Art and Bob and Ted’s Excellent Adventure

Back in mid-July, I was fortunate to have the chance to host a couple of beetle colleagues from out-of-state and to show them a few of Missouri’s premier habitats. Art Evans is a scarab specialist from Virginia who has published several books on beetles—most recently “Beetles of Eastern North America” and “Beetles of Western North America.” Bob Anderson is a weevil specialist from the Canadian National Collection in Ottawa. I’ve been in the field with Art a couple of times, both out west in Arizona, while this is my first time in the field with Bob. They are passing through as they work their way west for a 4-week collecting trip, and since neither has ever collected in Missouri it was a perfect opportunity to spend a few days together.


Sand Prairie Conservation Area

Photo by Art Evans.

We met up at Sand Prairie Conservation Area in southeastern Missouri, which contains a high quality sand prairie remnant—one of Missouri’s rarest and most endangered natural communities. This is the place where a few years ago I discovered two scarab beetles not previously known to occur in Missouri—including Strategus antaeus (smooth ox beetle), the second largest beetle in eastern North America.

L-R: Me, Art Evans, and Bob Anderson (representing 195 years of entomological expertise!).

I was hoping there might still be jewel beetles (family Buprestidae) on the Quercus spp. (oaks) and Diospyros virginiana (persimmon) trees ringing the perimeter of the sand blowout area, and my hopes increased when I swept two Dicerca obscura off high branches of the latter. Those would be the last buprestids I saw there. Sweeping the high branches of Quercus marilandica (blackjack oak) and southern Q. falcata (red oak) produced only epitragine tenebrionids.

Me using a long-handled net to shake loose buprestids and cerambycids from Diospyros virginiana (persimmon). Photo by Art Evans.

I looked for live individuals of S. antaeus but, as is typical, only saw bits and pieces of carcasses near the base of certain oaks. Under one I did collect an almost intact female carcass (missing only the head), and the head and pronotum of a super-major male.

Strategus antaeus carcass remnants found under Quercus marilandica (post oak) at sand blow perimeter in xeric sand prairie remnant. Photo by Art Evans.

Continued sweeping of high branches finally produced something besides epitragines—an Enoclerus sp. Returning to the vehicles, I exchanged my long-handled net for a sweep net and swept the Polygonellum americanum (American jointweed), collecting only a single Cryptocephalus sp. For their part, Bob and Art collected a small variety of beetles, including some weevils that Bob found interesting.

Stylisma pickeringii (Pickering’s dawnflower) in dry sand prairie remnant.

Otter Slough Conservation Area

After Art and Bob had seen enough of Sand Prairie, I took them to Otter Slough Conservation Area. This area features sloughs and wet bottomland forests where I’ve collected many good species, most notably Agrilus concinnus—formerly considered very rare until I determined it to be a later-season species associated with species of Hibiscus (rose mallow)—and an undescribed species of Taphrocerus that I’ve found on Carex hyalinolepis here but nowhere else in the state (despite finding the host plant). I alerted Art and Bob to these possibilities and started down the 2-track where I’ve seen these species before.

My beating sheet has seen better days. Photo by Art Evans.

Unfortunately, the 2-track was completely overgrown to the point of being impassable (it’s been many years since I last walked it), so I opted instead to walk the perimeter of Mudsnake Marsh where I’ve also seen the two species. The marsh was dry—first time I’ve seen it like that, allowing me to check Hibiscus plants in the marsh interior as well as along the edge. No A. concinnus were seen on any of the plants—just a few Paragrilus tenuis and good numbers of Chaetocnema quadricollis (hibiscus flea beetle).

Chaetocnema quadricollis on Hibiscus lasiocarpus (rose mallow) in lowland wetland.

Failing to find A. concinnus, I swept the Carex along the perimeter on the edge of the wet bottomland forest side and even in a spot where I had collected Taphrocerus abundantly in previous years, but to no avail and finding instead only one chrysomelid. Collecting was slim at Sand Prairie, and it was even slimmer here at Otter Slough, so it appears seems we have definitely entered the “summer doldrums” stage of the collecting season—at least for beating and sweeping.

Owl-fly (family Myrmeleontidae), probably genus Ululodes, sweeping sedges in lowland wetland.

Holly Ridge Natural Area

After going into bearby Dexter to get a motel room and eat some dinner (Dexter Bar-B-Q pulled pork—pretty good!), I took Art and Bob to Holly Ridge Natural Area for an evening of blacklighting. I wanted to come here to 1) avoid the hoards of aquatic insects that would come to our sheets if we blacklighted at Otter Slough and 2) give myself a chance (however outside) of getting Saperda obliqua—known from only a single specimen collected at Hawn State Park but likely here as well due to the stands of Alnus serrulata (hazel alder).

Photo by Art Evans.

Bob and Art agreed it looked like a good spot to blacklight when we arrived, so Art placed his light setup in the parking lot, while I placed mine about 100 meters into the mesic lowland deciduous forest. I had high hopes for the evening—it was warm (89°F) and humid, and we were two days past the full moon so moonlight would not be an issue for at least two hours after sunset.

Art Evans collects beetles at a mercury-vapor light.

A lot of insects ended up coming to the lights, but not a lot of cerambycids (and certainly not S. obliqua). Nevertheless, I picked up one Enaphalodes atomarius and a few Lepturges confluens, along with some telamonine treehoppers and other miscellaneous beetles—all at my light setup.

Enaphalodes atomarius (robust oak borer) at UV/MV light in mesic lowland deciduous forest.

We also saw Elytrimitatrix undata and several other miscellaneous beetles on the trunk of a large standing Quercus sp., and E. atomarius and another E. undata at Art’s lights, all of which I let him collect. He was also happy to see the many Lucanus capreolus and Neocicada hieroglyphica that came to both our lights. By 10:30 pm few additional insects were coming to the lights, so we broke them down and heads back to Dexter.

Elytrimitatrix undata (family Disteniidae) nocturnally on trunk of large standing Quercus sp. in mesic lowland deciduous forest.
Neocicada hieroglyphica (hieroglyphic cicada) at UV/MV light in mesic lowland deciduous forest.
Callosamia angulifera (tuliptree silkmoth) at UV/MV light in mesic lowland deciduous forest.
Phobetron pithecium (hag moth) at UV/MV light in mesic lowland deciduous forest.
Dolomedes albineus (whitebanded fishing spider) nocturnally on trunk of large standing Quercus sp. in mesic lowland deciduous forest.

Long Bald Glade Natural Area

It took us all morning to drive across southern Missouri to reach this near the easternmost limits of the White River Hills, a fantastic region in the extreme southwest of Missouri featuring dry oak/juniper woodland surrounding extensive dolomite glades atop rounded knobs. It is my favorite region of the state for collecting insects, and I wanted Art and Bob to see the area before they continue on to western Texas. I also have jug traps and a Lindgren funnel trap placed here, so coming here would give me a chance to service them before checking the rest during the next two days on the way back to St. Louis.

By the time we arrived, temperatures were already soaring, and I was disappointed to see that conditions were very dry. Nevertheless, you never know how collecting will be until you try. I started out by checking the Lindgren funnel trap, which I had placed in actively-restored dry post oak woodland and was pleased to see a variety of beetles, including a series of Neoclytus scutellatus. In addition to re-baiting with ethanol-only, I added a pheromone lure (Fuscumol Lure MR, #P655-MR, Chemtica International) to the trap to increase the attractiveness of the trap to cerambycids.

Bob Anderson watches as I service a Lindgren funnel trap.

The ethanol/red wine trap had a nice variety of beetles that caught the attention of Art and Bob and included Plinthocoelium suaveolens, Stenelytrana emarginata, Eburia quadrigeminata, undetermined elaphidiines, Euphoria fulgida, E. sepulchralis, Cnephus mutilatus, miscellaneous beetles, and a cicada. The ethanol-only trap, however, was much less productive, yielding no cerambycids and just one E. sepulchralis, a few C. mutilatus, and some miscellaneous beetles. This continues the trend noticed four weeks ago during the first round of trap collections, when it appeared that the ethanol/red wine traps yielded higher numbers and diversity of not only beetles but also bycatch of other insects (primarily moths, flies, and wasps), suggesting that red wine possesses additional components attractive to beetles that more than make up for the reduction in ethanol content resulting from mixing the two.

Servicing a “jug” trap.
“Jug trap” catch.

The condition of the glade vegetation was extremely dry, and as I walked between the traps I didn’t see a single plant in bloom. I encountered Bob on the way back to the car and mentioned this to him, and I suggested to both Art and Bob that instead of spending time here we should look at locations further west that may have gotten more rain. They agreed, so we cut our visit short and headed further west.


Mincy Conservation Area

As we traveled west, we passed two locations where I had placed traps, but the entire area still looked exceedingly dry so we didn’t stop. By the time we reached Branson, however, conditions looked much better, and I suggested to Art and Bob that we visit Mincy Conservation Area just south of town since we were now in an area that looked like it had received some rain. Mincy is another of my favorite localities in this area due to the presence of high-quality dolomite glades, and in fact it is one of the localities where I have placed jug traps. Beetle numbers and diversity were much higher in the ethanol/red wine trap, which had Plinthocoelium suaveolens, Eburia quadrigeminata, elaphidiines, Neoclytus scutellatus, Strangalia luteicornis, Acmaeodera texana, Cotinis nitida, Euphoria fulgida, E. sepulchralis, elaterids, Cnephus mutilatus, mordellids, and a cicada, while the ethanol-only trap yielded most of the same but in lower numbers. While servicing the traps, I noticed an Acmaeodera on the flower of Liatris hirsuta (hairy blazingstar). I at first assumed it was A. pulchella, a common summertime species here, but looking more closely at it I realized it was A. texana, a very uncommon species in Missouri that I have not seen for many years. I looked at other Liatris flowers but didn’t see any, so I swept the surrounding herbaceous vegetation of this xeric dolomite prairie and found one more (plus a few chrysomelids). We had originally planned to go further west to Roaring River State Park for an evening of blacklighting. Hiwever, seeing that the vegetation looked good here and that the area had obviously gotten some rain, I suggested that we stay here instead of taking a chance on moving to an area where we were not sure what conditions were like. This also would allow us to spend more time collecting—we could go into Branson for dinner and return here quickly rather than driving another hour to Roaring River. Art and Bob agreed this was a good idea, so we headed into town and enjoyed Mexican cuisine at Los Poblanos (I had crispy tacos and, in a true rarity, resisted the siren call of the beans, rice, chips and salsa that would have totally derailed my 3-week stretch of healthy eating).

Acmaeodera texana on flower of Liatris hirsuta (hairy blazingstar) in xeric dolomite prairie remnant.

We returned with still about an hour before it would start getting dark, so I decided to continue sweeping the glade vegetation to look for more Acmaeodera texana. I paid particular attention to any flowers (primarily Liatris hirsuta and Rudbeckia missouriensis), since those were the plants most likely to have the beetles on them. Most sweeps, while not yielding A. texana, did produce hispine leaf beetles in the genera Anisostena and Microrhopala—certainly interesting enough to collect and motivate me to continue sweeping. After going through one particular area and looking at the sweep contents on the net, I saw the unmistakable shape of an Agrilus. This was not, however, just any Agrilus, but rather A. impexus—one of North America’s rarest Buprestidae! I recognized it because some years ago I received two specimens from another person who collected them sweeping prairie vegetation at Ha Ha Tonka State Park in west-central Missouri. I identified them as this species but noted they were much larger than specimens collected commonly in the southwestern U.S. I sent the specimens to Henry Hespenheide, who not only confirmed their identity but also determined they were not conspecific with a more common but as yet undescribed southwestern species going under the same name. In reality, true A. impexus is very rare, known from only a handful of specimens—most many decades old—collected in the tallgrass as prairie region of the central U.S., and the common but unnamed southwestern species was described as A. paraimpexus. I have swept tallgrass prairies abundantly ever since but failed to find the species—until now. I alerted Art and Bob to the find and worked up to the brink of darkness sweeping the area to look for more. I never did, but Bob, on his last sweep of vegetation before closing darkness, found another (right in the same area where I had collected mine)!

Acanthepeira stellata (starbellied orbweaver) swept from herbaceous vegetation in xeric dolomite prairie remnant.

As exciting as this find was, darkness prevented continuing to look for it, and I had to take advantage of what little light remained to setup my lights. I felt a few sprinkles as I did this but didn’t think much about it, assuming it would pass, and fired up the generator. The sprinkles continued, however, and gradually increased to the point where I worried about the mercury-vapor bulb. Art had also set up his lights but waited on turning on the mercury-vapor light, and together we hoped against hope that it would blow over and we would be able to proceed with blacklighting. It was all in vain though, as temps continued to drop and light sprinkles turned to steady drizzle. With darkness well developed and absolutely no insects flying around the ultraviolet lights (which we had left on, it was clear that it would be pointless to continue. With that, we took our lights down and said our goodbyes, as their plans were to head towards Texas in the morning while I went to Roaring River to begin checking the rest of my jug traps as I worked my way back to St. Louis. It seemed an inappropriately inauspicious end to an otherwise successful stop and fun trip with two exceptional coleopterists. I’m already looking forward to the next chance I get to spend time in the field with each of them!

Asclepias viridiflora (green milkweed, green comet milkweed, green-flowered milkweed) in xeric dolomite prairie remnant.

Roaring River State Park—Chute Ridge Glade

First stop of the day to service jug traps, and the area has apparently gotten more rain as the glade vegetation looked reasonably lush. I was hoping to see more Glycobius speciosus here after getting a single individual in the ethanol/red wine trap last time, but that was not the case. The trap was, however, still full of beetles, including numerous Plinthocoelium suaveolens and a variety of other longhorns such as Eburia quadrigeminata, Elytrimitatrix undatus, and elaphidiines, scarabs like Cotinis nitida, Euphoria fulgida, and E. sepulchralis, and other beetles such ad elaterids, one Chrysobothris chlorocephala?, one Chariessa pilosa, mordellids, and Cnestus mutilatus. The ethanol-only trap, as before, had fewer individuals and lower diversity. Between checking the traps, I swept the herbaceous glade vegetation and didn’t get much, but after checking the second trap I swept along the roadside and got five more Agrilus impexus, effectively doubling the known series of this species! I was very happy to have collected more of this very rare species after the two that Bob and I collected yesterday at Mincy Conservation Area.

Asclepias verticillata (whirled milkweed) in xeric dolomite prairie remnant.

Hercules Glades Wilderness

The jug traps here didn’t produce much last time, and I wasn’t expecting much this time either. The ethanol/red wine trap had Plinthocoelium suaveolens, Eburia quadrigeminata, a few elaphidiines, one Cotinis nitida, several Euphoria fulgida, a couple of elaterids, and a clerid, while the ethanol-only trap had one cicada, one elaterid, and one miscellaneous beetle. I also collected Cicindela sexguttata and Cicindelidia rufiventris on the rocky-clay path through dry oak-juniper woodland; however, sweeping the herbaceous glade vegetation produced nothing. The area has been quite dry, but I believe the low trap numbers are also due to the lack of vegetational diversity here due to the lack of any management via prescribed burns.

Cicindelidia rufiventris (eastern red-bellied tiger beetle) on rocky-clay path through dry oak-juniper woodland.

Blackjack Knob

There is lots of Sideroxylon lanuginosum (gum bumelia) at this Mark Twain National Forest site along Hwy 160 in Taney Co., and last time I checked the jug traps here there was already Plinthocoelium suaveolens in them. As a result, I expected to see lots of them this time, and such was the case. The ethanol/red wine trap had nearly three dozen P. suaveolens along with Stenelytrana emarginata, Eburia quadrigeminata, a few elaphidiines, Cotinis nitida, Euphoria fulgida, E. sepulchralis, and elaterids m. The ethanol-only trap had far fewer P. suaveolens (only 2) but similar numbers and diversity of other beetles. The area looks quite dry, but I swept the herbaceous glade vegetation anyway in hopes that I would find more Agrilus impexus. I did not, but I did collect about a dozen Taphrocerushowardi” (the quotation marks are a story for another day), which was surprising to me given how crispy brown the vegetation looked.

View from Blackjack Knob.

Bald Hill Glade Natural Area

This was my best locality last time in terms of trap numbers (at least the ethanol/red wine trap, not so much the ethanol-only trap). This is also one of two localities where I have a Lindgren funnel trap, which had a nice diversity of cerambycids (Neoclytus scutellatus, Eburia quadrigeminata, Aegomorphus modestus, Xylotrechus colonus, an elaphidiine, and an undet. cerambycid), scarabs (Cotinis nitida, Anomala sp.), elaterids, (scolytines (C. mutilatus), and other miscellaneous beetles. The area looked quite dry—both in the glade proper and in the surrounding dry upland deciduous forest, but trap numbers (again, only in the former) were incredible. In fact, there were so many beetles in the ethanol/red wine trap that I skipped sorting them in the field and brought the whole unsorted catch back to the car to sort out later. Part of this decision was due to the dark line of clouds I noticed to the north and that seemed to be moving closer. I hustled to the ethanol-only trap and quickly sorted its much sparser contents (only three Plinthocoelium suaveolens and a few E. quadrigeminata, E. fulgida, and elaterids), all the time keeping an eye on the approaching front. As I started the half-mile hike back, the winds really started picking up and the temps began to drop, and it became clear it would be a race to reach the car before the skies opened up. There isn’t much that phases me when I’m out in the field, but bushwhacking a half-mile stretch of abandoned, overgrown 2-track under dark, ominous clouds and with increasingly gusty winds had me feeling a bit nervous. I almost made it to the car before the rain started but did get wet in the final stretch. Still, I was able to get out of the area and back onto asphalt before the real deluge started. All traps were re-baited with the same bait, and a pheromone lure (Fuscumol Lure MR, #P655-MR, Chemtica International) was added to the Lindgren funnel. Sweeping herbaceous glade vegetation leading to the ethanol-red wine trap produced only a single Exema sp.

Threatening clouds.

That evening in the hotel room, I sorted the contents of the ethanol/red wine trap and recovered a nice diversity of cerambycids (Plinthocoelium suaveolens, Stenelytrana emarginata, Enaphalodes atomarius, Eburia quadrigeminata, Neoclytus scutellatus, Lepturges confluens, and elaphidiines), one Chrysobothris sp., numerous cetoniine scarabs (Cotinis nitida, Euphoria fulgida, E. herbacea, E. sepulchralis), elaterids, and Cnestus mutilatus, a mordellid, and two miscellaneous beetles.

Sorted contents of an ethanol-red wine-baited “jug trap.”
Euphoria fulgida (emerald euphoria) in ethanol/red wine-baited jug trap in xeric dolomite prairie remnant. The pink morph (right) and pink/green morph (center) were the only ones out of many normal green morphs (left) trapped.

Peck Ranch Conservation Area

I wasn’t optimistic about the jug traps here, since it was rather unremarkable dry-mesic upland deciduous forest—a last minute replacement locality for a dolomite glade that I could not access due to a closed gate. The area got a good rain last night and looked lush anyway, and as it turned out there were huge numbers of beetles in the ethanol/red wine trap. These included one Plinthocoelium suaveolens—unexpected, since I didn’t think there was any gum bumelia in the area, a nice series of Stenelytrana emarginata, lots of Eburia quadrigeminata, Cotinis nitida, and Euphoria fulgida, single specimens of Neoclytus scutellatus, Strangalia luteicornis, and Batyle suturalis, Euphoria sepulchralis, Trigonopeltastes delta, and Chrysobothris sp., and a handful of elaterids, Cnestus mutilatus, and other miscellaneous beetles. As at most previous spots, the ethanol-only trap had fewer numbers and lower diversity of beetles, although this did include one Lepturges sp. not caught yet in ethanol/red wine. Given the success with sweeping at previous spots (Agrilus impexus as two locations and Taphrocerushowardi” at the last one), I decided sweep the herbaceous roadside vegetation in this dry mesic upland deciduous forest and got singletons of Taphrocerus nicolayi, Acmaeodera pulchella, and Exema sp.—the Taphrocerus definitely making the effort worthwhile.

Helenium amarum (bitterweed) along roadside in dry mesic upland deciduous forest.

