nature Museum.

"Beetle" Bill Smith, founder of Bill & Janet's Nature Museum, Hardtner, Kansas.

“Beetle Bill” Smith, founder of Bill & Janet’s Nature Museum, Hardtner, Kansas.

After the tour (and a delicious lunch at his house of fried crappie prepared by his wife Janet), we headed west of town and then south just across the state line into Oklahoma to a spot where Bill had found a blister beetle (family Meloidae) that Jeff was interested in finding. During lunch I mentioned a jewel beetle (family Buprestidae) that I had looked for in the area several times, but which had so far eluded me—Buprestis confluenta. Emerald green with a dense splattering of bright yellow flecks on the elytra, it is one of North America’s most striking jewel beetles and is known to breed in the trunks of dead cottonwoods (Populus deltoides). Bill mentioned that he had collected this species at the very spot where we were going, and when we arrived I was enticed by the sight of a cottonwood grove containing several large, dead standing trunks—perfect for B. confluenta.

Buprestis spp. love large, dead, barkless cottonwood trunks.

Buprestis spp. love large, dead, barkless cottonwood trunks.

I searched for more than one hour without seeing the species, though I did find a few individuals of the related (and equally striking) B. rufipes on the trunks of the large, dead trees. Once that amount of time passes I’m no longer really expecting to see what I’m looking for, but suddenly there it was in all of its unmistakable glory! It would be the only individual seen despite another hour of searching, but it still felt good for the first beetle of the trip to be one I’d been looking for more than 30 years!

Buprestis (Knulliobuprestis) confluenta, on large, dead Populus deltoides trunk | Woods Col., Oklahoma| USA: Oklahoma

Buprestis confluenta, on the trunk of a large, dead cottonwood (Populus deltoides) | Woods Col., Oklahoma| USA: Oklahoma

I usually wait until near the end of a collecting trip to take the requisite selfie, but on this trip I was sporting new headgear and anxious to document its maiden voyage. My previous headgear of choice, a vintage Mambosok (impossible to get now), finally disintegrated after 20 years of field use, and on the way out-of-town I picked up a genuine Buff® do-rag. I know many collectors prefer a brim, but I don’t like they way brims limit my field of vision or get in the way when I’m using a camera. Besides, I’m usually looking down on the ground or on vegetation, so sun on my face is not a big issue. And do I be stylin’ or wut?

A "selfie" makes the trip official.

A “selfie” makes the trip official.

We made it to our first locality in southeast Colorado by noon the next day—the vast, dry grasslands north of Las Animas. Jeff had collected a blister beetle of interest here on an earlier trip, but as I looked out across the desolate landscape I wondered what on Earth I could find here that would be even remotely interesting to me.

Shortgrass prairie habitat for Prionus integer.

Shortgrass prairie habitat for Prionus integer.

Letting Jeff have some time to look for his blister beetle, I started down the roadside and after a short time found a live female Prionus sp. (later determined to represent P. integer). The only female Prionus I had ever collected before was P. heroicus, a giant species out in Arizona, and that was almost 30 years ago, so I wasn’t immediately sure what it was. Eventually I decided it must be Prionus, and a quick stop to kick the dirt while Jeff looked for his beetle turned into an intense search for more Prionus that surely were there. I did find two male carcasses shortly thereafter, and then nothing more was seen for the next hour or so.

Prionus integer male | Bent Co., Colorado

Prionus integer male (found dead) | Bent Co., Colorado

During the time that I was searching, however, I started noticing strange burrows in the ground. I excavated a few—they were shallow but contained nothing. Nevertheless, they matched the size of the beetles perfectly—surely there was a connection?

Prionus integer adult burrow.

Prionus integer adult burrow.

I wondered if Jeff knew about the beetles occurring here, but when I showed him what I had found the surprised look on his face told me this was not the case. I showed him the burrows, and we both agreed they had to be connected. I got the shovel out of the truck and walked back to the area where I had seen the live female, then sunk the shovel deep into the ground next to one of the burrows and pried up a chuck of the soil containing the burrow in its entirety. As we broke apart the soil another female was revealed, and we immediately decided to set out some traps baited with prionic acid lures. We expected the beetles to become active during dusk, so we went into town to get something to eat and then check out another nearby locality before returning to the site at dusk. While we were gone it rained heavily at the site, so we weren’t sure if or how this would affect beetle activity and their possible attraction to the traps. However, as we approached the site (slipping and sliding on the muddy 2-track), we could actually see beetles crawling on the road from afar. What we found when we got out of the car was nothing short of mind-blowing—the beetles were everywhere, crawling on the road, crawling through the grass, and overflowing in the flooded traps! The vast majority were males, as expected, but we also found a fair number of the much more rarely collected females. This was significant, as the chance to observe mating and oviposition behavior made the encounter far more informative than if we had only found and collected the much more numerous males.

Prionus integer mating pair.

Prionus integer mating pair.

The following day we headed south into northeastern New Mexico to look at some shortgrass prairie sites near Gladstone (Union Co.) where two species of Prionus had been collected in recent years: P. fissicornis (the lone member of the subgenus Antennalia) and P. emarginatus (one of eight species in the poorly known subgenus Homaesthesis, found primarily in the Great Plains and Rocky Mountains). Fresh off of our experience the previous day, we were on the lookout for any suspicious looking “burrows” as we checked the roadsides at several spots in the area but found nothing, and while a few blister beetles piqued the interest of Jeff at one site, the complete absence of woody vegetation or flowering plants in general in the stark grassland landscape made the chances of me finding any other woodboring beetles remote. Eventually I became distracted by the lizards that darted through the vegetation around us, including this lesser earless lizard (Holbrookia maculata) and a collared lizard (better photos of both forthcoming).

Holbrookia maculata (lesser earless lizard) | Union Co., New Mexico.

Holbrookia maculata (lesser earless lizard) | Union Co., New Mexico.

Despite no clues to suggest that Prionus beetles were active in the area, we set out some traps at two sites with soil exposures that seemed similar to those seen the day before. As Jeff set the last pair of traps in place, my distraction with saurian subjects continued with a dusty hognose snake (Heterodon nasicus gloydi). While photographing the animal I looked down to my side, and what did I see but a male Prionus fissicornis crawling through the vegetation! I called out to Jeff, and for the next half an hour or so we scoured the surrounding area in a failed attempt to find more. We would not be back until the next morning to check the traps, so our curiosity about how abundant the beetles might be would have to wait another 18 hours. We cast an eye towards the north and watched late afternoon thunderstorms roll across the expansive landscape and decided to check out the habitat in nearby Mills Rim.

Thunderstorms over shortgrass prairie.

Thunderstorms over shortgrass prairie.

The rocky terrain with oak/pine/juniper woodlands at Mills Rim was a dramatic contrast to the gently rolling grasslands of the surrounding areas. We came here mostly out of curiosity, without any specific goal, but almost immediately after getting out of the car a huge Prionus male flew up to us—almost surely attracted by the scent of the lures we were carrying. Within a few minutes another male flew in, and then another. Because of their huge size and occurrence within oak woodland habitat, we concluded they must represent P. heroicus, more commonly encountered in the “Sky Islands” of southeastern Arizona. We stuck around to collect a few more, but as dusk approached we returned to the surrounding grasslands to set out some lures to see if we could attract other Prionus species. The frontal system that had waved across the landscape during the afternoon had left in its wake textured layers of clouds, producing spectacular colors as the sun sank inexorably below the horizon.

Sunset over shortgrass prairie.

Sunset over shortgrass prairie.