Stegall Mountain Natural Area

I was not a fan of this spot when I set the traps in mid-May given the fact that the spot and a large surrounding area had been recently burned. Despite that, I did get Purpuricenus in one of the traps last time, so I was more optimistic about it this time. As with the previous spot the area looked lush and got good rain yesterday, and as with the previous spot there were huge numbers of beetles in the ethanol/red wine trap, the most exciting of which were Stenelytrana emarginata, Purpuricenus humeralis, and Knullianaspinifera” (I think this may actually be an undescribed species). Other longhorns in the trap included lots of Eburia quadrigeminata, Neoclytus scutellatus (may include a few N. mucronatus), and elaterids and small numbers of Batyle suturalis, Enaphalodes atomarius, Lepturges sp., Chrysobothris/Actenodes, Euphoria fulgida, E. sepulchralis, Cnestus mutilatus and other miscellaneous beetles. I was especially happy to see Purpuricenus once again—the first of this trip. Again, the ethanol-only trap had fewer numbers and lower diversity of beetles, but this did include a single specimen of Phaenops aeneola—only the second known Missouri specimen (I collected the first many years ago at this very location)! Sweeping herbaceous vegetation in the xeric rhyolite prairie around Trap A and dry oak/pine woodland around Trap B produced nothing, but doing so along the roadside in the oak/pibe woodland produced Microrhopala vittata and Exema sp.

Liatris hirsuta (hairy blazingstar) in post-burn understory regrowth of dry post oak woodland.

Russell Mountain, Jakk’s Glade

This area also looked lush and got more good rains last night. The ethanol/red wine trap was down last time (not vandalized, I just failed to tie the knot on the carabiner securely), so I was anxious to see what it might produce. Like the previous spots on this side of the state, there were huge numbers of beetles in it, including Stenelytrana emarginata, Purpuricenus humeralis, Eburia quadrigeminata, Knullianaspinifera”, Neoclytus scutellatus (may include a few N. mucronatus), Enaphalodes atomarius, Strangalia luteicornis, undetermined elaphidiines, Chrysobothris sp., Cotinis nitida, Euphoria fulgida, E. sepulchralis, and undetermined elaterids (including many very tiny ones that I didn’t collect). The large series of Purpuricenus was unexpected and quite nice and means I’ve now collected species in this genus in all three of the igneous glades where I placed traps (but none in any of the dolomite glades—the opposite of what I expected given the number of individuals of this genus I’ve bait-trapped over the years at dolomite glades in Jefferson Co.). I was also surprised and happy to see the Knulliana, which I think is actually an undescribed species—they looked very similar coming out of the trap as the much more abundant Eburia, so I’ll have to check the collected specimens (both from here and other locations) more closely to see if there are other specimens that I’ve counted as Eburia. As expected, the ethanol-only trap again had far fewer numbers and lower diversity of the same species plus a single Lepturges sp. (same as the previous location). Both traps were re-baited with the same bait, though I started to doubt the value of continuing the bait comparison when ethanol/red wine already seemed far superior to ethanol-only. In the end, I decided to continue the continue the comparisons to allow for the possibility of other species later in the season that might show a different preference (and to allow publication of the study data even if this is not the case). Disappointingly, sweeping herbaceous vegetation in the xeric igneous prairie around both traps produced nothing.

Trimerotropis saxatilis (lichen grasshopper) on lichen-covered rhyolite exposures in xeric igneous prairie.

Hughes Mountain Natural Area

Again, like the previous spots today, the area looks lush and got more rain last night, but the numbers of beetles in the ethanol-red wine trap was not quite as overwhelming as in other locations. Nevertheless, there still a lot and two very good species : Sphenostethus taslei and Purpuricenus paraxillaris! I’ve only seen the former once before in a bait trap at Victoria Glades, and I’m always happy to see more individuals of the latter (the first new species I ever discovered!). Other beetles in the trap were Eburia quadrigeminata, an undetermined elaphidiine, Strangalia luteicornis, Typocerus velutinus, Xylotrechus colonus, Cotinis nitida, Euphoria fulgida, and various elaterids (including many tiny ones that I didn’t collect). In what is becoming a familiar refrain, the ethanol-only trap again had lower numbers and diversity, with Cnestus mutilatus being the only species not also caught in the ethanol/red wine-baited trap. Sweeping herbaceous vegetation, both in xeric igneous prairie around Trap A and dry post oak woodland around Trap B, produced nothing. This was the last location I was able to visit this day because of the unexpectedly large amount of time it took to sort the large numbers of beetles in many of the traps, so I saved the last two locations for the next day.

Sunset at Hughes Mountain.

St. Joe State Park

Like other areas on this side of the state, the vegetation was lush and got even more rain two nights ago. I’d been very anxious to see this spot, as there is lots of Sideroxylon lanuginosum (gum bumelia) in the small xeric dolomite glades remnants and extensive surrounding areas of restored dry post oak woodland, which promise to yield Plinthocoelium suaveolens. Unfortunately, the ethanol/red wine trap was completely missing—not fallen or pulled down by raccoons, but more likely stolen by humans (I probably should have placed this trap further away from the nearby paved bicycle trail). This is first case of trap theft I’ve experienced in a long time. If they had to steal one of the traps, I wish it would have been the ethanol-only trap rather than the ethanol/red wine trap—the former haven’t produced nearly as well as the latter, thus the loss of data would have been less. The ethanol-only trap had only a few beetles representing E. quadrigeminata, Batyle suturalis, Dichelonyx sp., Elateridae, and Cnestus mutilatus. I did not replace the ethanol/red wine trap, but instead re-baited the ethanol-only trap with ethanol/red wine. This means I will need to drop this location from the bait comparison analysis, but I wanted to give myself the best chance for trapping Plinthocoelium, and ethanol/red wine is clearly the better bait for accomplishing this. Sweeping herbaceous vegetation in the xeric dolomite prairie near Trap A and dry post oak woodland around Trap B produced nothing, but I did see an impressive Mydas tibialis (golden legged mydas fly) visiting flowers of Eryngium yuccifolium (rattlesnake master) in the dry post oak woodland.

Mydas tibialis (golden legged mydas fly) visiting flowers of Eryngium yuccifolium (rattlesnake master) in dry post oak woodland

Salt Lick Point Land & Water Preserve

Pflasterer’s Glade at Salt Lick Point Land & Water Preserve.

This is the only locality in Illinois that I placed traps—specifically because last fall I saw good numbers of Sideroxylon lanuginosum (gum bumelia), host for Plinthocoelium suaveolens (bumelia borer), and one in particular that showed signs of active infestation. Plinthocoelium has not been recorded from Illinois, so a voucher specimen will be important for publishing the record. The area apparently had gotten decent rains to this point, with vegetation in the woodlands looking lush and that in more exposed situations showing a tinge of wilt. Anticipation turned to disappointment when I looked inside the ethanol/red wine trap (placed right next to the infested bumelia tree) and saw lots of Eburia quadrigeminata and elaphidiines but only one Neoclytus acuminatus and no Plinthocoelium or other cerambycids. The infested teee was again churning out frass at the base of the trunk, so the infestation remains active—I will consider bringing screen with me next time so I can place a skirt around the base of the tree and check periodically for the adult once it emerges. In addition to the cerambycids noted above, the ethanol/red wine had Cotinis nitida, Enoclerus sp., Elateridae, Cnestus mutilatus, and other miscellaneous beetles. The ethanol-only trap had lower numbers of beetles, with Neoclytus scutellatus and Lepturges sp. being the only longhorns not represented in the ethanol/red wine trap. Interestingly, the trap also contained a few Buprestidae, which have been scarce in traps to this point (Actenodes sp.—prob. A. acornis or A. simi, Anthaxia sp.—prob. A. cyanella or A. dichroa, and Agrilus sp. Both traps were re-baited with ethanol-red wine to increase the likelihood of picking up P. suaveolens during the remainder of this season, and I will accordingly drop this locality from the bait comparison analysis (along with St. Joe State Park due to the ethanol-red wine trap being stolen) as well, thus limiting the analysis to the remaining 10 locations in Missouri.

Freshly-ejected frass at the base of a small living Sideroxylon lanuginosum (gum bumelia) tree indicates active larval infestation by Plinthocoelium suaveolens (bumelia borer)—a species not yet formally recorded from Illinois.

Sweeping herbaceous vegetation in the dry hilltop prairie near the traps produced nothing, but I did find a dead Lucanus capreolus (reddish brown stag beetle) dead on the trail though the dry-mesic upland deciduous forest leading up the them.

Lucanus capreolus (reddish brown stag beetle) female. Found dead on trail through dry upland deciduous forest.
Silphium integrifolium (prairie rosinweed) in dry upland deciduous forest.

©️ Ted C. MacRae 2022

2022 Oklahoma Insect Collecting Trip iReport

Welcome to the 11th “Collecting Trip iReport”; this one covering a very short (4 days) trip to northwestern Oklahoma on May 3–7, 2022. My collecting partner for this trip was long-time friend and hymenopterist Mike Arduser. Mike is one of the best natural historians that I know and, like me, has a special love for the often overlooked beauty of western Oklahoma and its fascinating insect fauna. It had been 13 years, however—too long, in my opinion, since our last joint field trip when we sampled the bee (Mike) and beetle (me) fauna at The Nature Conservancy’s Four Canyon Preserve in Ellis Co. Thus, I was happy for the chance to once again spend some time in the field with such a knowledgeable naturalist in an area we that both know and love.

As with all previous “iReports” in this series, this report is illustrated exclusively with iPhone photographs (thus the term “iReport”). Previous iReports in this series include:
2013 Oklahoma
2013 Great Basin
2014 Great Plains
2015 Texas
2018 New Mexico/Texas
2018 Arizona
2019 Arkansas/Oklahoma
2019 Arizona/California
2021 West Texas
2021 Texas/New Mexico/Arizona


Day 1 – Gloss Mountain State Park (Major Co.)
It took most of the day to get here—Tulsa threw us a couple of obstacles in the form of a construction-mediated wrong turn and a motorcycle engulfed in flames. I’ve been to Gloss Mountain a number of times, but never this early in the season. Skies were sunny (unlike St. Louis when we left this morning), but temps didn’t get much above 60°F and even dropped down into the upper 50s before we finished up at sunset.

Gloss Mountain State Park – early evening view from atop the mesa.

Surprisingly, despite the earliness of the season and cool temps, beating was quite productive. Working the low areas around the parking lot, I beat a fair number and diversity of beetles and hemipterans—mostly chrysomelids—but only a single Agrilus sp. off of Prosopis glandulosa.

This must be Oenothera macrocarpa (bigfruit evening primrose, Ozark sundrop, Missouri evening primrose), though it looks very different from populations south of St. Louis.

I knew there were other trees, principally Celtis reticulata (net-veined hackberry) and Sapindus drummondii (soapberry), on top of the mesa and wanted to see if anything was on them. Bingo! Even before reaching the top, I beat a few Agrilus (several spp.) from the Celtis, and up on top I beat quite a few more off the same. There were also additional mesquite trees up top, off which I again beat a single Agrilus sp. along with a few other things, notably a series of ceresine treehoppers. The Sapindus was just starting to leaf out, and I found nothing by beating them other than a single ceresine. A notable find was the pile of larval frass of Plinthocoelium suaveolens (bumelia borer) at the base of a living Sideroxylon lanuginosum (gum bumelia) tree—a sure sign of active infestation by a beetle I have yet to formally record from this place.

Frass at the base of a small Sideroxylon lanuginosum (gum bumelia)—evidence of an active larval infestation by Plinthocoelium suaveolens (bumelia borer).

On the way back down from the top, we hit the sunset perfectly as it “touched” a peak in the foreground! Despite my success here this evening, Mike saw no bees of interest on the few flowers that were found due to the cold temps and chilling winds, so tomorrow we will continue west hoping for warmer conditions on the western edge of Oklahoma.

Dramatic sunset “landing” on a small foreground peak.
The opportunity lasted for only a minute!

Back in town, we searched for an open sit-down restaurant—fruitlessly because of the late hour—and ended up with a mediocre breakfast burrito from a fast food shop I’ve never been to before. The local Buick dealership, however, with its 1950s neon lights shining brightly in the night sky, was a taste of Americana that makes these trips so enjoyable. Life on the road!

Jensen’s Buick, Fairview, Oklahoma.

Day 2 – Black Mesa State Park (Cimmaron Co.)
Welp! We awoke this morning to cold temps (low 60s), thick fog, and low hanging clouds, and the forecast for the area showed essentially no improvement through at least the day. Our plan had been to hit a spot about an hour southwest before heading back north to Beaver Dunes State Park, but the forecast for both those areas also was cold and wet. It was not until we looked at the forecast for Black Mesa—our last planned stop of the trip and a 4½-hour-drive to the west—that the forecast seemed to be in our favor, so we decided to blast on out there. We figured we would get there at about 2:00 pm and could spend the rest of the day there collecting, camp there tonight, and start heading back east tomorrow (assuming the forecast improved for the areas we missed).

Pronghorn antelope (Antilocapra americana).

Wrong! When we got there, it was not only cloudy and cold, but dry as a bone! Even if it had been sunny with warmer temps, there still would not have been any insect activity to speak of. The leaves of oaks and hackberries in the area were just barely starting to break bud, and the only flowers we saw at the park were a large willow in full bloom—but not a single insect visiting them. Knowing that there was no other place where conditions were better that we could drive to within the next couple of hours and collect for at least a short time, we instead decided to make it a hiking day and hike the High Point Trail at nearby Black Mesa Nature Preserve.

Black Mesa Nature Preserve (Cimmaron Co.)
When we arrived and looked at the signage, we learned that the hike to the oracle at the official high point would be a more than 8-mile hike! Just reaching the top of the mesa itself would be a more than 3-mile-hike, with the high point another mile on top. Not knowing if we had the appetite for such a distance (or time to do it before sunset) and with the wind cold and biting, we started out anyway and gave ourselves permission to turn around at any point if we felt like it.

View along High Point Trail.
View along High Point Trail.
Berlandiera lyrata (lyreleaf greeneyes).

Nevertheless, we persevered. We checked the cholla (Cylindropuntia imbricata) along the way hoping to see Coenopoeus palmeri (one of the cactus longhorns, which I’m not sure has been recorded from Oklahoma) or at least one of the more widespread Moneilema species, but none were seen (nor really expected). The trail up the side of the mesa was steep and spectacular, and the trail atop the mesa was surreal—especially given the cold winds and low-hanging clouds. Eventually, we made it to the official high point and enjoyed the fun facts carved into each side of the granite obolisk marking the spot.

View along High Point Trail about halfway up the climb.
Juniperus monosperma (one-seed juniper).
Oklahoma High Point obolisk.
Oklahoma High Point marker.
Mike (right) and me at Oklahoma High Point.

Coming back down was not much easier than going up, the steepness of the trail jamming my toes into the toe box of my new hiking boots (which performed admirably!), but I did find an insect—a largish black weevil torpidly crawling on the trail. Even on the relatively level lower portion of the trail once we got there was difficult, our legs really starting to feel the miles now. As we hiked the last mile back to the car, the temperature continued to plummet as it started to sprinkle, turning to rain soon after we reached the car and then heavy rain as we headed down the highway back to the east. The irony of the situation—rain coming to a parched landscape just when we are ready to leave—did not escape us. We’ll spend the night in Boise City and hope for a better forecast tomorrow!

Descending the upper slope on the High Point Trail.

Day 3 – Beaver Dunes State Park (Beaver Co.)
Temps were down in the mid-40s when we awoke this morning, but skies were sunny and we were heartened by a promising forecast of continued sun and highs in the low to mid-60s. Our first destination—Beaver Dunes—was a relatively short 2-hour drive further east, and when we arrived sunny skies still prevailed. Unfortunately, temps still hovered in the mid-50s with a biting wind that made using the beating sheet difficult to impossible.

Dunes at Beaver Dunes State Park.

That said, I managed to beat a fair series of Agrilus spp. (probably mostly one species) and a few other beetles off living Celtis reticulata (net-veined hackberry) dotting the roadside along the entrance to the Picnic Area. Under the main group of hackberries I noticed new growth of Cucurbita foetedissima (buffalo gourd) along with last year’s dead stems. I’ve never collected Dorcasta cinerea (a longhorn beetle that utilizes buffalo gourd as a larval host), so I began splitting open the old stems to see if I could find unemerged adults. I didn’t, but what I did note inside the stems was evidence of boring by some insects and, eventually, the tiniest little scolytine bark beetles that I’ve ever seen. They were always found right at the node, usually in pairs (perhaps male and female?), and I ended up collecting a series of about a dozen specimens from two different stems.

Tradescantia occidentalis (prairie spiderwort, western spiderwort).

Also in the main group of hackberries, I noticed a dead branch hanging from the tree, which had fallen but gotten snagged on a lower branch to remain off the ground. The branch was obviously infested and showed a few emergence holes indicative of both buprestids and cerambycids, and when I broke into it I found two unemerged adult Agrilus (different species), which caused me to cut and bundle the branch to being back for rearing. At the entrance, I went to examine the stand of yellow flowers that greeted our arrival, determining them to be Pyrrhopappus pauciflorus (smallflower desert-chicory, Texas false dandelion). While I was on the ground photographing the flowers, I noticed a red and black hister beetle that proved to be Margarinotus bipustulatus—aptly named considering the two red maculations on the elytra. I also noticed a couple of tiger beetle larval burrows in the hard-packed sandy soil and found a long, thin plant stem to “fish” the larvae out. I managed to snag the larva in one of the burrows, which I believe is Tetracha carolina (Carolina metallic tiger beetle) by virtue of the thin white margin around the prothorax and the open habitat in which the larval burrow occurred. If this is true, then it is a second instar because it is slightly smaller than a typical Cicindela sp. third-instar larva.

Pyrrhopappus pauciflorus (smallflower desert-chicory, Texas false dandelion).
Pyrrhopappus pauciflorus (smallflower desert-chicory, Texas false dandelion).
Margarinotus bipustulatus (family Histeridae).

Afterwards, I went over to the dunes to see if Mike had found anything, but temps were still too cold to see anything flying. He did, however, show me an interesting stand of Penstemon that he’d found and that we determined to be P. fendleri (Fendler’s penstemon). The plants were all on the north side of the dune in apparently protected spots, and I noted that on iNaturalist our observation was the northernmost record for the species (save one suspicious, disjunct Colorado record).

Penstemon fendleri (Fendler’s penstemon).
Penstemon fendleri (Fendler’s penstemon).

On the way back to the car, I beat a few more beetles off living Celtis reticulata. By now, we’d seen all we needed to see here and decided to head southeast to one of the Brachys barberi locations (that were the reason for this trip in the first place).

5 mi E of Harmon (Ellis Co.)
This

Recently, another coleopterist collected Brachys barberi—more typically a southwestern species—on Quercus harvardii (shinnery oak) at this spot. I’ve not managed to find the species myself yet, and as it was collected on May 3rd last year I hoped the timing would be right. Quercus havardii dominated the landscape at this spot, mostly as thick stands of low-growing shrubs but also as a copse of small trees.

Quercus havardii (shinnery oak, shin oak, Havard oak).

At first, I swept the lowest-growing plants, collecting a variety of mostly chrysomelids and curculionids and even one Agrilus sp., before moving to beating along the sunny edges of the patches of taller shrubs and collecting similar species (but no Agrilus sp.). Just to the north, I noticed a stand of individuals tall enough to be considered trees (presumably a clonal stand) and began beating them. Immediately I began collecting not only the chrysomelids and curculionds that I was collecting before, but also several Agrilus spp. and what must be Agrilaxia texana—a species represented in my cabinet by just two specimens that I collected in northeastern Texas way back in 1984.

I worked nearly the full perimeter of the copse, noticing that most of the beetles were being collected only on the south-facing sunny (and leeward) side. When I was just about ready to call it quits, a much larger black and yellow beetle landed on the sheet. For an instant I thought it was a lycid, but it moved characteristically like a longhorned beetle, and I quickly realized that I had collected Elytroleptus floridanus—a quite rare southeastern U.S. species that I have only seen once before when I reared a single individual from dead oak that I collected in the Missouri bootheel (and representing a new state record). I wasn’t sure the species had ever been recorded from Oklahoma, so I found Gryzmala’s revision of the genus online and saw that it had been previously recorded from the state—but all the way over on the east side near the border of Arkansas. All records from Texas as well are from the eastern side of the state, so today’s capture appears to represent a significant northwestern extension of the species’ known geographic range by about 300 miles!