This attempt to collect grassland Prionus beetles would not be successful, and as dusk progressed we became distracted collecting cactus beetles (Moneilema sp., family Cerambycidae) from prickly pear cactus plants (Opuntia sp.) before darkness ended our day’s efforts. This did not mean, however, that all of our efforts were done—there are still night active insects, and in the Great Plains what better nocturnal insect to look for than North America’s largest tiger beetle, the Great Plains giant tiger beetle (Amblycheila cylindriformis, family Cicindelidae—or subfamily Cicindelinae—or supertribe Cicindelitae, depending on who you talk to)?! We kept our eyes on the headlamp illuminated 2-track as we drove back to the highway and then turned down another road that led into promising looking habitat. Within a half-mile of the highway we saw one, so I got out to pick it up and then started walking. I walked another half-mile or so on the road but didn’t see anything except a few Eleodes darkling beetles (family Tenebrionidae), then turned around and walked the habitat alongside the road on the way back. As I walked, tiny little rodents—looking like a cross between a mouse and a vole—flashed in and out of my headlight beam as they hopped and scurried through the vegetation in front of me. Most fled frantically in response to my attempted approach, but one, for some reason, froze long enough under my lamp to allow me this one photo. When I posted the photo on my Facebook page, opinions on its identity ranged from kangaroo rat (Dipodomys sp.) silky pocket mouse (Perognathus flavus) to jumping mouse (Zapus sp.). Beats me.

silky pocket mouse? Zapus sp., jumping mouse? | Union Co., New Mexico.

Kangaroo rat? Silky pocket mouse? Jumping mouse? | Union Co., New Mexico.

Almost as if by command, it rained during the early evening hours where we had set the traps, and the following morning we were rewarded with traps brimming with Prionus fissicornis males. Not only were the traps full, but males were still running around in the vicinity, and we even found a few females, one of which was in the act of ovipositing into the soil at the base of a plant.

Prionic acid-bated traps w/ Prionus fissicornis males.

Prionic acid-bated traps w/ Prionus fissicornis males.

Prionic acid-bated traps w/ Prionus fissicornis males.

Prionic acid-bated traps w/ Prionus fissicornis males.

Prionus fissicornis male | Harding Co., New Mexico.

Prionus fissicornis male | Harding Co., New Mexico.

Prionus fissicornis oviposition hole.

Prionus fissicornis oviposition hole.

Eventually P. fissicornis activity subsided, and we decided to go back to the area around Mills Rim to see what beetles we might find in the woodland habitats. We also still were not sure about the Prionus beetles we had collected there the previous day and whether they truly represented P. heroicus. The scrubby oaks and conifers screamed “Beat me!”, and doing so proved extraordinarily productive, with at least a half-dozen species of jewel beetles collected—including a nice series of a rather large Chrysobothris sp. from the oaks that I do not recognize and a single specimen of the uncommonly collected Phaenops piniedulis off of the pines.

Oak/juniper woodland at Mills Canyon, habitat for Prionus heroicus.

Oak/juniper woodland at Mills Canyon, habitat for Prionus heroicus.

Not only is the scenery at Mills Rim Campground beyond spectacular, it also boasts some of the most adoringly cute reptiles known to man—such as this delightfully spiky horned lizard (I prefer the more colloquial name “horny toad”!). I’m probably going to regret not having photographed this fine specimen with the big camera.

Phrynosoma cornutum (Texas horned lizard) | Harding Co., New Mexico.

Phrynosoma cornutum (Texas horned lizard) | Harding Co., New Mexico.

Fresh diggings beside a rock always invite a peek inside. You never know who might be peeking out.

Who's home?

Who’s home?

Peek-a-boo!

Peek-a-boo!

The trip having reached the halfway point, we debated whether to continue further south to the sand dunes of southern New Mexico (with its consequential solid two-day drive back to Wichita) or turn back north and have the ability to collect our way back. We chose the latter, primarily because we had not yet had a chance to explore the area around Vogel Canyon south of Las Animas, Colorado. We had actually planned to visit this area on the day we encountered P. integer in the shortgrass prairie north of town, and a quick visit before going back to check the traps that evening showed that the area had apparently experienced good rains as shown by the cholla cactus (Cylindropuntia imbricata) in full bloom.