Sadly, I never saw Brachys barberi, but collecting Elytroleptus floridanus (in Oklahoma!) was a pretty good consolation prize.😊


Day 4 – Prologue (“Good to Go” coffee shop)
We awoke to bright sunny skies, and though a tad chilly it was still warmer than the previous mornings and with a good forecast to boot! It would take about an hour to drive to the day’s collecting spot—the one and only Gloss Mountain State Park (where we visited briefly a few days ago to start the trip), but not until after an unexpected and hilariously bizarre experience at a coffee shop in town called “Good to Go”.

“Good to Go” coffee shop lounge.

Mike was the first to notice the velociraptor in the lounge—saddled up for a ride! Okay, that’s cute. Then he noticed the sign on the outdoor display that read “Stegosaurs roamed the Earth about 5,000 years ago.” At first I thought, okay, they’re a little confused on the timeline, but what they’re trying to say is that dinosaurs lived a long time ago.

“Stegosaurs roamed the Earth about 5,000 years ago.”

Then I noticed a granite plaque in the background that clearly read “The Holy Bible”, and it dawned on me that we had entered a creationist’s den! Had we not already ordered our coffee, I might have surreptitiously tiptoed my long-haired hippy butt out of there before somebody pointed at me and began slowly chanting “Lucifer!”

Apparently this is overwhelming evidence that humans saw living dinosaurs.

Once we were outside the shop, our coffee secured and the need for hushed tones no longer muffling our reactions, we took a quick walk with the dinosaurs to admire their seeming scientific accuracy. I was impressed with the T. rex in particular, it’s body axis realistically horizontal with the tail straight and strong—not the lumbering, upright, tail-dragging version that I learned about as a kid. At least they were accepting some of the current body of scientific evidence on dinosaurs and ignoring only that dealing with their age—or so I thought…

A remarkably scientifically accurate rendition of T. rex with the more recently advocated horizontal posture.

The stegosaur as well appeared to be fairly accurately rendered, its tail also straight and strong and a youngster trailing closely behind, until I noticed something atop the adult—an angel riding it! ‘God’s creatures big and small’, I guess.

Note the angel riding the stegosaur!

The coup de grace was the information plaque behind the stegosaur. Rather than providing information on dinosaurs, I was instead treated to a barrage of hilariously unsupported claims advocating the idea that humans and dinosaurs once lived together. Each “factoid” on the plaque was more bizarre and quotable than the one before. Did you know that the adult stegosaur probably died 4,000 years ago in the Great Flood, but that the baby—happily—likely survived by getting a ride on the Ark with Noah! And all that scientific evidence that pinpoints the Cretaceous extinction to 65 million years ago? Apparently it has merely been fabricated as part of a global conspiracy because scientists just don’t want to agree with the Bible. I just about lost it, however, when I reached “It is uncertain if humans ever rode Dinosaurs, but there is overwhelming evidence that humans saw living dinosaurs.” I mean—What?!

I don’t even know where to begin!

Our unplanned morning entertainment now done, we hit the road for our next—and final—collecting spot for the trip.

Gloss Mountain State Park (Major Co.)
We arrived at about 10 am with a plan to spend the rest of the day there—whether the collecting was good or bad, this would be our final stand. We hiked up to the mesa, stopping at an accessible spot about halfway up to work the trees (me) or set out pan traps (Mike). Beating the Celtis reticulata (net-veined hackberry) yielded a similar assortment of beetles as last time—a couple of Agrilus spp. along with the occasional chrysomelid or curculionoid and a few other beetles, and the same was true with Prosopis glandulosa (mesquite), with the exception that I did not find any Agrilus this time.

Gloss Mountain State Park – view west from atop gypsum-capped red clay mesa.

Atop the mesa, I decided to do an entire perimeter hike—something I’ve always wanted to do but never actually accomplished. The idea was to beat all of the C. reticulata, P. glandulosa, and Sapindus drummondii (soapberry) that I could find in an effort to “leave no stone unturned” in my quest for beetles. Soon after starting out, I saw a nice Pasimachus elongatus ground beetle running across the mesa top and “forced” it to cooperate for photos by pinning a hind tarsus to the ground with my finger tip (barely visible in the upper left side of the photo). I collected it, as well as another that I saw a short distance away, and then proceeded with the beatings! Beating the C. reticulata was quite productive, with perhaps three Agrilus spp. and numerous other beetles being collected off of nearly every tree that I beat. Beating P. glandulosa also was productive for various beetles, though again no Agrilus were encountered. The biggest surprise came when I started beating S. drummondii, most of which were still in the earliest stages of leafing out. I got nothing from most of the trees (the majority of which were clustered in a small copse near the front of the mesa), but in the back part of the cluster were a couple of trees with noticeably more foliage—beating them yielded perhaps a dozen Agrilus limpiae, a soapberry specialist that I haven’t seen in numbers since 1986 when I collected a series on soapberry in south-central Kansas.

Gloss Mountain State Park – view west from atop gypsum-capped red clay mesa.
Pasimachus elongatus (family Carabidae) atop gypsum-capped red clay mesa.

I rarely get anything beating Sideroxylon lanuginosum (gum bumelia), but I beat most of the trees that I saw anyway and collected one cryptocephaline chrysomelid and two curculionoids. A single Eleodes hispilabris (apparently on its last leg) was seen near the north end of the mesa, which I photographed and collected, and on the way back I encountered a small patch of Sphaeralcea coccinea (scarlet globemallow) in bloom, from the flowers of which I collected a few small melyrid-type beetles and a small halictid bee for Mike. Also on the north part of the mesa I saw a young eastern collared lizard (Crotaphytus collaris), who posed just long enough for me to get off a shot before blasting away from my approaching lens.

Eleodes hispilabris (family Tenebrioindae) atop gypsum-capped red clay mesa.
Sphaeralcea coccinea (scarlet globemallow) atop gypsum-capped red clay mesa.
Crotaphytus collaris (eastern collared lizard) atop gypsum-capped red clay mesa.

Throughout the hike atop the mesa I kept my eye out for “new-to-me” plants (of which there are many), finding for the first time Toxicodendron rydbergii (western poison ivy) and blooming individuals of Chaetopappa ericoides (rose heath). Physaria gordonii (Gordon’s bladderpod)—a relative of the federally threatened P. filiformis (Missouri bladderpod)—was blooming abundantly atop the mesa. At this point, Mike and I rejoined and relayed to each other our more notable findings. For Mike’s part, he had seen a couple of cacti that I had missed—Escobaria missouriensis (Missouri foxtail cactus) and Echinocereus reichenbachii perbellus (black lace cactus)—and took me to the spots where he had seen them. While retracing our steps, we also found Gaillardia suavis (pincushion daisy, perfumeballs) and the strikingly beautiful Penstemon cobaea (cobaea beardtongue, prairie beardtongue, foxglove penstemon).

Toxicodendron rydbergii (western poison ivy) atop gypsum-capped red clay mesa.
Chaetopappa ericoides (rose heath) atop gypsum-capped red clay mesa.
Physaria gordonii (Gordon’s bladderpod) atop gypsum-capped red clay mesa.
Escobaria missouriensis (Missouri foxtail cactus) atop gypsum-capped red clay mesa.
Escobaria missouriensis (Missouri foxtail cactus) atop gypsum-capped red clay mesa.
Echinocereus reichenbachii perbellus (black lace cactus) atop gypsum-capped red clay mesa.
Echinocereus reichenbachii perbellus (black lace cactus) atop gypsum-capped red clay mesa.
Gaillardia suavis (pincushion daisy, perfumeballs) atop gypsum-capped red clay mesa.
Penstemon cobaea (cobaea beardtongue, prairie beardtongue, foxglove penstemon) atop gypsum-capped red clay mesa.
Penstemon cobaea (cobaea beardtongue, prairie beardtongue, foxglove penstemon) atop gypsum-capped red clay mesa.

By this time, I had been on the mesa top for five hours, and even though temperatures were mild (mid-70s) I desperately needed food and water. Mike, for his part, had also had a wildly successful day with bees, capturing many at the flowers and many more in the various pan traps (both in top and halfway up the slope). I descended the steep slope with its mixture of metal steps, cut rock, and wooden planks and enjoyed a quick feast of sardines and Triscuits (a decades-long bug-collecting-trip staple) washed down with Gatorade before getting back to work on the mesquite around the parking lot. I was committed to trying to find Agrilus on the plants—a single individual of which I’d beaten from the plants three days earlier, and after beating several plants and seeing none (but collecting a great number of clytrine and cryptocephaline chrysomelids along with other insects) I finally found one! I continued to work the trees and collect primarily chrysomelids, but no more Agrilus were seen. I am hopeful that it will be a southwestern species not currently known from Oklahoma—a situation I have found with several other Prosopis-associated beetles in this part of northwestern Oklahoma.

Gloss Mountain State Park – view north from atop gypsum-capped red clay mesa.
Gloss Mountain State Park – history of the name.

I hadn’t intended to work any additional Prosopis beyond the road into the parking lot, but there were a few particularly large trees along the front of the park next to the highway rest stop. The first one I beat yielded a very large cryptocephaline that I had not seen on any of the other Prosopis, so I continued beating them and collected a nice series along with a few other clytrines, pachybrachines, and curculionoids. At the furthest point west, I recalled having seen during a previous visit a western diamondback rattlesnake a bit further to the west, so I continued to the spot hoping to see another. No such luck, so I tiptoed through the tall grass back to safety and made my way back to the car to wrap up seven and a half hours of collecting on a spectacular day—sadly, the last of the trip!


Epilogue
This trip was just a warm-up. In just over one week, I will head out again—this time to western Texas and southern Arizona for sure, and maybe elsewhere depending on how things go. At three weeks, it will be the longest collecting trip I’ve done since I went to South Africa in 1999 and Ecuador 10 years before that. I’m also looking forward to meeting up with a number of other coleopterists at various points during the trip—Jason Hansen, Joshua Basham, and Tyler Hedlund in Texas, and Norm Woodley and Steve Lingafelter in Arizona. If there is time, I may stop off at a place or two in northeastern New Mexico and at Black Mesa on the way back. Look for an iReport on that trip sometime in early-mid June!

©️ Ted C. MacRae 2022

2021 Texas/New Mexico/Arizona Insect Collecting Trip iReport

This is the 10th “Collecting Trip iReport”; this one covering 13 days of collecting in western Texas, southern New Mexico, and southeastern Arizona July 19–31, 2021. This trip was a “two-parter”—the first week with frequent field mate Jeff Huether (our seventh joint collecting trip) as we made our way from western Texas through southern New Mexico and into southeastern Arizona on our way to a memorial celebration for Jim Wappes at the home of Steve Lingafelter and Norm Woodley (for which I took the day off from collecting), and the second week visiting various locations in the “Sky Islands” of southeastern Arizona with several other entomologists.

As with all previous “iReports” in this series, this report is illustrated exclusively with iPhone photographs (thus the term “iReport”). Previous iReports in this series include:
2013 Oklahoma
2013 Great Basin
2014 Great Plains
2015 Texas
2018 New Mexico/Texas
2018 Arizona
2019 Arkansas/Oklahoma
2019 Arizona/California
2021 West Texas


Day 1 – Monahans State Park, Texas
This was my first stop on the previous trip back in late April and early May, and what a difference a couple of months with good rains makes—dry as a bone then but bursting with a great variety of wildflowers now. Like last time we stopped at the Shin Oak Picnic Area first, and almost immediately Jeff got an Acmaeodera gibbula on living Prosopis glandulosa (mesquite). I started beating the trees and got a good series of that species along with a good series of an Actenodes sp. (prob. A. mendax). There were also some mesquites that had been killed (apparently by herbicide), and when I started beating them I got several more A. gibbula and one Paratyndaris sp. It went from blazing hot when we arrived to raining about an hour later, and for a while after the rain moved through it stayed cloudy and quite comfortable. Eventually we decided to look for another spot with more mesquite to beat.

Monahans Sandhills State Park with rain moving in!
Poecilanthrax effrenus (family Bombyliidae) on flower of Thelosperma megapotamicum (rayless greenthread).
Acanthochalcis nigricans (family Chalicididae) female looking to oviposit on dead Prosopis glandulosa (mesquite).
Palafoxia sphacelata (othake).
Palafoxia sphacelata (othake).
Asclepias arenaria (sand milkweed).
Asclepias arenaria (sand milkweed).
Penstemon ambiguus (gilia beardtongue).

We found an area closer to the entrance (Equestrian Area) with lots of mesquite and also sunflowers, which Jeff was interested in looking at to search for meloids (blister beetles). Before I even reached the first mesquite I saw an Acmaeodera sp. (maybe A. obtusa) sitting on the flower of Thelosperma megapotamicum (rayless greenthread) and later found another plus one A. immaculata. Off the mesquite I beat just one Actenodes sp. prob. mendax and a few treehoppers, while another and a Chrysobothris sp. got away (it was by now quite hot and they bolted!). I continued beating mesquite but just wasn’t seeing anything, so we decided to take a look at the area around the main dunes and another picnic area.

Helianthus petiolaris (prairie sunflower).

At the Pump Jack Picnic Area, we saw a lot of Thelosperma megapotamicum (rayless greenthread) in bloom and decided to check the flowers for Acmaeodera. We each got a nice series of what appear to be A. obtusa and A. immaculata. Also, I finally found a single Acmaeodera immaculata on the flower of Hymenoppapus flavescens (collegeflower), which I’d been looking at all day thinking it must be a good Acmaeodera flower host. On the way back to the vehicle, I scared up a cicada that had been singing on a nearby plant—I’d been hearing them all day but assumed they were grasshoppers or katydids. I listened for another and saw it perched on the stem of Mentzelia nuda (bractless blazingstar), and netted it—a fine male, smallish and with a very white venter. I wanted to find one more—catching another A. gibbula in flight, and saw one singing in a mesquite tree. This time I took some photos of it (working carefully not to alarm it) and then hand-caught it (later identified as Diceroprocta texana). A nice end to the visit.

Male Diceroprocta texana singing on Prosopis glandulosa (mesquite).
Mentzelia nuda (bractless blazingstar).
Mentzelia nuda (bractless blazingstar).

After dinner, we returned to Shin Oak Picnic Area for night collecting. I setup my new Mercury-vapor (MV) station (first time using the gas-powered generator and tripod—slick setup) and my two ultraviolet (UV) stations. Jeff set out three prionic acid lures. Large numbers of Polyphylla monahanensis (which came mostly to the UV stations) and P. pottsorum (which came mostly to the MV station), but otherwise few beetles showed up and not a single longhorn. Other insects were also limited mostly to large numbers of ground-nesting bees and several big grasshoppers. No Prionus came to the lures—not surprising since we are in the tail end of the season for P. arenarius (April to July) and too early for P. spinipennis (mostly August). I’ve only gotten a few P. monahanensis and P. pottsorum before now, so it’s nice to have good series of each, but I would have preferred to collect some longhorns.

Blacklights humming as dusk settles over the dunes.
Polyphylla pottsorum at the MV sheet.

Day 2 – Toyahvale, Texas
We’re on our way to the Davis Mountains, and along the way I decided to stop at the “Agrilus cochisei” spot we found (on a tip from Jason) back during the April trip. I swept the roadsides—not just the host plant (Artemisia occidentalis, western ragweed) but a variety of other plants in bloom but did not find any A. cochisei. I did collect a few meloids (which were on Solanum eleagnifolium) and Zygogramma leaf beetles but nothing else.

Proboscidea parviflora (family Martyniaceae)—doubleclaw or red devil’s-claw.

A bit further down the road from the last stop, we noticed this memorial to the many horses that have been transported along this highway on their way to slaughter.

Davis Mountains, Ft. Davis, Texas
Another roadside stop for one of the places where I collected during last April’s trip. I was hoping we would not be too late for Acmaeodera—there were plenty of plants in bloom but we did not see any. Senegalia greggii (formerly Acacia greggii, commonly called cat-claw acacia) and Vachellia constricta (formerly Acacia constricta, commonly called whitethorn acacia) were both in bloom, and off the former I got the obligatory Stenaspis solitaria male/female pair as well as a Lampetis drummondii, but I collected nothing off the latter. Tried for a couple of cicadas and missed ‘em both!

Stenaspis solitaria male on flowering Senegalia greggii (cat-claw acacia).
Lampetis drummundii captured on flowering Senegalia greggii (cat-claw acacia).
Eurema nicippe (sleepy oranges) on yellow asteraceous flower.

Davis Mountains, Boy Scout Rd, Texas
This looked like a good spot, with water in the creek and lots of butterflies flying around. Beating, however, yielded nothing but lots of lep larvae. It seems we are in the mid-summer lull—too late for spring things, but too early for late summer-fall species. I think we’ll try some higher elevations and see what it is like.

Argemone aenea (golden prickly poppy).
Argemone aenea (golden prickly poppy).
Argemone albiflora (white prickly poppy).
Argemone albiflora (white prickly poppy).

We were headed back towards the highway when I spotted a stand of Thelosperma megapotamicum along the road. We got out so I could take a look at them and immediately encountered this Texas horned lizard who seems to be saying “WTF?!” I swept through the Thelosperma and picked up two Batyle sp., one Enocleris sp., and a couple of species of meloids. There were lots of other plants in bloom, too, including several that are typically attractive to beetles such as Sphaeralcea and Ratibida. However, nothing was seen on them, further reinforcing our desire to go to higher elevations to see if that would improve the collecting.

Phrynosoma cornutum (Texas horned lizard).

Davis Mountains, Madera Canyon, Texas
We wanted to get up to higher elevation to see if that might improve the insect collecting. It is strange—the Davis Mountains are greener than I’ve ever seen them, yet there are almost no insects, no flowers. Jeff and I were wondering if the deep freeze Texas experienced this past winter might have knocked out insect populations. We beat along the way but just we’re not seeing anything on the beating sheets. We hiked our way up to the overlook, and up there I ran into a few species of tenebrionids running along the trail and doing their famous “headstands” when we disturbed them. On the way back down I saw a few large, red and black clytrine chrysomelids on what I take to be a fall-flowering helianthoid aster (old flowering stalks were 5–6 feet tall), so I picked up a few for Shawn (my scope of insects that I’ll collect expands greatly when I’m not finding anything in the groups that I study). I think we’ve had it with the Davis Mountains, and tomorrow we’ll travel further west and try our luck around Fabens.

Lower part of the Madera Canyon Trail.
The trail at upper elevations.
A small Eleodes sp. doing the headstand.
The larger Eleodes carbonaria doing the headstand.
Same individual as in the previous photo.

Day 3 – Van Horn, Texas
Just a quick stop along the highway when we saw a variety of plants in bloom. Things are different at these lowers elevations compared to the Davis Mountains. Sweeping yielded a number of Agrilus sp. (vittate) and two Agaeocera gentilis—I suspect they were on the Sphaeralcea angustifolia (narrow-leaved globemallow), along with an Acmaeodera sp. and assorted other beetles. On a much taller globemallow I found two Tylosis jiminezi (male/female) perched on the foliage—a first for me! I did a little more sweeping further to the south but came up with only a few melyrids and a blister beetle (Epicauta segmenta). Nice stop!

Tylosis jiminezi on Sphaeralcea angustifolia (narrowleaf globemallow).
Sphaeralcea angustifolia (narrowleaf globemallow).
Sphaeralcea angustifolia (narrowleaf globemallow).