Cylindropuntia imbricata | Otero Co., Colorado.

Cylindropuntia imbricata | Otero Co., Colorado.

Whenever I see cholla plants I can’t help myself—I have to look for cactus beetles (Moneilema spp.). It had rained even more since our previous visit a few days ago, and accordingly insects were much more abundant. Several Moneilema adults were seen on the cholla, one of which I spent a good bit of time photographing. The iPhone photo below is just a preview of the photos I got with the big camera (which also included some very impressive-sized cicadas—both singing males and ovipositing females). The cactus spines impaled in the camera’s flash control unit serve as a fitting testament to the hazards of photographing cactus insects!

Moneilema sp. on Cylindropuntia imbracata } Otero Co., Colorado.

Moneilema sp. on Cylindropuntia imbracata } Otero Co., Colorado.

The hazards of photographing cactus beetles.

The hazards of photographing cactus beetles.

Later in the afternoon we hiked down into the canyon itself, and while insects were active we didn’t find much out of the ordinary. We did observe some petroglyphs on the sandstone walls of the canyon dating from the 1200s to the 1700s—all, sadly, defaced by vandals. Despite the rather uninspiring collecting, we stayed in the area for two reasons: 1) Jeff wanted to setup blacklights at the canyon head in hopes of collecting a blister beetle that had been caught there on an earlier trip, and 2) I had noted numerous Amblycheila larval burrows in the area (and even fished out a very large larva from one of them) and wanted to search the area at night to see if I could find adults. Jeff was not successful in his goal, and for a time I thought I would also not succeed in mine until we closed up shop and started driving the road out of the canyon. By then it was after 11 p.m. and we managed to find about a half-dozen A. cylindriformis adults. This was now the third time that I’ve found adults of this species, and interestingly all three times I’ve not seen any beetles despite intense searching until after 11 p.m and up until around midnight.

Lithographs on canyon wall | Mills Canyon, Colorado.

Lithographs on canyon wall | Mills Canyon, Colorado.

The next morning we found ourselves with two days left in the trip but several hundred miles west of Wichita, where I needed to drop Jeff off for his flight back home before I continued on home to St. Louis. I had hoped we could make it to the Glass Mountains just east of the Oklahoma panhandle to see what Prionus species might be living in the shortgrass prairies there (and also to show Jeff this remarkable place where I’ve found several new state records over the past few years). As we headed in that direction, I realized our path would take us near Black Mesa at the western tip of the Oklahoma pandhandle, and having been skunked on my first visit to the area last year due to dry conditions but nevertheless intrigued by its very un-Oklahoma terrain and habitat I suggested we stop by the area and have a look around before continuing on to the Glass Mountains. We arrived in the area mid-afternoon and headed straight for a rock outcropping colonized by scrub oak (Quercus sp.) and pinyon pine (Pinus sp.)—very unusual for western Oklahoma—that I had found during my previous trip.

The author looks pensively out over the Black Mesa landscape.

The area around Black Mesa couldn’t be more unlike the perception that most people have of Oklahoma.

I wanted to beat the oaks for buprestids—surely there would be a state record or two just sitting there waiting for me to find them, but as I started walking from the car towards the oaks the approach of a loud buzz caught my attention. I turned around to see—would you believe—a large Prionus beetle circling the air around me and was fortunate to net it despite its fast and agile flight. I hurried back to the car to show Jeff what I had found; we looked at each other and said, “Let’s collect here for a while.” The beetle had apparently been attracted to the lures in the car, so we got them out, set them up with some traps, and went about beating the oaks and watching for beetles to fly to the lure. Sadly, no  jewel beetles were collected on the oaks, although I did find evidence of their larval workings in some dead branches (which were promptly collected for rearing). Every once in a while, however, a Prionus beetle would fly in, apparently attracted to the lure but, curiously, never flying directly to it and falling into the trap. Many times they would land nearby and crawl through the vegetation as if searching but never actually find the trap. However, just as often they would approach the trap in flight and not land, but rather continue circling around in the air for a short time and before suddenly turning and flying away (forcing me to watch forlornly as they disappeared in the distance). Based on their very large size, blackish coloration and broad pronotum, we surmised (and later confirmed) these must also be P. heroicus, despite thinking (and later confirming) that the species was not known as far east as Oklahoma. Not only had we found a new state record, but we had also recorded a significant eastern range extension for the species. And to think that we only came to Black Mesa because I wanted to beat the oaks!