San Felipe Park, El Paso Co., Texas
When we first arrived, we were not at all optimistic—it looked like it hadn’t rained in years. However, the Larrea tridentata (creosote) and Prosopis glandulosa (mesquite) were both in full bloom, so we began looking about. Almost immediately I saw Stenaspis solitaria in the mesquites, which in itself is not exciting, but two of the first three I saw were the form that has the distinctly reddish-brown pronotum, which I’ve never seen before. I saw a couple more and tried to get photographs, but they were too skittish. As I searched for them, I caught one Aethecerinus latecinctus (a second got away), one Plionoma sp. (not sure if it’s suturalis or rubens), one Chrysobothris sp. (prob. C. octocola) and one Acmaeodera gibbula. The real fun began, though, when I walked by a creosote and saw a Gyascutus planicosta (should be subsp. obliteratus in this area) take flight. I tracked it to see where it landed, caught it, and then put most of my effort into getting a decent series of individuals. I succeeded, but it took more than four hours with the heat maxing out at 96°F! In addition to the Gyascutus, Jeff was quite excited to see the bright green and orange blister beetle Eupompha fissiceps abundant on the creosote in mating pairs and feeding on the petals of the flowers. I finished off the blister beetle fun by finding Cysteodemus wislizeni (black bladder-bodied meloid) crawling in the sand, which, despite the common name, was decidedly bluish.

The author looks out over the vast creosote/mesquite scrub.
Eupompha fissiceps on Larrea tridentata (creosote).
Eupompha fissiceps feeds on Larrea tridentata (creosote) as “Mr. Meloid” (Jeff Huether) looks on.

After going into town to restock on supplies and catch some dinner, we returned to the park to do some lighting and night collecting. The moon is almost full, which generally puts the kibosh on longhorns coming to the lights, but we decided to try anyway because of the high amount of activity during the day in a rarely-visited location. I set up the Mercury-vapor (MV) light only and skipped going through the trouble to put up the ultraviolet lights also. I also wanted to beat the mesquite since I didn’t have much chance during the day, spending most of my time hunting Gyascutus with an aerial net, and once I got the MV setup going I started whacking the mesquite. Almost immediately, I got three Aethecerinus latecinctus, which came off the first two plants I beat. This motivated me further and caused me to commit to beating for the next hour or so—never even getting a single beetle of any kind! By then the lights had been going for awhile, and I was pleased to see several Derobrachus hovorei (palo verde root borer) crawling on the sand near the light. Even though it is a common species, I’ve not seen many myself, so I was happy to have a nice series to take up beaucoup room in one of my prionid drawers. Otherwise, very few beetles came to the lights, or most other insects as well—the sheet being covered primarily by dozens of white-lined Sphinx moths and lots of wasp/bee-type things. We did enjoy the evening, however, as we sat in our chairs and drank a cold brew between checkings of the light.

Mercury-vapor light setup at dusk.
Mercury-vapor light just turned on at sunset.
Mercury-vapor light going strong as darkened settles.
Derobrachus hovorei (palo verde root borer).

Day 4 – Chaparral, New Mexico
We had planned to take NM-213 north to White Sands, but public access was blocked at Ft. Bliss. While backtracking, I spotted Sphaeralcea angustifolia (narrow-leaved globemallow) growing along the roadside and stopped to check it out. There was nothing on any of the plants, despite good growth and appearing to be coming into flowering. I did a little beating on Prosopis glandulosa (mesquite) as well—again very little going on, just a few treehoppers, tiny blister beetles, and one weevil. Larrea tridentata (creosote) was in full bloom, just like at Fabens, and here too there were many Eupompha fissiceps on the flowers and in mating pairs. I couldn’t resist collecting just a few and even made a short video of a mating pair engaged in some interesting behavior. I did see one Gyascutus planicosta as it flew by, but I could not track it to see where it landed. Otherwise all I picked up was another Cysteodemus wislizeni (black bladder-bodied meloid)—along with a photo, and a male cicada (maybe Diceroprocta texana) singing on a dry yucca stalk (hand-collected!).

Cysteodemus wislizeni (black bladder-bodied meloid).
Eupompha fissiceps mating pair engaged in some interesting behavior.

Point of Sands, New Mexico
Jeff wanted to stop here to look for Pleurospasta mirabilis, a really cool-looking blister beetle that looks unlike anything else. He found just one by disturbing the host plant (small purple blooms), and I found none—seems we are right at the tail end of their activity period. I looked around for other things also, but there was not much out. I did catch a couple more cicadas (males singing), and near an Ephedra sp. bush I found a mostly-intact carcass of Sphaerobothris ulkei. The most interesting find, however, was a couple of apparently lost pitfall traps—the barriers had fallen over, and the cops were filled with sand and the carcasses of numerous tenebrionids that had fallen into the traps and never been retrieved. I pulled up the cups and filled in the holes to prevent further loss, finding a few live tenebrionids and trogids and one Pasimachus sp. ground beetle that had not yet succumbed in the process.

Point of Sands, New Mexico.
“Lost” pitfall trap.

Hatch, New Mexico
While passing through the town of Hatch, I couldn’t resist taking the opportunity to stop and photograph some interesting town characters.

An interesting take on the Sinclair dinosaur.
Chili Pepper Man!
Cast of characters.

Deming, New Mexico
Another roadside stop in an area that was green with flowers and also had Yucca (to look for Tragidion) and Ephedra (to look for Sphaerobothris). Neither of those insects were found, and no buprestids or cerambycids were seen on or swept from any of the many composite flowers about including Thelosperma megapotamicum. I did find a couple of the meloid Lytta biguttata, one on flowers of Cirsium sp. and another on an unidentified yellow composite flower, and Jeff found a huge aggregation of another meloid, Epicauta costata, on herbicide-treated Kali tragus (prickly Russian thistle). I finished off the stop by finding a spectacular ridged tenebrionid beetle walking about after the sinking sun went behind some clouds.

Lytta biguttata on yellow composite flower.
Epicauta costata on herbicide-treated Kali tragus (prickly Russian thistle).
Epicauta costata on herbicide-treated Kali tragus (prickly Russian thistle).

Day 5 – Sunshine, New Mexico
We saw a nice stand of what proved to be Picradeniopsis absinthifolia (formerly Bahia absinthifolia, hairyseed bahia) and stopped to check them for buprestids/meloids. None were seen, just a few bees and lots of bee flies (genus Geron?). We did find a fair number of Cysteodemus wislizeni (black bladder-bodied meloid) crawling on the roadsides in a patch of Solanum elaeagnifolium (silverleaf nightshade).

Geron? sp. (family Bombyliidae) on flower of Picradeniopsis absinthifolia (hairyseed bahia).
Geron? sp. (family Bombyliidae) on flower of Picradeniopsis absinthifolia (hairyseed bahia).

Columbus, New Mexico
We found a moist drainage along the roadside with plants blooming in abundance, including Isocoma tenuisecta (burroweed) and Sphaeralcea angustifolia (narrow-leaved globemallow); however, insects were very scarce. I didn’t see any buprestids or cerambycids at all on any of the plants, only picking up one Lytta biguttata and one Cotinis mutabilis on the Isocoma and sweeping single examples of a small black/red Cleridae from the latter and a yellow-flowered composite.

Pepsis thisbe (Thisbe’s tarantula-hawk wasp) on flowers of Isocoma tenuisecta (burroweed).

Animas, New Mexico
We drove a fair distance west hoping to get into a different rainfall system in hope that insects would be present and saw a roadside in good bloom with the surrounding creosote scrub also green and blooming. As soon as we got out of the car we saw big beetles flying overhead and tracked them back to several creosote bushes very near the car with an aggregation of yet another blister beetle, Pyrota postica, which were mating and feeding on the flowers and leaves. After taking a few photos (and collecting my small series), I started sweeping through the variety of plants in bloom along the roadsides. I did not see anything on the flowers themselves (including Baileya multiradiata, pretty good buprestid flower) but collected a series of clytrine chrysomelids and one Dectes sp. While sweeping through Sphaeralcea hastulata (spear globemallow), I got one Agaeocera gentilis, and sweeping though a mix of S. hastulata and S. angustifolia (narrow-leaved globemallow) I got three Agrilus sp. (perhaps the same as I collected south of Van Horn, Texas). Finally, on the latter, I found one more A. gentilis perched on the leaf.

Pyrota postica female on flowering Larrea tridentata (creosote).
Pyrota postica female feeding on the flowers of Larrea tridentata (creosote).
Sphaeralcea hastulata (spear globemallow).

Portal, Arizona
Finally made it to Arizona, and for the first stop I wanted to try a spot below Portal where I’ve had limited success finding Sphaerobothris ulkei on Ephedra. Last time I was I here I found a few, but not until after being distracted by Gyascutus caelatus and Hippomelas sphenicus in the acacias and mesquites. I vowed not to let that happen this time and weaved a zigzag pattern looking at every Ephedra I could find. While I was doing that. I did see one G. caelatus take flight and then caught another that I saw sitting on Vachellia constricta (white-thorn acacia) but remained focused on looking at the Ephedra. Eventually, after not seeing any S. ulkei, I started looking for other buprestids. The acacias were just beginning to flower—only a few plants had open flowers, and Jeff noticed the tiny silhouette of an insect in flight approaching the flowers. He netted it and showed it to me, and to my surprise it was one of the species in the A. stigmata group (black with two red apical spots)—none of which I have ever collected before but which I believe could be A. davidsoni. We spent the next hour watching for and netting the silhouettes on the few trees we could find with flowers, and then I got the beating sheet out and beat more off of the trees (whether in flower or not) to end up with a nice little series. I’ll be anxious to confirm whether these are A. davidsoni.

Mozena arizonensis on Vachellia constricta (white-thorn acacia).
Vachellia constricta (white-thorn acacia).

South of Willcox, Arizona
We got to Willcox around dusk, and the cool, breezy conditions told my gut that it would be pointless to set up the lights. Still, I couldn’t let myself not try, and without much opportunity to look for a good place to set up I just went down Blu Sky Rd to E Moonlight Rd and hoped for the best. My optimism waned rapidly, as conditions continued getting colder and breezier, and not a single insect came to the light—I should’ve listened to my gut!

Moonrise. A twilight zone moment occurred when I realized that I had taken this photo on Moonlight Rd!😮

Day 6 – Jim Wappes Celebration, Hereford, Arizona
Fun day with lots of fellow coleopterists at the home of Steven Lingafelter and Norm Woodley in memory of Jim Wappes. We got there in the early afternoon and enjoyed eats, conversation, and war stories from the field.

Min, Sangmi Lee, Lisa Lee, Candy Kuckartz, the author, Margarethe Brummermann, Jason Botz, & Andrew Johnston (photo by Steven Lingafelter).
The author, Andrew Johnston, Norm Woodley, & Ed Riley (photo by Margarethe Brummermann).

It was a good day for a party, as rain made insect collecting a no-go. As dusk settled, I admired the incredible view from Steve’s and Norm’s back patio!

Low clouds hang over the Huachuca Mountains.

The fun extended well into the evening hours. My thanks to Steve and Norm for hosting the celebration—what fun to see and talk to so many entomologists in one place.

Steve Lingafelter, Gino Nearns, “Kira Brummermann,” & the author (photo by Margarethe Brummermann).
Bill Warner, Norm Woodley, Paul Skelley, Jeff Huether, Steve Lingafelter, Andrew Johnston, & the author (photo by Margarethe Brummermann)

Day 7 – Superstition Mountains, Weaver’s Needle Vista Viewpoint, Arizona
We awoke to rain yesterday morning in Willcox, and it has stayed with us since—first on our way to Hereford for the Wappes Celebration, then up to Phoenix this morning—our efforts to escape the rain by coming north thus proving futile. I came to this spot on a tip that I might find Agrilus cavifrons on Celtis pallida (spiny hackberry) (although maybe a bit early), and I’d hoped despite the light rain I would still be able to find it. I did not—though I found the plants, but I did get a Chrysobothris sp. that I don’t recognize while beating Senegalia greggii (formerly Acacia greggii, cat-claw acacia). Of course, it was on the first plant that I beat, so I ended up beating for another hour with nothing to show for it! While walking the short paved trail, I found Cercidium sp. (palo verde) tree that had been cut up and showed evidence of buprestid infestation in one of the larger branches, so I retrieved and cut it up for rearing.

Water… water… everywhere!
Cylindropuntia fulgida (chain-fruit cholla, jumping cholla, hanging chain cholla).

Lost Dutchman State Park, Arizona
We made a quick stop here to admire its incredible scenery before heading back south.

Superstition Mountains as seen from Lost Dutchman State Park.
Stands of Cylindropuntia fulgida (chain-fruit cholla) frame this view of the Superstition Mountains from Lost Dutchman State Park.
Carnegiea gigantea (saguaro) in front of the Superstition Mountains.
Carnegiea gigantea (saguaro).

Upper Tanque Verde Falls Trailhead, Tucson, Arizona
The rain finally moved out and it was sunny for the drive back to Tucson, but with still-cool temps we weren’t sure if lighting would be worthwhile. We decided to try this spot—not too high (based on a tip by Bill Warner), and I setup the lights using a technique recommended by Roy Morris that involved placing one ultraviolet (UV) light on each side of the sheet, extending the Mercury-vapor lamp above to the top of the sheet, and periodically shutting off the latter to allow the UV lights the pull in the “shyer” insects before turning it back on. While I waited for the lights to started pulling things in, I did some beating around the area. I only collected one specimen, but it was a Cleridae that I don’t recognize, which was beaten off of Prosopis glandulosa (mesquite). Temps ended up in the lower 70s to upper 60s, but at the lights we still got a few longhorned beetles (Aneflomorpha sp.), a few melolonthine scarabs, and a nice series of two species of Pachybrachis (that I will send to Bob Barney).

Jeff looks for beetles at the light sheet.
Lethocerus medius at ultraviolet/mercury-vapor light in mesquite/acacia scrub.

Day 8 – Atascosa Mountains, Ruby Rd near Atascosa Lookout Trailhead, Arizona
The primary quarry here was Acmaeodera chuckbellamyi, a species I described in 2014 from a single specimen collected by my friend and hymenopterist, Mike Arduser, at this location on flowers of Aloysia sp. Several people have tried to find it since—without success, and in June 2011 the area was severely burned by the 27,550-hectare Murphy Fire. I was hoping enough time has passed to allow the area (and beetle population) to recover in this, my first attempt, at finding the species myself. I knew it was a long-shot, and long sorry short I did not find either the beetle or it’s Aloysia host flowers. I did collect a number of other beetles, however, including Lycus lorises, a few longhorned beetles and pachybrachine leaf beetles, and one Acmaeodera quadrivittatoides on flowers of Eysenhardtia orthocarpa (desert kidneywood), several Aneflomorpha sp. and a few pachybrachines on Quercus oblongifolia (Mexican blue oak), and more pachybrachines on Senegalia greggii (catclaw acacia), Mimosa dysocarpa (velvetpod mimosa)—the latter also yielding a few tiny Chrysobothris spp. (one looking like C. lucanus), and Propopis glandulosa (mesquite). Jeff also collected and gave to me a couple of Acmaeodera parkeri on flowers of Talinum aurantiacum (orange flameflower). I hiked 0.7 mi E on Ruby Rd to a spot where I swept a few beetles from low vegetation, and on the underside of a large, fallen branch of Q. oblingifolium I found a large female Polycesta arizonica. Finally, about halfway back to the trailhead I encountered a few tiger beetles on the road near standing puddles of water from the recent rains.

View from Ruby Rd looking south toward Mexico.
View from Ruby Rd looking southwest toward Mexico.
Lycus loripes on flowers of Eysenhardtia orthocarpa (desert kidneywood).
Lycus loripes on flowers of Eysenhardtia orthocarpa (desert kidneywood).
Talinum aurantiacum (orange flameflower).

Atascosa Mountains, Peña Blanca Lake, Arizona
We came here to look for Deltaspis tumacacorii, which like many rare beetles the odds are against finding it despite it having been taken in the area on several occasions. Again, this would not be one of those occasions, but I was happy to find a tiny tiger beetle (Cylindera viridisticta arizonensis) along the creek and even happier to beat a series of two species of Paratyndaris, one large Lampetis webbii (only my fifth specimen). and one Aneflomorpha sp. from mostly dead Senegalia greggii (catclaw acacia).

Leptinotarsa lineata on Ambrosia monogyra (cheeseweed burrobrush)—adult.
Leptinotarsa lineata on Ambrosia monogyra (cheeseweed burrobrush)—larvae.

Huachuca Mountains, lower Carr Canyon, Arizona
After dropping Jeff off at his hotel in Tucson (he flies home in the morning), I high-tailed it down to Carr Canyon to do some light collecting. It has been a long time since I’ve seen my sheet covered so quickly and thoroughly with insects! My quarry was longhorned beetles—of which I got a nice variety, but who can resist also the variety of scarabs, ground beetles, tenebrionids, blister beetles, and even planthoppers that flock to the lights in the mountains of southeastern Arizona? I’ll have to control myself a little better in the coming nights! Walking about along the roadsides (hoping to see Amblycheila giant tiger beetles), I encountered a gorgeous male Aphonopelma chalcodes (Arizona blonde tarantula).

Mercury-vapor/ultraviolet light setup.
Enaphalodes cortiphagus at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Parabyrsopolis chihuahuae at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Phileurus truncatus (triceratops beetle) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Chrysina beyeri (Beyer’s jewel scarab) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Chrysina gloriosa (glorious jewel scarab) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Pachysphinx occidentalis (western poplar sphinx) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Manduca florestan (Florestan sphinx) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Aphonopelma chalcodes (Arizona blonde tarantula) on ground in in oak/juniper woodland.
Aphonopelma chalcodes (Arizona blonde tarantula) on ground in in oak/juniper woodland.
Aphonopelma chalcodes (Arizona blonde tarantula) on ground in in oak/juniper woodland.

Day 9 – Huachuca Mountains, Copper Canyon, Arizona
I met Steve Lingafelter and Norm Woodley at their house, and together we drove to the famed Copper Canyon on the south side of the Huachucas. On the way I got a nice primer about the species that have been collected there and the plants they have been collected on. Steve and I started out walking the trail up the canyon while Norm swept the area down below. I pretty much beat every oak along the way, for a while only getting a smattering of beetles—Agrilaxia sp. on Quercus emoryi (Emory oak) and also on Q. arizonica (Arizona white oak) along with Sternidius decorus?. About a half-mile up the trail I beat dead branches of Q. hypoleucoides (silverleaf oak) and, not seeing anything at first, said “I’m going back; I’m literally not getting anything.” Right then, a small black beetle on the sheet caught my eye. I looked at it closely and realized it was a Mastogenius (prob. M. robustus)! I popped it in the bottle and beat more dead branches from the same tree and got not only another Mastogenius but also Tigrinestola tigrina. Freshly motivated, I spent the next half-hour working all the oaks in the area—and, as often happens, did not see another beetle! Nevertheless, it was hard not to be happy with the beetles that I’d gotten.

An unidentified treehopper nymph, likely something in the subfamily Smiliinae, tribe Telamonini, on twig of Quercus arizonicus (Arizona white oak).

I walked back down the trail and met Norm, who was just starting up. He filled me in on the results from below, which included sweeping a few Agrilaxia hespenheidei—one of my target species—and an Agrilus sp. on Bouvardia ternifolia (firecrackerbush). I decided to work the slope under the road, reasoning that Norm had likely already worked the flat ground below. On the first B. ternifolia I approached, I saw an A. hespenheidei on the flower, gave the plant a sweep, and caught not only the A. hespenheidei but also an Agrilus sp. (maybe A. latifrons). Over the next hour I would sweep a nice series of A. hespenheidei from B. ternifolia (but not another Agrilus sp.). While I was doing that, I also swept the numerous stands of Acaciella angustissima (formerly Acacia angustissima, prairie acacia) looking for the large, spectacular Agrilus cavatus. I would find two, and considering that I swept perhaps 50 or more stands they were well earned. Also, in the meantime, I noticed Acmaeodera parkeri on small blue flowers that I eventually identified as Evolvulus arizonicus (Arizona blue-eyes). The flowers were few in number and the beetles difficult to catch, so I only ended up with two specimens. Further down the slope the flowers much more abundant, but there was not a beetle to be found on them. In the waning moments of my visit, I encountered two Trichodes peninsularis horni on flowers of Lasianthaea podocephala (San Pedro daisy).

Bouvardia ternifolia (firecrackerbush), flower host for Agrilaxia hespenheidei.
Acaciella angustissima (prairie acacia), host for Agrilus cavatus.
Evolvulus arizonicus (Arizona blue-eyes), flower host for Acmaeodera parkeri.
Lasianthaea podocephala, flower host for Trichodes peninsularis horni.

Eventually we all met up at the car, compared our catches (not surprisingly, Norm did very well with buprestids and Steve did very well with cerambycids), and I did okay in both counts. We headed back over Montezuma Pass and were greeted with stunning views looking down into the Coronado National Monument!

Coronado National Monument from near Montezuma Pass.

Huachuca Mountains, Miller Canyon, Arizona
After dropping Steve off, Norm and I went to nearby Miller Canyon to look at a spot where he has collected three species of Taphrocerus (I’ve only collected two, but only once way back in 1987 at a spot in the lower canyon). We thought it might be too late in the season, but it was at least worth a shot. The sedges were lush and green, but the “sedgy wedgies” were absent. Looks like I’ll have to just come back out in April or May!

After striking out with Taphrocerus, I went back down to the lower elevations of the canyon to look for Tragidion on the stand of Baccharis sarothroides (desert broom) that occurs there. Once again, I was likely too late to find them, but as with Taphrocerus it was certainly worth a shot, and again I would not find any despite looking and most of the large plants in the area. I did find a few Euphoria leucographa feeding on the sap flows and a very large red/black clytrine, so it wasn’t all for naught. Just another reason to come out earlier in the season.

Euphoria leucographa feeding on a sap flow on stem of Baccharis sarothroides (desert broom).

Day 10 – Santa Rita Mountains, upper Box Canyon, Arizona
After getting reports of buprestid activity near Madera Canyon, I decided to head to the Santa Rita Mountains today instead of continuing in the Huachucas. The shortest route to this most famous of canyons in the Santa Ritas goes down Box Canyon, a less-well-known but still-fantastic canyon in its own right and where I’ve had good luck collecting the two previous times I’ve been there (August 2018 w/ Art Evans, Steve Lingafelter, and Norm Woodley; and September 2019 w/ Jeff Huether). I stopped at the “dry falls” and worked my way back up the road to a point where I’ve collected the majority of my insects there. Along the way, I beat the flowering Eysenhardtia orthocarpa (desert kidneywood)—insects were not numerous on the plants, but over the course of the trips up and back I got Acmaeodera gibbula, A. cazieri, two Aneflomorpha sp., and a few Lycus sp. I also swept the just-beginning-to-flower Mimosa dysocarpa (velvetpod mimosa) but got just a single Sphaenothecus bivittatus. When I reached the top of the canyon, I looked for a small patch of Allionia incarnata (creeping four-o’clock) in the steep road bank, off the flowers of which I have previously collected Acmaeodera cazieri, A. parkeri, and A. yuccavora. I found the patch, but the plants were not yet in flower. What was in flower on the flats above the bank, however, was Talinum aurantiacum (orange flameflower), off which I collected all three species (A. cazieri being the most abundant and only a single A. cazieri). As at previous stops this week, it seems that insect activity is beginning but is still a bit shy of coming into full peak.

Larva of Euscirrhopterus gloveri (purslane moth), which was present in outbreak numbers on a portulacaceous plant growing amongst Talinum aurantiacum.

While I was collecting, a caravan of cars came by. They turned out to be filled with entomologists attending the Invertebrates in Education Conference, one of whom I knew—Tad Yankoski of the Sophia M. Sachs Butterfly House in my hometown of St. Louis. He handed me a vial containing a large, live individual of Polycesta aruensis. I was excited to see this, especially when he told me he found it and saw many more in the flats below Madera Canyon, where I had planned to go next!

Santa Rita Mountains, flats below Madera Canyon, Arizona
It was a frustrating afternoon on several fronts. Starting off, I had trouble finding the Polycesta aruensis locality, and when I finally did find it there was nary a Polycesta to be seen. Perhaps they sleep during the heat of the day.🤷 After that, there was little time to go anywhere but Madera Canyon, where I spent a half-hour beating Quercus oblongifolia (Mexican blue oak) hoping to see Chrysobothris chalcophoroides (I didn’t) and another two hours checking out Baccharis sarothroides (desert broom) in the area where I collected Stenaspis verticalis and Tragidion deceptus two years ago (also fruitless). Of course, the dreadfully common Stenaspis solitaria was everywhere, but all I ended up collecting was a tenebrionid on the 2-track, a clytrine beaten from Cercidium aculeata (retama), and one Euphoria leucographa along with a nondescript cerambycid (Heaperophanini maybe) on B. sarothroides. On the way back to the vehicle, I encountered a dead, mostly skeletonized deer, and while I rarely collect from carcasses, I noticed a little green beetle crawling on the jaw bone. I figured it must be the cosmopolitan clerid, Necrobia rufipes—something I’d not seen before, so I collected it and tried to collect but missed another one.

Rain clouds gather over Madera Canyon.
Another view of the mouth of Madera Canyon.
Magusa sp. (one of the narrow-wings) caterpillar feeding on Sarcomphalus obtusifolius (lotebush).

Santa Rita Mountains, lower Florida Canyon, Arizona
I suppose I can credit my frustrating afternoon for one of my best nights of lighting ever. I stumbled upon this spot at the bottom of Florida Canyon during this afternoon’s Polycesta wild-goose chase and immediately thought, “Wow, what a perfect spot to set up a light!” A nice place to pull off the road with a small, level clearing embedded within low-elevation oak woodland. There was even a babbling creek in the background! It was close to dark by the time I returned and set up the lights (would’ve been even later if I’d gone into town for a “real” dinner). Ironically, there were neither the diversity nor quantity of beetles as two nights ago in lower Carr Canyon. But the cerambycids brought it… and kept bringing it! It seemed like every time I got up to check the sheet there were another 4–5 individuals. I ended up leaving the sheet up for four full hours and collected perhaps 40–50 specimens representing a dozen or more species. A few I don’t recognize, and most of those that I do recognize have resided in my cabinet in precious few numbers until now. It’s been years literally since I’ve had a night like this, and it’s a nice shot of motivation leading into the last few days of what is starting to feel like a long trip.

Sometimes circumstances dictate an unusual dinner.
Enaphalodes niveitectus at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Tigrinestola tigrinus at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Coenopoeus palmeri at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Syssphinx hubbardi (Hubbard’s silk moth) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Eacles oslari (Oslar’s imperial moth) at Mercury-vapor/ultraviolet light in oak/juniper woodland.

Day 11 – Madera Canyon Rd, Continental, Arizona
I came back to the spot where Tad Yankoski had seen Polycesta aruensis so abundantly yesterday morning but which was completely absent by the time I got there in the afternoon. I did not see any adults on the trees this morning either and was about to give up when I spotted a few partially dead trees with very large, apparently fresh emergence holes in the main trunks that were the perfect size for P. aruensis—good thing I brought my chainsaw! I cut a trunk with its branches and segregated the cut up wood into age (fresh dead versus older) and size (twigs, medium branches, and main trunk) classes. While I was doing this, a Polycesta adult dropped off one of the fresh-dead, medium-sized branches! I beat the remaining branches on the tree and on nearby trees but did not see any more, so whatever Tad witnessed yesterday morning must have been an ephemeral event, perhaps related to synchronized emergence from the very trees among which I collected the wood. NOTE: don’t let anyone tell you that cutting up wood for rearing beetles is anything but a sweaty, exhausting endeavor, even with temps still in the mid-80s and decent cloud cover!

This camouflaged tanker marks the spot—trees on the W side of the tanker.
Job half-done—wood cut up.
Job complete—wood segregated and bundled.

Santa Rita Mountains, lower Florida Canyon, Arizona
I had such good night collecting here last night that I thought I’d come back and see what I could find during the day in this ribbon of riparian oak/hackberry woodland. Almost immediately I beat a Paratyndaris sp. off of dead Celtis reticulata (netleaf hackberry). The tree was very dead, but I knew Paratyndaris spp. like old, dead wood, so I split open some of the branches and found larvae inside and also a dead and unemerged but perfectly intact adult of a very tiny Chrysobothris sp. inside one of the smaller branches. Beating on other plants in the area was, in general, fruitless, but occasionally (and just often enough) I encountered something of interest that motivated me to continue working: Paratyndaris sp. and Agrilus sp. on Quercus oblongifolia (Arizona white oak), a small red/brown elaterid on Prosopis glandulosa (mesquite), and Acmaeoderopsis sp., Paratyndaris sp., and a few clytrines on Senegalia greggii (cat-claw acacia).

Lower Florida Canyon.
Celtis reticulata (netleaf hackberry).

Santa Rita Mountains, Gardner Canyon, Arizona
Last night, I got a message from fellow buprestophile Robert Velten, who told me that he would be in Gardner Canyon the next day, so after finishing at Florida Canyon I drove over to Gardner Canyon to meet up with him. Despite being longtime correspondents, Rob and I had never actually met face-to-face, so I was thrilled to have the chance to do so and spend some time with him in the field. Joining him were his mothing buddies Steve McElfresh and Paul Tuskes, and a little later our mutual friend and Arizona coleopterist-extraordinaire, Margarethe Brummermann, also joined us for a night of lighting. It was great to spend time at the lights with so many like-minded folks! There were three light stations between us, but the weather was less than cooperative—a persistent cool breeze accompanied constant lighting and thunder in the mountains above. Eventually, the threat was realized when the skies opened up, prompting a hasty dismantling and storage of all my lighting equipment safely inside the vehicle. Nevertheless, in the time that I was able to collect, I got a small number of longhorned beetles (half of which came to my light in the moments I was taking it down—longhorns typically become very active right before a storm) along with a variety of showy scarabs and clerids. The rain ended as quickly as it began, so the socializing continued. The entire evening I was continuously taunted, however, by a large prionid sitting inside its emergence hole on the trunk of a large Quercus emoryi (Emory oak). It only showed its jaws and antennae, and if I even touched the tree to boost myself up for a closer look it withdrew deep into the hole. I’m convinced it was Nothopleura madericus—a species I’ve never collected. I can still hear it laughing at me! My attempt to find one out and about by scanning the trunks and branches of of the other oaks in the area with my headlight was not successful, although I did collect another elaphidiine longhorn in such manner.

Polyphylla decemlineata (or perhaps a new species) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Lucanus mazama (cottonwood stag beetle) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Xyloryctes thestalus (western rhinoceros beetle) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Hemiphileurus illatus at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Cymatodera horni at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Manduca rustica (rustic sphinx) at Mercury-vapor/ultraviolet light in oak/juniper woodland.

Day 12 – Santa Rita Mountains, Gardner Canyon, Arizona
Rob had noticed a stand of Anisacanthus thurberi (desert honeysuckle)—a host for Spectralia cuprescens—along the road into Gardner Canyon, so together (after morning coffee!) we checked the spot on the way out. The plants were in the early stages of leafing out, and after visually inspecting them for a while and not seeing anything I decided to get out the beating sheet to sample the stand more thoroughly. My eyes did not deceive me—neither one of us found any. Too early? Low population? Who knows! I did beat one clytrine off the plants and collected a few weevils by beating Prosopis glandulosa (mesquite) before finding Acmaeodera parkeri and A. cazieri on the flowers of Allionia incarnata (creeping four o’clock). The flowers were common in the area around the road, but no Acmaeodera were seen until I started scaling the steep hillside nearby—a similar situation in which I’ve found these species on this flower in Box Canyon.

Rob Velten and the author enjoying morning coffee (photo by Margarethe Brummermann).
Dense stand of Anisacanthus thurberi (desert honeysuckle) in Gardner Canyon.
Gardner Creek running full after last night’s rain.
Yours truly with buprestophile extraordinaire Rob Velten.

Huachuca Mountains, upper Carr Canyon, Reef Township Campground, Arizona
I’ve been wanting to explore the higher reaches of Carr Canyon ever since I arrived in Arizona a full week ago. It is the only high canyon in the Huachucas that has a road leading all the way into its upper reaches. At these high elevations the forest is Pinus ponderosa (ponderosa pine) and Quercus hypoleucoides (silverleaf oak). There are many species of woodboring beetles at these high elevations that simply aren’t found down below. Another reason is the recent discovery up here of one of the rarest and most enigmatic of North American longhorned beetles, Placoschema dimorpha. Not even know to science until it was described from Mexico in 2007, it has since popped up here and a few other places in southeast Arizona—some of which have been heavily studied by coleopterists or many decades. Now, I didn’t actually <u>expect</u> to find such a rare thing, but maybe I could get lucky or at least find some other unusual species. The specimen at this location was photographed on a burned pine tree; however, I do not think that is the host (as far as I am aware, no species in the tribe utilizes gymnosperms as larval hosts). I think the host must be oak, as is the case for many trachderines. So, while I kept an eye out for burned pine trees, I also looked for oak, and especially recently dead oaks showing signs of woodboring beetle infestation. I did some of beating on Q. hypoleucoides and had collected just a clerid (Enoclerus bimaculatus) when I came upon a recently fallen Q. hypoleucoides that showed a few buprestid(ish) emergence holes and looked to be “the right age.” Cutting away the bark of the trunk revealed galleries, and chopping into sapwood revealed buprestid larvae in their galleries. I tagged it for retrieval, eventually cutting it up and segregating the trunk sections from the branches. Very nearby, I found another dead Q. hypoleucoides, this one much smaller and apparently cut rather than fallen. Unlike the previous one, however, this one showed the round holes with ejecting frass that indicated infestation by cerambycid rather than buprestid larvae. Cutting into the wood confirmed the presence of such, and so this one also was later cut up and bundled for bringing back. I saw no beetles on the trunks of any of the many fire-scarred pines lofting overhead, but at one point I spotted in the distance the telltale brown flagging of a recently died pine up the slope. Hiking up to it took some effort, but when I reached it the first thing I saw was a giant click beetle—Chalcolepidius apacheanus (Apache click beetle)—nestled against the ground at the base of the trunk (apparently ovipositing?). Inspecting the trunk of the tree itself, I noted just a few buprestid emergence holes that seemed fresh. Once again, chopping away the bark revealed the frass-packed galleries, and chopping into the heartwood revealed a large pre-pupal buprestid larvae. This was put into a vial, and I noted the location so I could return with the chainsaw and “bring ‘er down.” As I was cutting up the oaks, I found some small, recently cut pine branches near where I had parked the car. I found (and accidentally killed) a woodboring beetle larva of some type (I mangled it pretty good, but I think it was a buprestid) in one of them, so that was good enough to earn a spot in the rearing tubs. Unfortunately, I was not able to retrieve the dead pine tree—cutting up the oaks took a fair bit of time, during which darkening clouds gathered over the nearby peaks. Eventually cracks of thunder began piercing the air. It was all I could do to get the oaks into the car and all of the equipment put away before heavy rain drops began pelting the car. I had no idea what the storm would bring, but the last thing I wanted to be was stranded on the top of a mountain on my last full day of field collecting. As it turned out, the storm was more bark than bite (although the sharp drop in temperatures would kill lighting later in the evening). Perhaps I’ll be able to get back up the mountain in the morning and retrieve the pine tree.

Chalcolepidius apacheanus (Apache click beetle) on trunk base of dead Pinus ponderosa (ponderosa pine).

Huachuca Mountains, lower Carr Canyon, Arizona
It seems like forever ago that I began this trip, and now the last night of collecting has arrived. I decided to come back to this spot where I’d had such good luck earlier this week, but I wasn’t optimistic given how rain chased me out of the canyon earlier. My pessimism was warranted, and while I did picked up a variety of things, it included only two longhorns—both rather pedestrian species. No, the real charm of the night came not from collecting insects, but after the lights were down and my mind was free to wander as I leaned back in my chair and gazed into the crisp, dark, starry Arizona sky—its perimeter along the horizon bound by a craggy silhouette of nearby oak trees and distant peaks; from listening to the sounds of the night, alternately focusing on the individual cricket or distant coyote versus the chorus as a whole. Only to the north could I see the faint glow of city lights—the only sign that anything beyond me and this moment exist. These moments happen only once on a trip (maybe twice), and they are to be savored; indelibly stamped into the memory banks for future enjoyment; one of those experiences that, when recalled, is guaranteed to trigger euphoric recall.

Campground set for a final night of lighting.
Manduca sexta (Carolina sphinx) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Sphinx dollii (Doll’s sphinx) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Sphinx dollii (Doll’s sphinx) at Mercury-vapor/ultraviolet light in oak/juniper woodland.

Day 13 – Huachuca Mountains, upper Carr Canyon, Reef Township Campground, Arizona
I went back up to the top of Carr Canyon to retrieve the dead Pinus ponderosa that I found yesterday. Good thing I did, as I also found my favorite hatchet (which I’d inadvertently left behind yesterday). The 9” diameter trunk was almost too big a job for my Stihl MiniBoss chainsaw, but I kept at it and finally felled the the 25’ tall tree. It took three trips up and down the steep slope—each round trip almost a half-mile—to haul out the upper 10 ft of trunk and associated branches, which I segregated into three batches for rearing: trunk, 1–2” día. branches, and <1” dia. twigs. Now let’s hope the effort was worth it and I get some good species out of the wood.

Pinus ponderosa (ponderosa pine) dominates the high elevations in the Huachuca Mountains.

Huachuca Mountains, near Carr Canyon, Waterfall, Arizona
One last stop to take in the terrifyingly magnificent views from atop Carr Canyon! You can see the road that I traveled up this morning snaking back down the right side of the mountain. The massif to the left is the highest point that you can see from the valley below, but there are much higher peaks behind me.

Terrifyingly steep, magnificently endless drop!
Carr Canyon Rd snakes up the right side of the mountain.
This massif is the highest point visible from the mouth of the canyon.

Epilogue
Sadly, I could squeeze no more stops into the trip—I’d allowed myself two days to make the 24-hour drive back to St. Louis, and it was already almost noon on the first of the two days. I left, however, with a rack full of vials filled with insects and a renewed love for Arizona and the desert southwest that first captured my heart some 37 years ago!

©️ Ted C. MacRae 2021

2021 West Texas Insect Collecting Trip iReport

Alternative title: Rich and Ted’s “Excellenter” Adventure.

This is the ninth “Collecting Trip iReport”; this one covering a 10-day trip to western Texas from April 27 to May 6, 2021 with friend and local collecting buddy Rich Thoma. Rich and I have done many shorter collecting trips (up to five days) throughout Missouri and in the neighboring states of Kansas, Nebraska, and Oklahoma (in fact, our first joint trip was to Barber County, Kansas way back in May 1986!). This trip, however, was our first truly long one together—10 days of collecting plus a travel day on each end. To take full advantage of the amount of time we had, we chose western Texas; an area that I have visited several times from the mid-90s through 2004 but not since. We wanted to make the trip during early to mid-May, but scheduling conflicts forced us to go earlier. I reasoned that even if it was a bit too early in the season, I could still collect infested wood for rearing—as I did with great success during my April 2004 trip. For Rich, who is more of a general insect collector, the trip provided him an opportunity for extended collecting in an area that he’d not previously spent a lot of time.

As with all previous “iReports” in this series, this report is illustrated exclusively with iPhone photographs (thus the term “iReport”), with previous versions including the following:
2013 Oklahoma
2013 Great Basin
2014 Great Plains
2015 Texas
2018 New Mexico/Texas
2018 Arizona
2019 Arkansas/Oklahoma
2019 Arizona/California


Day 1 – Monahans State Park, Shin Oak Picnic Area
First stop of the trip. I was hoping to see beetles on flowers and maybe some tiger beetles, but unfortunately the area hasn’t had any rain yet this spring (according to the ranger). The mesquite was in bloom, but the only beetles I beat from it were a few tiny weevils. A few other plants were in bloom, but only one—Hymenopappus flavescens—had beetles on it (mordellids, which I picked up for Enrico Ruzzier). After a lot of walking I noticed Quercus havardii (shin oak) with flagged branches of dead leaves—a bit of investigation revealed it had been attacked by what must be Chrysobothris mescalero, so I collected as many flagged branches as I could find (7 total) and will bring them back for rearing.

Monahans Sandhills State Park.
Oenethera berlandieri (Berlandier’s sundrops).
Penstemon buckleyi (Buckley’s beardtongue).
Hymenopappus flavescens (collegeflower)—host flower for mordellids).
Chaetopappa ericoides (rose-heath).
Quercus havardii (shin oak) attacked by buprestid, presumably Chrysobothris mescalero.