Prionus heroicus male

Prionus heroicus male

Bite from Prionus heroicus male.

Proof that Prionus heroicus males can bite hard enough to draw blood!

We each collected a nice series of the beetles, and despite never witnessing the beetles actually going to the traps a few more were found in the traps the next morning after spending the night in a local bed & breakfast. I also found a dove’s nest with two eggs hidden in the vegetation, and as we were arranging for our room at the bed & breakfast a fellow drove up and dropped off a freshly quarried dinosaur footprint (the sandstone, mudstone, and shale deposits around Black Mesa are the same dinosaur fossil bearing deposits made more famous at places like Utah’s Dinosaur National Monument).

Dove's nest w/ eggs.

Dove’s nest w/ eggs.

Dinosaur fossil footprint

Freshly quarried dinosaur fossil footprint

By the way, if you ever visit the area, the Hitching Post at Black Mesa is a great place to stay. A longhorn skull on the barn above an authentic 1882 stagecoach give a hint at the ambiance, and breakfast was almost as good as what my wife Lynne can do (almost! 🙂 ).

Longhorn skull on barn at our Bead and Breakfast.

Longhorn skull on barn at our Bead and Breakfast.

132-year-old stagecoach - model!

132-year-old stagecoach – model!

After breakfast we contemplated the long drive that lay between us and our arrival in Wichita that evening—our longer than expected stay in the area had virtually eliminated the possibility to collect in the Glass Mountains. Nevertheless, there was one more thing that I wanted to see before we left—the dinosaur footprints laying in a trackway along Carrizo Creek north of the mesa. I only knew they were in the area based on a note on a map, but as there were no signs our attempt to find them the previous day was not successful. Armed with detailed directions from the B&B owners, however, we decided to give it one more shot. Again, even after we found the site I didn’t see them immediately, I suppose because I was expecting to see distinct depressions in dry, solid rock. Only after the reflections of light from an alternating series of small puddles—each measuring a good 10–12″ in diameter—did I realize we had found them. Recent rains had left the normally dry creek bed filled with mud, with the footprints themselves still filled with water.

Dinosaur tracks | vic. Black Mesa, Oklahoma.

Dinosaur tracks | vic. Black Mesa, Oklahoma.

It is not surprising that I would be so excited to find the tracks, but what did surprise me was the effect they had on me. Seeing the actual signs of near mythical beasts that lived an incomprehensible 100 million years ago invites contemplation and reminds us that our time here on Earth has, indeed, been short!

Dinosaur tracks | vic. Black Mesa, Oklahoma.

Dinosaur tracks | vic. Black Mesa, Oklahoma.

By this time, we had no choice but to succumb to the long drive ahead. We did manage to carve out a short stop at the very first locality of the trip in an effort to find more Buprestis confluens (finding only a few more B. rufipes), but otherwise the day was spent adhering to our goal of reaching Wichita before nightfall. Jeff was home and sipping tea before lunchtime the next day, while I endured one more solid day of driving before making it back to St. Louis in time for dinner with the family. At that point, the trip already could have been considered a success, but how successful it ultimately ends up being depends on what beetles emerge during the next season or two from these batches of infested wood that I collected at the various spots we visited.

Wood collected for rearing wood-boring beetles.

Wood collected for rearing wood-boring beetles.

If you like this Collecting Trip iReport, you might also like the iReports that I posted for my 2013 Oklahoma and 2013 Great Basin collecting trips as well.

© Ted C. MacRae 2014