Monahans State Park, Sandhills Picnic Area
The big dunes are in this area. We didn’t expect to see any insects but brought our nets anyway. As we were walking the ridge we saw two grouse-like birds in the distance. We tracked them for a bit before I decided to go back and get my binoculars. They kept us at bay, but eventually I was able to get close enough to get a good look at them—they turned out to be scaled quail, a new bird for me. We continued tracking them and eventually they were joined by two more individuals. Handsome birds!

Rich scans the vast sand dunes.
Endless dunes!

Monahans State Park, Shin Oak Picnic Area
After going into town and picking up some dinner, we came back out to the park and setup the ultraviolet lights. I didn’t have much optimism based on the lack of insect activity we saw during the day, but the temperatures were still plenty warm (well into the 80s) and we had nothing better to do. We returned to the Shin Oak Picnic Area since it had a mix of open and more vegetated dunes. Glad we did because two male Prionus arenarius, one Megacyllene antennata, and a tiny, unidentified elaphidiine came to the lights. I also found two small darkling beetles crawling on the sand nearby. I searched the surrounding sand hoping to find more males looking for females, or perhaps even a female herself, but found none. Wolf spiders, however, were common on the sand, their glowing eyes drawing attention beyond their abundance. I guess they are a species of Hogna, but I’m not certain—I photographed two individuals. Also got a large bostrichid (Apatides fortis?) at the light. Before we took down the lights, Rich called me over to see a tiny, slender, worm-like snake that we eventually determined was one of the blind snakes (Leptotyphlum sp.)—definitely a first for me.

Blacklights setup and humming.
Prionus arenarius male in front of the blacklight. This species is restricted to sand dune systems in west Texas and southeastern New Mexico.
Megacyllene antennata at ultraviolet light.
Hogna sp. (burrow-living wolf spider)—individual #1. This one appears to be a large female.
Hogna sp. (burrow-living wolf spider)—individual #2.
Hogna sp. (burrow-living wolf spider)—individual #2.

Day 2 – Toyahyale
We stopped here on a tip from Jason Hansen and Tyler Hedlund, who swept good numbers of Agrilus cochisei off of Ambrosia psilostachya (western ragweed)—albeit, a few weeks later during May. I found the plants, but they were very small and low to the ground. Nevertheless, adults could be swept abundantly from the plants, and I was able to take good photos of singles and a mating pair with the big camera (iPhone photo here just to show what they loook like). Also got a single specimen of an apparently undescribed Acmaeodera sp. while sweeping for A. cochisei and one of two A. cochisei adults that I saw on flowers of Sphaeralcea sp.

Ambrosia psilostachya (western ragweed).
Agrilus cochesei mating pair on Ambrosia psilostachya (western ragweed).

Davis Mountains, 15.8 mi NE Ft. Davis
We stopped here to look for the undescribed species of Acmaeodera, which Jason had found in good numbers during May on blooms of Lygodesmia and Convolvulus. Both plants were present, but neither was in bloom. Still, I found one adult on flowers of Verbina sp. and swept another from roadside vegetation. Ambrosia polystachia (western ragweed) was also present—I looked visually for Agrilus cochisei and did not see any, but I did get one adult and a couple of cryptocephalines in the sweeping that produced the second Acmaeodera.

Verbina sp. (host for Acmaeodera sp.).
Verbina sp. (host for Acmaeodera sp.).

Day 3 – Point of Rocks Roadside Park
The weather turned decidedly cool in the Davis Mountains—first time I’ve ever frozen camping out on a collecting trip. The high temps are expected to stay in the 50s to 60s with a chance of rain for the next few days, so we decided to head down to the Big Bend area where there is still a chance of rain but warmer temps (up to the high 70s). Maybe we’ll come back to this area next week. Before leaving, however, I wanted to check the Quercus vasseyana (vassey oak) at Point of Rocks, where in the past I collected a good series of Mastogenius texanus even earlier in April (it was actually undescribed at the time). I’ve also collected Elytroleptus lycid-mimicking cerambycids on soapberry flowers here in June, although I knew the soapberry would not be in bloom. There was nothing on the oaks, but I did collect a few miscellaneous beetles beating Prosopis glandulosa (mesquite) and found one vassey oak branch with evidence of wood-boring beetle larval feeding and which I collected for rearing.

Point of Rocks Roadside Park.

17 mi S Alpine
This is the first of two picnic areas along Hwy 118 going south from Alpine towards Big Bend National Park. There are lots of big Quercus vasseyana and Q. grisea here, so I stopped to see if I could find any infested wood. Bingo—one of the Q. grisea had a dead branch hanging from it that looked recently dead, and breaking apart a few of its smaller branches revealed fresh larval workings of some kind of buprestid (perhaps Polycesta arizonica). I cut of the branch and will bring back the bundle for rearing.

Dead Quercus grisea (gray oak) cut and bundled for rearing.

26 mi S Alpine
This is the second of two picnic areas south of Alpine on Hwy 118 towards Big Bend National Park. Rich and I have both stopped here before, and Rich brought back infested wood (apparently Juglans sp.) from which I reared Chrysobothris comanche, so that was the plan again unless we saw active insects. We did not, so I scanned the trees and found a small Celtis laevigata (sugarberry) that had recently died—the bark was peeling, but there were no emergence holes that I could see. I started chopping into the trunk wood and quickly encountered a large buprestid larvae in its “pre-pupal fold”. This could be Texania fulleri based on host and location, so I cut a couple of bolts from the trunk to bring back for rearing. The branches also showed fresh larval workings, so I cut up one along with its smaller branchlets to also bring back for rearing.

Texania sp. prob. fulleri larva in trunk sapwood of dead Celtis laevigata (sugarberry).

3.3 mi W of Hwy 118 on Agua Fria Rd.
Last stop of the day, which I was told could have water with tiger beetles. The creekbed was bone dry, and I collected but a single Chrysobothris sp. beating Prosopis glandulosa in flower—amazing given the proliferation of wildflowers that were in bloom. We did find a nautiloid/ammonite-type fossil in the bone-dry creekbed, which Rich says is of Cretaceous origin based on clam fossils in the underlying layer, and I tracked a common poorwill (Phalaenoptilus nuttallii) for a little bit, eventually getting close enough for the rare iPhone bird photo. Interesting position it assumed upon landing with its wings outstretched above its back.

Amazing wildflower displays in the area.
Seam in sedimentary layer of bedrock. I’m not sure if it is of volcanic provenance.
Nautiloid/ammonite-type fossil—large (about 8” diameter).
Common poorwill (Phalaenoptilus nuttallii) feigning injury.

Big Bend Ranch State Park, West Contrabondo Campground
We arrived in Study Butte with just enough time to check into a motel and get dinner before heading out to Big Bend Ranch State Park. The drive through the park was incredible as we searched for a spot to setup the lights. After finding such spot, however, we were greeted as we got out of the car by a stiff, chilly wind. I knew there was no point in going through the trouble to setup, so instead we drove further down the 2-track to an amazing scenic overlook into an impressive box canyon. Words cannot describe the contortions this acrophobiac took to find good position for these photos, but it was well worth the views.

Dusk along Hwy 170 approaching Big Bend Ranch State Park.
Sunset over canyon near West Contrabondo Campground.
The closest I will ever get to the edge of a canyon!

Day 4 – Big Bend National Park, Boquillas Canyon Trail
Well, the rain and cold continue to follow us. Rather than trying once again to drive somewhere else to escape, we decided to just sit this day out and visit the national park (not a bad Plan B!). Boquillas Canyon is an amazing slice through the rocks along the course of the Rio Grande River, and we hiked as far into the canyon as we could before sheet rock on the left and deep water on the right prevented any further progress. We saw only two insects—a tiger beetle larva that I “fished” out of one of the many larval burrows we saw (definitely Tetracha, and likely T. carolina) and a velvet ant (black head and pronotum, red abdomen).

Rio Grande River from Boquillas Canyon Trail.
Tetracha sp. prob. carolina (Carolina metallic tiger beetle) larva extracted from its borrow.
Mouth of Boquillas Canyon.
Mouth of Boquillas Canyon.
Cobblestone view of Boquillas Canyon.
Rio Grande River in Boquillas Canyon.
Rich contemplates emigration.
Still contemplating.

Big Bend National Park, near Panther Junction
Driving towards the Chisos Mountains after hiking the Boquillas Canyon Trail, we encountered this fine adult male Aphonopelmis hentzii (Texas tarantula) crossing the road.

Male Aphonopelmis hentzii (Texas tarantula).

Big Bend National Park, Chisos Basin, Window Trail
After lunching at Panther Junction, we headed up the into the Chisos Mountains towards Chisos Basin. Heavy clouds shrouded the peaks, so we weren’t sure what we would encounter up there, and once in the cloud zone and then heading down into the basin we could hardly see anything. Suddenly the western side of the basin came into view, still overcast and drizzly but at least free from the heavy fog that shrouded the eastern half of the basin. That made our decision of which trail to hike easy—the Lost Mines Trail under heavy fog versus the Window Trail with semi-clear views. I’ve hiked the Window Trail several times, but the last time was 17 years ago, and Rich in his single attempt a year or two later did not make it to the “Window” due to an impatient 10-year old son in tow. The views on the way down the canyon were spectacular—not despite the rain and clouds but because of it. It is a rare opportunity to see richly moist desert mountains shrouded in mist. At one point on the way down, a Woodhouse’s scrub jay (Aphelocoma woodhouseii) caught our attention—sitting very nearby in a tree before hopping down to the ground and nonchalantly pecking for bugs. Another soon joined him, first landing on a branch just a few feet above me and returning my captivated stare for a few moments before joining his mate on the trail ahead of us… followed shortly by a third individual. Their soft chirpings were a charming contrast to their more familiar raucous calls, and Rich and I soaked in the moment until they moved on. The trail is not an easy hike—nearly 7 miles round trip, dropping over a thousand feet on the way down, and then gaining over a thousand feet on the way back. The “Window,” however, is a sight to behold—a narrow gap in the rocks soaring high overhead with a view out onto the desert floor almost a thousand feet below. There is tempting danger at the window—its smooth, water-carved rocks are deceptively slippery even in dry conditions, and with the rain of the day they were especially so. I would not be surprised to learn that at least one person had made a fatal error in judging how close to the window one can get. They would have had plenty of time to think about that mistake on the way down! The views on the way back up were even more breathtaking, as fog enshrouded the high peaks towering above us. Periodically the sun attempted to push through the clouds, creating surreal lighting in a battle of sun versus rain, but eventually the rain won out and fell steadily on us for the last, switchback-laden mile back to the trailhead. As for insects, we actually did see some despite the rain—a few blister beetles resting torpidly on yellow composite flowers.

Window Trailhead.
Chisos Mountains’ South Rim from Window Trail.
Chisos Mountains east rim from Window Trail.
Beginning the descent to the “Window.”
Rich photographs a Woodhouse’s scrub-jay.
Yellow composites bloom en masse.
Resting point halfway down—Rich’s prior turnaround point.
The descent steepens!
Steps carved into the rock aid the traverse across slippery rocks.
The “Window” from as close as I was willing.
The author (left) and Rich document their arrival at the “Window.”
Looking back at the “Window” from a bit further back up the trail.
Beginning the rugged, 1,000-ft ascent back up to the basin.
Clouds and mist shroud the surrounding peaks.
A rainy last few miles provides a spectacular last look at from whence we came.

Day 5 – Big Bend National Park, Sotol Overlook
We’re on our way to Santa Elena Canyon and stopped at this overlook. From a distance of 14 air miles, the canyon entrance looks like a tiny split in the rocks, belying the 1000-foot canyon walls that await us. Cacti were nicely in bloom, if a bit rain battered—two species of yellow-flowered Opuntia (pricklypear) and the always extraordinary pink flowers of Cylindropuntia imbricata (tree cholla). No insects were to be found, but we did find a live Orthoporus ornatus (desert millipede)—the first that we’ve seen on this trip—who obliged us by coiling into its classic defensive pose.

View towards Santa Elena Canyon—some 15 miles to the south—from Sotol Vista Overlook.
Orthoporus ornatus (desert millipede).
Cylindropuntia imbricata (tree cholla).

Big Bend National Park, Santa Elena Canyon
From 14 air miles away, Santa Elena Canyon looks like a tiny split in a little cliff (see previous post). Up close, however, it’s soaring walls tower 1000 feet overhead! The hike into the canyon features a tortuous staircase to bypass a narrows, followed by a leisurely stroll along the canyon bottoms along the Rio Grande River. Rain last night has triggered an en masse millipede emergence, and even a few insects were seen: velvet ant; Acmaeodera mixta, Trichodes sp. and Gnathium sp. on yellow composite flowers; and Omorgus sp. crawling in the sand.

Undetermined yellow composite in Rio Grande River floodplain.
Euodynerus pratensis on flower of undetermined yellow composite.
Mouth of Santa Elena Canyon.
The Rio Grande River spills forth from Santa Elena Canyon.
Agave lechuguilla (lechuguilla).
View of Mexican side of Santa Elena Canyon from the U.S. side.
The Santa Elena Canyon Trail probes deeper into the canyon.
Narrowing canyon walls.
No more land!

Big Bend National Park, Cerro Castellan (Castolon Peak)
The layers visible in Cerro Castellan reveal millions of years of volcanic events. Stacked in this tower are several lava flows and volcanic tuffs (ash deposits), with layers of gravel and clay from periods of erosion between eruptions. Cerro Castellan’s cap rock is the same lava that formed the Chisos’ South Rim. The lighter orange and gray layers beneath are tuffs.

Cerro Castellan (Castolon Peak).
A smaller peak northwest of Cerro Castellan rises above volcanic tuffs (ash deposits) in the foreground.
Cerro Castellan (Castolon Peak).

Big Bend National Park, Tuff Canyon Trail
Some of the oldest layers of volcanic rocks lie at the bottom of Tuff Canyon. It is dry most of the time, but summer thunderstorm runoff churns through the canyon, cutting it deeper. This canyon is narrower and deeper than most others in Big Bend, partly because the light gray volcanic tuff is relatively cohesive. Swift, powerful floodwaters will cut down through any kind of bedrock, but the tuff is better able to resist the widening effects of sideward erosion.

Looking down into Tuff Canyon.
Northern branch of Tuff Canyon.
Northern branch of Tuff Canyon.
Southern branch of Tuff Canyon.
Fouquieria splendens (ocotillo).
Entering the south end.
Chilopsis linearis (desert willow).
Rich entering the southern end of the canyon.
Deeper into the canyon.
Deeper still.
Eucnide bartonioides (rock nettle, yellow stingbush).
Eucnide bartonioides (rock nettle, yellow stingbush).
Cairns (not a natural feature).

4.6 mi W of Langtry
After leaving Big Bend National Park we started making our way towards the Del Rio area in Val Verde Co., where we plan to meet up tomorrow with a few other beetle collectors (Dan Heffern, Brian Raber, and Ed Riley). I noted that our path took us right by the type locality of the recently described tiger beetle, Amblycheila katzi. I didn’t have much hope of actually seeing the species, given that the season seems to have even really started yet and the earliest record of the species is the 23rd of May. Nevertheless, since we happened by the spot right as dusk was falling it seems a good idea to at least try. First we walked the limestone 2-track just to see what was out and about (just a few darkling beetles), then we started checking the limestone ledges where the tiger beetle can be found. We checked about 100 m of ledge without seeing any, and I was about ready to call it a night when we finally spotted one. It was running in a seam about 2 m above the ground and was unmistakable. My attempt tiger an in situ photo failed at first, and it almost escaped deep into a crevice before I pulled out my long forceps and pulled him out by a tarsus. It gave me a healthy pinch when I grabbed it while fumbling in my pack for a bottle, but eventually I prevailed. Later on I placed it back on the ledge and covered it with the Nalgene bottle cap, waited for it to calm down, then carefully lifted the cap and got a couple of shots before it began scurrying again. We checked another 50 m of ledge without seeing any and decided to call it a night.

Rich scanning the ground at dusk for nocturnal insects.
A wolf spider in the subfamily Lycosinae.
Amblycheila katzi (Trans-Pecos giant tiger beetle).
Amblycheila katzi (Trans-Pecos giant tiger beetle).
Selenops actophilus, one of the so-called “flatties.”

Day 6 – Comstock (prologue)
The owner of the motel in which we stayed was super friendly and kind enough to leave a key in the door for our very late arrival last night. Settling up this morning, I saw this on the wall (right next to his vaccination card—two doses) and just had to get a pic. He was only too happy to oblige when I told him how awesome it was and could I get a picture. Hey, no reason to reveal true political leanings if it means we can all just get along.

Trump Lost LOL!
Illegal tender.

Amistad National Recreation Area, Spur 406 Campground
A quick stop here on the way to meet up with Dan, Brian, and Ed near Devil’s River. There were lots of dead and dying Acacia constricta (whitethorn acacias), off of which I beat a diversity of cerambycids and buprestids from both the dead and dying branches. I found one small sapling of the same with evidence of fresh woodboring beetle larval feeding, so I collected it as well for rearing. Other than that I just collected a few weevils off of living Prosopis glandulosa (mesquite) for weevil-specialist Bob Anderson.

Apiomerus spissipes, one of the bee assassins, in flower of Opuntia engelmannii (Engelmann’s pricklypear)

Devils River near Dry Devils River
We met up with Dan, Brian, and Ed north of Del Rio and, after exchanging peasantries, followed them into a private resort surrounding a stretch of the Devils River*—considered by some to be the most unspoiled river in Texas. Dan had arranged for access after befriending Dave Barker, a commercial herpetologist who had built a home on property overlooking the Devils River and also a guest cabin on property overlooking nearby Gold Mine Canyon. We met up at the cabin and then carpooled to a spot along the Devils River where Dan and Brian had placed a variety of traps that needed servicing. While they took care of that, Rich, Ed, and I collected in the area around where the traps had been placed. I started off beating dead branches of Vachellia farnesiana (huisache), sweeping blooming Salvia sp. (sage), and beating dead branches of Acacia rigidula (blackbrush acacia) down by the river but collected only a smattering of beetles. I then clambered up the rocks and found good numbers of Acmaeodera spp. visiting flowers of Echinocereus enneacanthus (strawberry cactus) and Opuntia engelmannii (Engelmann’s pricklypear). After collecting my fill of those beetles, I returned to the riverbanks and noticed some large Carya illinoensis (pecan), from which I beat a few Anthaxia (Haplanthaxia) sp. (hoping they are one of the recently described taxa). By then, Dan and Brian had finished servicing their traps and gave me a few specimens that had been collected in their ethanol-baited Lingren funnel trap.

* The accepted usage of the name is without an apostrophe, although the reason for this is a matter of debate.

Dan (right) and Brian service a malaise trap.
Echinocereus enneacanthus (strawberry cactus).

After finishing along the Devils River, Dave invited us to his home for a few post-collecting beers. Spectacular views overlooking the river.

The author (left) with (L-R): Dan Heffern, Brian Raber, Dave Barker, Rich Thoma, and Ed Riley.

Gold Mine Canyon
We setup a variety of light stations at Dave’s cabin a little east of the river. It was warm and dry, so conditions were good, if a bit windy. My two ultraviolet light stations a bit north of the cabin ended up catching the lion’s share of cerambycids, although it was mostly elaphidiines and a Lepturges sp. We also picked up a few tenebrionids and a Carabidae crawling on the ground near the lights. Ed’s mercury vapor/ ultraviolet station on the road west of the cabin attracted a few more cerambycids, including a Lagocheirus sp. Dan, however, got the catch of the night—a Goes that came to Dan’s $6 battery-powered lantern on the road south of Ed’s station. We at first thought it might be G. novus, but that Dan later decided it was just a very lightly marked G. tesselatus. Zephyranthes chlorosolen (Brazos rain lily) blossoms were beautiful at night, their stark whiteness catching the beam of the headlamp.

Zephyranthes chlorosolen (Brazos rain lily).

Day 7 – Gold Mine Canyon
Rich and I spent the morning walking the grounds around Dave’s cabin while the others packed up and got ready to leave. I found some oak (Quercus vasseyana) saplings infested with cerambycid larvae, which I cut and bundled to bring back for rearing. Acmaeodera were already coming to the flowers—a couple of small ones on an undetermined white composite, three different species on Opuntia engelmannii (Engelmann’s pricklypear) and Echinocereus enneacanthus (strawberry cactus) flowers, and a couple on Coreopsis? flowers. I beat some of the Diospyros texana (Texas persimmon) looking for Spectralia robusta but did not find any. It got hotter than blazes real quick!

Opuntia engelmannii (Engelmann’s pricklypear).

After Dan, Brian, and Ed left, Rich and I went down to the canyon entrance to beat on the oaks and Texas persimmons that dot the sides of the canyon. Nothing was on either plant, however, and I ended up again concentrating on the diversity of Acmaeodera that were coming to flowers of Coreopsis sp., Opuntia engelmannii, and an unidentified yellow composite. I did beat a single Cleridae off a dead branch of mesquite.

Gold Mine Canyon.

Devils River near Dry Devils River
After finishing at Gold Mine Canyon, we came back to the Devils River crossing near the first stop we made here for our final stop of the day. Temperatures had maxed out at 99°F! and I wasn’t too motivated to collect much more today, but when we arrived at the spot I noticed some declining Platanus occidentalis (American sycamore) with large emergence holes suggestive of Mallodon dasystomus and old fallen branches with the same suggestive of Polycesta elata. The tree with the Mallodon holes was much too large to cut (and embedded within a thicket of poison ivy), so I occupied myself by collecting a few more Acmaeodera off of Opuntia engelmannii flowers. As I walked the roadway I noticed more sycamore with some smaller trees in the grove that looked recently dead. One was dead from about three feet up and had buprestid workings under loose, peeling bark. I cut just above the live portion (3–4” diameter) and took three 4-ft sections of the trunk above that point, each cut showing internal galleries. If P. elata emerges from these pieces of wood I will be “elated” [Later edit: I did rear the species!].

Devils River crossing.
The water was too deep for my Ford Escape.
Next time I’ll have a higher-clearance vehicle.

Gold Mine Canyon
For blacklighting tonight we decided to bring the lights down to the mouth of the canyon where we collected this afternoon so we could have acces to anything associated with the oaks. Unfortunately it was a much slower night than last night and cooled off quickly despite the high heat earlier in the day. I only got three cerambycids (one Ecyrus and two Aneflomorpha) and a few clerids at the lights. I also walked the jeep track leading to the mouth of the canyon and the main road outside and didn’t see anything until I almost got back, when I noticed a beetle sitting on the trail that looked a bit odd. When I picked it up I realized it was a buprestid in the genus Melanophila—what the heck?! Totally unexpected to see this beetle at night and especially on the ground instead of on a tree. I suppose it is one of the juniper-feeding species (since pine doesn’t occur here).

Sinking sun over Gold Mine Canyon.

Juniper cadaver in late-evening light.

Day 8 – Gold Mine Canyon
Our plan today was to head over to some spots further west in Val Verde Co., but before leaving the cabin we did a bit of walking around and took a last few photographs.

Gold Mine Canyon in the morning.
Epithelantha micromeris (button cactus).

22 mi N Del Rio
I had noted a few scattered plants of Senna roemeriana (two-leaved senna) at this spot a couple of days ago when meeting up with Dan but didn’t have the chance to sample them for Agrilus obtusus—one of my target species for the trip. I found one on the second plant I checked, so I went back to the truck to get my big camera hoping to photograph one in situ. I didn’t see anything on the next plant, but when I tapped it over my net there was another one! I did that for the next hour or so—inspecting and tapping—and never saw another one. Rich did get one sweeping the S. roemeriana (in an area I’d already worked) and was gracious enough to give it to me. There were also tiny bruchids and clerids on the plant. Other than that I got a couple of Acmaeodera mixta sweeping, a couple of Canthon sp. in flight, and a Euphoria kerni on the flower of Zephyranthes chlorosolen.

Senna roemeriana (two-leaved senna).

Hwy 90 at Del Rio River
I first visited this spot nearby 30 years ago based on a tip by Dan Heffern, who had reared a Polycesta elata from Fraxinus greggii (Gregg ash). I found the ash on that visit, though I didn’t find any wood infested with that species here, but what I did find was Diospyros texana (Texas persimmon) infested with Spectralia robusta and managed to rear out a few individuals. That was my quarry today, but when I arrived the abandoned road on the northwest side of the bridge was fenced and posted. I took a look on the southwest side and found open access up top and decided to hike down towards the ravine from that point. Things seems to be about as far along here as they were at Devils River, with not much activity except for Acmaeodera coming to the Opuntia engelmannii flowers, albeit not quite the diversity. I found a few more also on flowers of an undetermined yellow composite, Coreposis sp., and an undetermined white composite. Closer towards the ravine I found just a single large F. microphylla (with no signs of infestation) and several D. texana—two of which had the half-live/half-dead branches in which S. robusta larvae live and showing the emergence holes of adults. I collected both branches and will bring them back for rearing.

Hwy 90 bridge over Pecos River.
Adult emergence hole of Spectralia robusta in live/dead trunk of Diospyros texana (Texas persimmon).

Amistad National Recreation Area, Pecos River Access Nature Trail
Just a quick stop at the Pecos River Access on the east side to walk the short nature trail and gaze at the 300-ft high, 100 million-year-old (Cretaceous Period) limestone bluffs that the Pecos River has cut near the junction with the Rio Grande River (the latter can be seen on the left side of photo 2). The first photo also shows the old road that was originally used to cross the river snaking down the west bluff—traffic today uses the tall bridge in the right side of the photo.

Limestone bluffs over the Pecos River
Pecos River junction with the Rio Grande River.
Pecos River Access Nature Trail.

Seminole Canyon State Park, Canyon Rim Trail
We came here looking for oak potentially infested with Spectralia roburella. We didn’t find any oak on this trail, but I did find a Acacia rigidula (blackbrush acacia) showing signs of infestation by buprestid larvae (difficult to find such this year because the freeze in February apparently killed or severely knocked back most of this species). I cut up and bundled the wood to bring back for rearing.

To insects, the collectors’ shadows loom large.

Amistad National Recreation Area, Spur 406 Campground
We got to Seminole Canyon State Park too late to check with the supervisor about setting up our blacklights at the park, so we came back to Spur 406 Campground where we’d collected a few things two days earlier. Temps were okay and there was no moon or wind, but it was still a very slow night—for me just a couple of elaphidiines, two trogids, two Digitinthophagus gazella (why do I continue to pick these things up?), and a bostrichid.

Ready for another night of blacklighting.

Day 9 – Seminole Canyon State Park, Windmill Trail
We came back to the state park since we ran out of time to look for oak yesterday. The park staff were extraordinarily helpful—both in getting me checked in with my permit and in directing me to the spots where I might be able to find oak. Their first tip—along the Window Trail—paid off, where we found a nice cluster of Quercus fusiformis (plateau live oak) clinging to the upper canyon walls. Most of them had dead branches on them, and I did some beating to see if by some chance the beetles would be out already. They were not, but on the second tree that I examined I found a main branch from near the base with the outer 4–6 ft dead but the bark not peeling and small living sprouts about 2 ft from the base. Pulling apart the dead portion revealed buprestid larval workings, likely my quarry—Spectralia roburella, but these could be old. I cut the branch at the base, however, and found fresh larval galleries in the sapwood of the still-living portion even extending into the trunk—success! I’ll bring this back for rearing and will hopefully get S. roburella out of it. Further along the trail I found a single Senna roemeriana (two-leaved senna), inspected it carefully and didn’t see anything, then tapped the plant over my beating sheet and a single Agrilus obtusus fell onto it to add to the three that I got yesterday. I really wish I could see these things before I beat them off the plants so I could take an in situ photo!

The Maker of Peace, a bronze sculpture by Texas artist Bill Worrell.
View of Seminole Canyon to the east.
View of Seminole Canyon to the west. The Fate Bell rock shelter is on the right at the bend.
A vulture soars overhead.
Panoramic view of Seminole Canyon.
The author admires a fine stand of Quercus fusiformis (plateau live oak). No oaks were harmed in the making of this photo!😊
My souvenir for the trip!

Seminole Canyon State Park, Canyon Rim Trail
Another place the park staff recommended to find oaks was along the Canyon Rim Trail. We hiked that trail yesterday for a bit and didn’t see any oaks, but it turns out they were farther down the trail then we went. We headed back out on the trail to find them, along the way checking Opuntia engelmannii flowers for Acmaeodera and seeing only one for the time being. Just past the first of two east-facing ravines where we expected to find oaks, we found one on the canyon edge that looked rather bedraggled. There were some completely dead branches with bark already sloughed but also one large fresher-looking dead branch that had one live branchlet coming out of it about a third of the way up (meaning there was at least a strip of live wood within the branch). I broke of one of the dead branches near the live/dead junction, and there in its gallery was a smallish buprestid larva that almost certainly is Spectralia roburella! I took the entire branch and cut it up to bring back for rearing. We continued hiking along the canyon rim and saw the most amazing views—sheer Cretaceous limestone walls towering 300 feet above the narrow canyon bottoms! Farther down the trail we finally started seeing Acmaeodera on O. engelmannii flowers. By then we’d hiked more than a mile and a half down the trail and temps were beginning to soar, so we turned back, picked up the wood we’d cut as we came back by, and finished the long, hot slog back to the truck.

A mirid bug (Oncerometopus sp.) on flower of Viguiera dentata.
View of cave dwelling area.
Top of a Canyon!
Seminole Canyon stretches from one side to the other.
Seminole Canyon walls.

Comstock
As Rich and I were lunching after our last stop, I got a text from Ed Riley about a spot near Comstock where he’d collected what he believed to be Acmaeodera starrae—a species I’ve never encountered. It just so happened that we would be passing by Comstock on our way back east this afternoon, so we stopped to see if we could find it. Bingo—right where and in the flowers he said it would be (an undetermined white composite that I later determined to be Aphanostephus ramosissimus [lazy daisy]). Together we found about 15 specimens, and interestingly about 25% have red rather than yellow elytra markings. [EDIT: I’m not convinced these are A. starrae, but I do not yet know what they are.]

Aphanostephus ramosissimus (lazy daisy).
Aphanostephus ramosissimus (lazy daisy).

Day 10 – Garner State Park, Wild Horse Creek/Highway/Campos Trails
It’s the final day of collecting for the trip, and for our last stop we picked Garner State Park along the Frío River. I was last here back in the mid 90s—nearly 30 years ago, Acmaeodera ornatoides and Polycesta elata being the two species of note that I remember finding. I remember during that first visit that the area reminded me of my beloved Ozark Mountains, especially the White River Hills region in southwestern Missouri—scraggly forests of oak and juniper on steep, rocky slopes over craggy hill and lazy dale. It still does, although the species are a bit different—Juniperus ashei (Ashe juniper) dominates instead of J. virginiana (eastern red-cedar), and a variety of other oaks replace the familiar Ozarkian Quercus stellata and Q. marilandica (post and blackjack oaks, respectively). We hiked a series of trails on the western side of the park, thinking the west-facing slopes would tend to be drier and result in more open, glade-like habitats, and for the most part this was true. Almost immediately after reaching the first glade along Wild Horse Creek Trail, we found A. ornatoides and at least two smaller congeners on flowers of Coreopsis sp. Flowers of Viguiera dentata have been uncharacteristically depauperate of buprestids on this trip, but I picked up a couple of Acmaeodera neglecta/neoneglecta nearby as well. On the Highway Trail a good series of Acmaeodera was found on flowers of an undetermined small white composite, and a few were also found on flowers of Senna roemeriana (two-leaved senna)—though no Agrilus obtusus. The Campos Trail ascended steeply and ruggedly to a nice overlook, where I found one Acmaeodera sp. on the flower of Zephyranthes chlorosolen and then the mother-load—the biggest diversity and abundance of Acmaeodera I’ve ever seen on cactus flowers occurred nearby in a single flowering Opuntia engelmannii. The final specimen of the day’s “Acmaeodera-a-thon” was taken a bit further up the trail on the flower of Echinocereus enneacanthus (strawberry cactus). Fortunately, the trail was all downhill from there (albeit a bit too steep and rocky at times for these no-longer-nimble legs!). We finished off the hike back along the Wild Horse Creek Trail by collecting a branch off a fallen oak that I hope proves fruitful in the rearing box back home and had some lunch. As we were getting ready to leave, I noticed most of the trees in the camping area were Carya illinoensis (pecans)—a great host for buprestids (especially Xenorhipis brendeli), so I picked up several fallen branches from under the trees to complete the wood collecting portion of the trip.

Acmaeodera ornatoides on flower of Coreopsis sp.
Overlook from atop the Campos Trail.
Xeric limestone prairie (glade) habitat.
Echinocereus enneacanthus (strawberry cactus).
Echinocereus enneacanthus (strawberry cactus).

Garner State Park, Brazos River (epilogue)
We visited the nearby Frio River for one last look at the park, took a shower, and settled in for the 15-hour trek back to St. Louis.

Brazos River at Garner Stare Park, Texas.

Postscript!

Somewhere near Rising Star, Texas.

©️ Ted C. MacRae 2021

2019 Arizona/New Mexico/California Insect Collecting Trip iReport

This is the eighth “Collecting Trip iReport”; this one covering a one-week trip to southern Arizona, New Mexico, and California from September 7–14, 2019 with meloid/cerambycid-enthusiast Jeff Huether. Jeff has been a frequent collecting trip partner during recent years, this being our sixth joint outing since 2012. Our initial objective on this trip was to collect cerambycid beetles of the genus Crossidius occurring across southern Arizona/California—part of a larger effort to sample as many of the named subspecific taxa as possible from multiple locations (including type locations when possible) for future molecular studies. We had good success, though we did not collect every taxon that we were after (we were a tad early in soCal). Also, the fact that we had Crossidius as our primary goal did not mean that we would not concurrently be on the lookout for buprestids (me), meloids (Jeff), or other cerambycids (both of us)—and in that regard we were also successful.

As with all previous “iReports” in this series, this report is illustrated exclusively with iPhone photographs (thus the term “iReport”), with previous versions including the following:
2013 Oklahoma
2013 Great Basin
2014 Great Plains
2015 Texas
2018 New Mexico/Texas
2018 Arizona
2019 Arkansas/Oklahoma


Day 1 – Dripping Springs Mountains, Arizona
First stop of the trip, and we’re heading east to Safford. As soon as we got east of Superior up into the mountains we saw a place where Heterotheca subaxillaris was in bloom abundantly along the roadsides and pulled over. There were four species of Acmaeodera on the flowers, and I also found a fifth species on the flower of a small white aster. Nice first stop for the trip!

Dripping Spring Mountains.
Acmaeodera gibbula on flower of Heterotheca subaxillaris.
Acmaeodera rubronotata on flower of Heterotheca subaxillaris.
Acmaeodera alicia on flower of Heterotheca subaxillaris.

3.6 mi NW Bylas on US-70, Arizona
Continuing our way to Safford, Jeff saw some patches of sunflower and wanted to look for Epicauta phoenix. I found the first two (but not in sunflower), and then Jeff found two more. As we were walking back to the car I noticed a Crossidius suturalis sitting on Isocoma tenuisecta that was not quite in bloom, and then another nearby on the same plant. We searched the area again, but the only plants were those few right around the car.

I’ve never seen an orange jumping spider (family Salticidae) before!

5.7 mi NE Safford, Arizona
After getting a hotel in Safford, we had time to come back to a spot where Jeff had collected Epicauta phoenix back in July. We found quite a few (see photo) on plants nearby the original collection spot. Looking around more I found an Acmaeodera convicta perched on the tip of a shrub—first time I’ve collected this species! There were several species of tenebrionids crawling on the ground, perhaps prompted to activity by cooling temps as rain whipped up in the distance. I kept one eye on the skies and the other on the plants and eventually found two more A. convicta perched together on the same type of shrub just as rain began pelting my back. We made a quick dash back to the car and called an end to Day 1 in Arizona.

Epicauta phoenix (order Coleoptera, family Meloidae).

Day 2 – 1.9 mi S Artesia, Arizona
We started seeing Isocoma tenuisecta just coming into bloom as we headed south of town so stopped to see if we could find any Crossidius. I looked at a lot of plants before finding a single C. suturalis sitting on one of the non-blooming plants and in the meantime found one Trichodes peninsularis and a fair number of Zonitis dunniana on the flowers. Looking around on other plants, I found one large Chrysobothris sp. (not C. octocola, but longer and narrower) on the branch of a living Acacia constricta [Edit: this is C. knulli—a new one for me!] and one Acmaeodera disjuncta, several more Z. dunniana and T. peninsularis on flowers of Hymenothrix wislizeni. Finally, I did some sweeping of the bunch grasses in the area and got a nice series of what I presume to be Agrilus rubrovittatus—first time I’ve collected that species!

Crossidius suturalis on pre-blooming Isocoma tenuisecta.
Trichodes peninsularis on Isocoma tenuisecta.
Zonitis dunniana on flowers of Isocoma tenuisecta.

17.7 mi S Artesia, Arizona
Another stop with both Isocoma tenuisecta and also Gutierrezia microcephala coming into bloom. We immediately began finding Crossidius pulchellus on the latter and eventually collected a good series of them and also Trichodes peninsularis off the plants When I returned to the first plant we had checked (in fullest flower), a Lampetis webbii landed on it right in front of me! I eventually found C. suturalis on Isocoma tenuisecta, as well as Trichodes sp. and a few C. pulchellus. There was a tall-stemmed malvaceous shrub off which I got a male/female pair of Tylosis maculata, and sweeping produced a couple more Agrilus rubronotata, a few more T. peninsularis, and one Acmaeodera scalaris. I saw a couple of Acmaeodera disjuncta on Baileya multiradiata flowers but missed them both!

Stagmomantis limbata (bordered mantis) on Gutierrezia microcephala.
Lampetis webbii on Gutierrezia microcephala.

1.1 mi N Rodeo, New Mexico
We slipped just inside the New Mexico border to visit the area around the type locality of Crossidius hurdi. We found a spot where there were good stands of Isocoma tenuisecta along the roadsides and checked them out. Like the other spots today they were just starting to come into bloom, and rain had just moved through the area. We found perhaps 20 Crossidius individuals total, and honestly they were so variable that I don’t know whether they represent C. suturalis, C. hurdi, or both! [Edit: they are all C. suturalis] I also collected one Sphaenothecus bivittatus and several individuals each of three species of clerids on the flowers of these plants. A male Oncideres rhodosticta was found on the twig of Prosopis glandulosa, and I also found a cool meloid that I’ve never seen before—Megetra punctata!

A particularly well-marked female Crossidius suturalis on flowers of Isocoma tenuisecta.
The coloration of Megetra punctata screams “Don’t eat me or you’ll be sorry!”
Oncideres rhodosticta on Prosopis glandulosa (mesquite).
Mule deerly departed.
Fence row to the Chiricahua Mountains.

Willcox Playa, Arizona
We plan to visit Willcox Playa tomorrow (my inaugural visit!), but we had some time at the end of the day and decided to come take a look. There were some stands of Isocoma tenuisecta at the north end of the playa, and I found just a couple of Crossidius individuals on them, presumably C. suturalis, but it looks like they are bedding down for the evening. Also got a couple of Enoclerus sp. on the flowers.

North end of Willcox Playa.

Day 3 – 8.4 mi SE Willcox, Arizona
On our way towards the Chiricahua Mountains to see if we can find any Crossidius host plant stands. We found patches of Isocoma tenuisecta and Gutierrezia microcephala along Hwy 186 southeast of town—the former was just coming into bloom, but there were plenty of Acmaeodera (scalaris, disjuncta, and amplicollis) on the flowers, including on the unopened heads. We found perhaps a dozen Crossidius suturalis on them as well, and Jeff found one small female that looks like C, hurdi [Edit: it is C. suturalis]. I looked at a lot of Gutierrezia before finding a single C. pulchellus sitting on one of the plants. The same diversity of Acmaeodera as well as a few A. gibbula and T. peninsularis was also found on flowers of Hymenothrix wislizeni, and I took a series of about 10 specimens of what I looks like A. parkeri on flowers of what appears to be Stephanomeria pauciflora. There were also some tiny membracine treehoppers on a thorny shrub (maybe Condalia?) being tended by ants—both adults and young, and I collected a few of the adults.

Acmaeodera amplicollis on flowers of Isocoma tenuisecta.
Acmaeodera disjuncta on flowers of Isocoma tenuisecta.
Acmaeodera scalaris on flowers of Hymenothrix wislizeni.

Jct AZ-186 & AZ-181, Arizona
After passing over a small range towards the Chiricahuas we didn’t see any Isocoma tenuisecta until we got to Hwy 181. There were some Baccharis sarothroides at the junction also, so we stopped and looked around. The Isocoma was just barely coming into bloom, but I found two Crossidius on them—one male C. suturalis and one small female that may be C. hurdi [Edit: nope, it is C. sururalis]. Heterotheca subaxillaris was in bloom abundantly, but there were no Acmaeodera on them and the area in general looked quite dry. I did find two A. decipiens on Sphaeralcea sp., and in the way back to the car I spotted a huge Lampetis webbii hanging on Ericameria nauseosa (which we’re not even close to blooming)—surely an incidental record.

Chiricahua Mountains in the distance.

4.1 mi SE Willcox, Arizona
We came back towards town where things seemed to be further along and found stands of Isocoma tenuisecta in full bloom at the junction of Blue Sky Rd (a classic Arizona collecting locality). Crossidius suturalis were out in numbers on the flowers! Every now and then I got one that seemed too heavily maculated, making me think it could be C hurdi, but in the end I decided that all represented C. suturalis.

Crossidius suturalis mating pair on flowers of Isocoma tenuisecta. Note the difference in antennal length between the male (top) and female.

Willcox Playa, Arizona
We went to the Playa to see if there were any tiger beetles to be had. I hiked to the edge of the Playa, and within a few minutes I saw a Cicindela pimeriana—just the second one I’ve encountered (the first was last night at gas station lights)! With that promise of more, I hiked the entire playa edge and never saw another one! I only saw one other tiger beetle—Cylindera lemniscata—seems I’m a bit late in the season for the Willcox Playa tiger beetles. Nevertheless, it’s a cool place and was fun to see. I’ll definitely be back during the summer, not just for here but for nearby Blue Sky Rd. Arriving back at the car, I did find one Moneilema sp. (I think M. appressum) on cholla. There were Crossidius suturalis abundant on the Isocoma tenuisecta, which, like the last spot, was in full bloom, but I’d gotten my fill of them at the previous spot and didn’t collect any.

Stalking tiger beetles.
Jumping spider out in the playa.
Eking out a living.
Cow tracks.
These Gnathium sp. were the tiniest blister beetles I’ve ever seen.

Willcox, Arizona (epilogue)
Collecting the insects from the field is only the beginning. Each night they must be processed for storage until they can be mounted once back in the lab.

Processing the day’s catch.

Day 4 – Santa Rita Mountains, Box Canyon, Arizona
We passed through Box Canyon on our way to Madera Canyon, so we decided to stop near the dry falls where last year I’d collected such a nice diversity of Acmaeodera spp. on flowers of Allionia incarnata. There was evidence of recent rain, and we found the patch nicely in bloom with four species (scalaris, decipiens, cazieri, and parkeri) on the flowers. Nearby in the wash before it crossed the road was a yellow composite (Xanthisma gracile), from which I collected the first three as well as gibbula, rubronotata, and disjuncta. Euphoria verticalis scarabs we’re flying plentifully around the flowers also—first time I’ve seen the species.

Allionia incarnata (trailing four o’clock) blooming the canyon slope.

Flats below Madera Canyon, Arizona
There are records of Deltaspis tumacacorii from Madera Canton at Proctor Rd collected on Croton, so we stopped by on our way south to give it a try. This seems to be a rather hard-to-find bug, so I didn’t have high expectations, and that’s a good thing because I didn’t see the beetle nor anything that even remotely resembled Croton. I ended up checking out the desert broom (Baccharis sarothroides) in the area on which I’d collected Stenaspis verticalis arizonensis and Tragidion spp. (also without high expectations). There were some interesting congregations of Euphoria leucographa feeding at sap flows on the stems and a few Stenaspis solitaria but otherwise litttle of note. I did find one Hippomelas planicauda hanger-on on a low fabaceous shrub (not Mimosa biuncifera), and inspecting the Gutierrezia microcephala plants revealed nothing but a single Acmaeodera rubronotata.

Stenaspis solitaria on Baccharis sarothroides (desert broom).
Peucetia viridans (green lynx spider) feeding on Acanthocephala thomasi twice its size!
Euphoria leucographa and a Polistes paper wasp feeding at a sap flow on Baccharis sarothroides (desert broom).
Euphoria leucographa on Baccharis sarothroides (desert broom).
Taenipoda eques (lubber grasshopper). The striking coloration is a warning to potential predators that it is chemically protected.

Madera Canyon Rd, Arizona
We stopped real quick down the road on the way out of Madera Canyon because we saw stands of Isocoma tenuisecta, although they were still just shy of blooming. We looked at quite a few and found a single Crossidius suturalis—probably we are a tad early, and the area looks like it could use a good rain to pop things out and bring the Isocoma into bloom. We also saw low plants that could be the Croton that Deltaspis tumacacorii has been found on [Edit: I do not believe these are the plants, as they are too low]. Would be good to revisit this spot after a good rain!

The author with Ferocactus wislizenii (fishhook barrel cactus)—also called “compass barrel” due to its habit of leaning to the south.

Tumacacori Mountains, Walker Canyon, Arizona
Our second shot at Deltaspis tumacacorii, which has also been taken in Walker Canyon. We found thick stands of knee-high flowers that we immediately took to be the Croton—just as described by our contact—on which the beetles have been taken. However, we quickly began doubting that ID and decided the plant must be some type of composite. That would make more sense from a host plant standpoint, as all known host plants for Crossidius spp. are composites (subsequently determined to be Pseudognaphalium leucocephalum, family Asteraceae). We looked at the dense stands for quite some time but didn’t see any beetles (or much of anything else) before deciding that we were probably too early—had the beetles already emerged we would have at least found some stragglers. I did take a few Acmaeodera on the flowers (scalaris and rubronotata), as well as a large cantharid (Chauliognathus profundus). I also took single A. amplicollis and A. rubronotata individuals off of a large helianthoid composite (Viguiera cordifolia) and one A. rubronotata on a small yellow composite (Xanthisma gracile). There were a multitude of darkling beetles crawling in the ground—in one spot I saw five individuals of several species all within a one-square foot area. We’ll have one more shot at D. tumacacorii tomorrow at Kitt Peak.

Pseudognaphalium leucocephalum (white rabbit-tobacco) blooms profusely in the dry wash.
Cantharid vs. cantharid! Chauliognathus profundus (right) feeds on a C. lewisi that it has captured.

Day 5 – Pan Tak, Arizona (road to Kitt Peak)
Today’s destination is Kitt Peak to look for Deltaspis tumacacorii and Acmaeodera resplendens, but at the entrance we saw some Isocoma tenuisecta just coming into bloom and decided to check it out. We found a half-dozen Crossidius suturalis but had to really work for them. Alliona incarnata was also nicely in bloom, but I got only one Acmaeodera parkeri? and one A. alicia off of the flowers. There was some Gutierrezia microcephala present, also not quite in bloom, off of which Jeff got a pair of C. suturalis and gave me one. Kinda dry but lots of flowers—wish there would have been more beetles coming to them.

Moneilema sp. on Cylindropuntia imbricata.

Kitt Peak National Observatory, Arizona
Our last chance to find Deltaspis tumacacorii, and I also got a tip that Acmaeodera resplendens has also been taken up here. We immediately found several species of Acmaeodera (amabilis, amplicollis, decipiens, and rubronotata) abundantly on several composite flowers—Heliomeris longifolia, Heterotheca fulcrata, Hymenothrix wrightii, Solidago velutina, and Gutierrezia microcephala, and I found a single A. solitaria on a pink malvaceous flower, but no A. resplendens. We also searched thoroughly for any Croton-like plant for D. tumacacorii but found nothing. The Kitt Peak records of that species are older than the Walker Canyon, Peña Blanca, and Madera Canyon records, and most of the records seem to be in August rather than September, so I suspect we are a bit late for both the species and its host plant. My plan at this point is to return sometime during the middle of August and enlist the help of the source of one of the recent records to accompany me.

View north from Kitt Peak National Observatory.
Multiple species of Acmaeodera visiting flower of Heliomeris longifolia.
Acmaeodera amabilis on flower of Heliomeris longifolia.

Road to Kitt Peak, Arizona
We had noticed Gutierrezia microcephala and some other yellow composites in bloom about halfway up the mountain on our way to Kitt Peak and decided then to stop and take a look around on the way down. I took “down” the mountain, Jeff took “up.” I hadn’t walked very far when I saw what I at first thought was the cantharid Chauliognathus profundus (which I had seen yesterday at Walker Canyon preying on another cantharid) on G. microcephala flowers, but something about it gave me pause—it was too cylindrical and robust. I leaned closer to get a better look and realized it was a cerambycid—one that I did not recognize, a beautiful orange color with black elytral apices and pronotal spots! I quickly grabbed it with my right hand, immediately saw another elsewhere on the bush and grabbed it with my left hand, and as I stood there trying to fumble a vial out of my pack to put them in I saw a third individual taking flight from the bush and spiraling into the air and out of reach! I shouted out to Jeff, who came down to where I was, and showed him what I’d found, and together we decided that it must be Mannophorus forreri—a very uncommonly encountered species and more than adequate consolation for not finding Deltaspis tumacacorii earlier in the day. We spent the next hour searching up and down the roadsides, and I ended up with two more individuals from Gutierrezia flowers and two from Heterotheca fulcrata. Jeff found an additional individual on flowers of Thelesperma sp. I also picked up a few black and white Enoclerus sp., one on flowers of G. microcephala and a mating pair on flowers of Acacia berlandieri. We have a long drive to California in front of us now, and it sure is good going into the drive with such a great find under our belts.

Lower slopes of the road to Kitt Peak Observatory.
Thasus neocalifornicus (giant mesquite bugs) congregate on their host plant (Prosopis glandulosa).

Day 6 – Cajon Pass, California
Finally made it into California! Once we turned off the interstate, we made a quick stop to look at the roadside habitat where we spotted a good stand of Isocoma sp. in full bloom. We looked at quite a few plants but didn’t find any beetles on them. There were also good numbers of Ericameria nauseosa plants as well (host for Crossidius coralinus), but they weren’t quite yet in bloom yet and the only thing I found on them was a mating pair of Agrilus walsinghami. Moved on quickly to the next spot!

Lancaster, California
We met up with Ron Alten and traveled to a classic “Crossidius” collecting site (up to four species have been taken there). We’d stopped at a couple of places on the way there but not found anything—either the host plants were not yet blooming or no beetles were found, so we had the feeling that we might be a week or two early. We had to drive into the habitat a ways before we started seeing host plants—in this case Ericameria nauseosa—but eventually we found a nice large area with the plants in full bloom. It didn’t take long before we found Crossidius coralinus (populations in this area are assigned to subspecies ascendens) on the blossoms. We worked the area for a couple of hours in the heat (97°F) and got a sufficient series for study with some individuals in ethanol for DNA analysis. Males exhibit quite a bit of variability in the degree of development of the elytral markings (thin to moderately expanded sutural marking), while females were quite consistently fully expanded. Males also outnumbered females by 3:1, and all of the individuals I collected were perfect and not damaged—both suggesting that the species is just beginning to emerge. Perhaps that is why we did not find individuals of the other species (mojavensis, suturalis, and testaceus). What I did find, however, was a small trachyderine cerambycid that none of us recognized! It was on the flowers of E. nauseosa—just like C. coralinus—and at first I thought it might be a small, aberrant C. coralinus, but the elytra are completely blue-black and the size was significantly smaller than the smallest C. coralinus male that we saw. I scanned BugGuide and didn’t find anything that matched, so this will have to remain a mystery for now. [Edit: I later determined this to be a heavily marked C. discoideus blandus. In the field I couldn’t see the orange laterals on the elytra.]

Crossidius coralinus ascendens (male) on flowers of Ericameria nauseosa.
Crossidius coralinus ascendens (female) on flowers of Ericameria nauseosa.
Crossidius coralinus ascendens (mating pair) on flowers of Ericameria nauseosa.

Day 7 – Santa Catalina Mountains, Mt. Lemmon, Arizona
We decided we were just a bit to early for things in California and decided to come back to Arizona where we’d been having better success. I wanted to take another shot at Acmaeodera resplendens and had been told that Oracle Ridge Trail was a good locality for them, though maybe a bit late. We began seeing them soon after getting out of the car—unmistakable by their brilliant metallic green to copper color. They were not numerous, so I had to work for them and walked the trail about 2 miles out collecting them off a variety of flowers. The majority were on Bahia dissecta, and I also found occasional individuals of them and other species of Acmaeodera (amabilis, amplicollis, decipiens, and rubronotata) on flowers of Heliomeris longifolia, Heterotheca fulcrata, Hymenothrix wrightii, Ageratina herbarea, Achillea millefolium, sweeping, Cirsium sp., and prob. Viguiera dentata. One other beetle I found was a Megacyllene sp. sitting on a plant under a stand of Robinia neomexicana [Edit: this is M. snowi snowi—another new one for my collection!].

View from Oracle Ridge Trail @ 1 mile north of the trailhead.
Acmaeodera resplendens on flower of Heliomeris longifolia.
Acmaeodera resplendens on flower of Heliomeris longifolia.

Scenic Overlook, Santa Catalina Mountains, Arizona
A quick stop on the way back down the mountain at a spot where we’d seen Gutierrezia microcephala and Heterotheca subaxillaris blooming along the sides of the road. There wasn’t much going on—a couple of Acmaeodera amplicollis and one A. rubronotata on the flowers of H. subaxillaris, one Enoclerus sp. on Solidago velutina, one A. solitaria on Baccharis sarothroides, and another swept from grasses. I did see Peucetia viridans (green lynx spider) feeding on a very bristly tachinid fly.

A clearwing moth (family Sesiidae).
Peucetia viridans (green lynx spider) feeding on a hairy tachinid fly.

Day 8 – Santa Rita Mountains, Montosa Canyon, Arizona (halfway up)
We decided to visit Montosa Canyon to take another shot at Deltaspis tumacacorii and also see if maybe we could find more Acmaeodera resplendens. We didn’t see many flowers along the way up the canyon until about the halfway point. When we did start seeing them we made a quick stop to see what might be on them. I collected some of the more common Acmaeodera (rubronotata, decipiens, and amplicollis) off a few different yellow composite flowers, but we quickly decided to take a look at the higher elevations.

Panoramic view from halfway up the canyon.
Apyrrothrix araxes (dull firetip skipper) on flowers of Baccharis salicifolia.
The larvae of these large skippers feed on oaks.

Santa Rita Mountains, Montosa Canyon, Arizona (entrance to Whipple Observatory)
The road was gated past the km-13 point—Jeff took the roadsides, and I took a ridgetop trail off to the south for a little over a mile. The panoramic views were spectacular, and at the southern terminus I stood at the edge amidst gale-force winds admiring the landscape! Acmaeodera were diverse and abundant, though not quite as abundant as yesterday on Mt. Lemmon or a few days ago on Kitt Peak. However, I did get another nice series of A. resplendens, along with decipiens, rubronotata, amplicollis, and amabilis. Host flowers were collected for most of these: Hymenothrix wrightii, Erigeron neomexicanus, Linum neomexicanum, Heliomeris multiflora, Verbesina enselioides, Heliopsis parvifolia, Heterotheca subaxillaris, and Machaeranthera tanacetifolia. I also collected a small series of A. decipiens perching on grass stems and a very cool-looking wasp—black with a bright orange thorax and whitish abdominal apex [edit: I believe this is the scoliid Psorthaspis portiae].

Vista from the southern terminus of the ridgetop trail off Mt. Hopkins Rd at km 13.
Acmaeodera decipiens on flower of Machaeranthera tanacetifolia.
A curious assemblage of bees on this Heliomeris longifolia flower. They were not active, just sitting. [Edit: these are Dufourea sp. (short-faced bees, family Halictidae)].

Santa Rita Mountains, lower Montosa Canyon, Arizona
We stopped at a spot near the bottom of the canyon on the way out to see what was going on at the lower elevations. The answer—not much! There were a variety of woody shrubs and other plants in bloom, but the area seemed rather “wilty”. I think this area has a lot of potential, we just didn’t hit it at the right time. I did take an impressively huge tarantula hawk, just because.

Fouquieria splendens (ocotillo) resembles cacti but is not a true cactus. Distantly related to persimmons, blueberries and acacias, it is now placed in its own family (Fouquieriaceae).
Apiomerus flaviventris (yellow-bellied bee assassin) with prey.
The prey is a soldier beetle (likely Chauliognathus lewisi).
Robber flies not only mate tail-to-tail, they fly coupled in the position also. This tandem of giant robber flies (possibly Promachus nigrialbus), flew by me and landed in the bushes. The male (right) tried to take flight again and pulled the female’s hind legs off her perch. She stood firm, however, forcing the male to grab a nearby branch with just his from and middle legs and leave his hind legs dangling also. Note that the female is also feeding on a honey bee—so much natural history going on here!

Flats below Montosa Canyon, Arizona
I was a bit disappointed at not finding any beetles at what seemed would be the last collecting stop of the trip. But on our way out we saw a patch of Isocoma tenuisecta in bloom in the lowlands some distance west of the entrance to the canyon—just what we were looking for! Jeff and I each quickly found Crossidius suturalis individuals on flowers of the plants and continued searching up and down along the roadway. We didn’t find any more for awhile but when I got back to the area where I started I spotted another one sitting on a plant on the other side of the barbed-wire fence. I extended my net handle to the max, maneuvered it in position, took an assertive swipe, and got it. Just as I was putting it into the bottle, I saw another one take flight from a plant right beside me. I hadn’t closed the bottle yet but didn’t want the other one to get away, so I capped my thumb over the opening, awkwardly wielded my net into position one-handed, chased after it and took a swing and got it, too! (More often than not these situations end up with me losing both specimens!). There was also a good amount of Hymenothrix wislizeni along the roadside, off the flowers of which I collected several Acmaeodera gibbula, A. disjuncta, and A. rubronotata. This is probably the last collecting locality of the trip, so I’m happy to end up having success with this subspecies of Crossidius (C. suturalis intermedius), which we havn’t found in large numbers on this trip. Just after leaving the site, we saw a bobcat on the side of the road—my first one! Unusual to see one in the middle of the day—it was a small one, must’ve been quite hungry!

Santa Rita Mountains from the highway.

Phoenix, Arizona (epilogue)
Bill Warner was kind enough to host Jeff and I for our last night in Arizona prior to returning home tomorrow. What an amazing collection he has built, and his use of flight-intercept traps in recent years has turned up even more amazing beetles. I was happy to also meet Andrew Johnston and Evan Waite, who joined us for dinner.

Bill Warner, an icon among Arizona beetle collectors, sits amidst newly collected I material waiting to be processed.
Sunset in Phoenix!

©️ Ted C. MacRae 2019

2019 Arkansas/Oklahoma Insect Collecting Trip iReport

Alternative title: Rich and Ted’s Excellent Adventure

This is the seventh “Collecting Trip iReport”; this one covering a 5-day trip to Arkansas and Oklahoma from June 7–11, 2019 with my friend and local collecting buddy Richard Thoma. As with all previous “iReports” in this series, this one too is illustrated exclusively with iPhone photographs.

Previous iReports include the following
2013 Oklahoma
2013 Great Basin
2014 Great Plains
2015 Texas
2018 New Mexico/Texas
2018 Arizona

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…and so it begins!

Day 1 – Ozark National Forest, vic. Calico Rock, Arkansas
It’s been many years since I’ve visited these sandstone glades overlooking the White River near Calico Rock. Conditions were partly sunny when we arrived, but water on the ground suggested rain earlier in the day. We had only a short time to start exploring before the wind started blowing up and the smell of rain filled the air. I did manage to beat one Amniscus sexguttata from a branch of living Pinus echinata and collect a couple of Strigoderma sp. from Coreopsis lanceolata flowers before steady rain forced us to retreat.

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White River near Calico Rock, Arkansas—before the rain.
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White River near Calico Rock, Arkansas—rain’s a comin’!

Day 2 – Ouachita National Forest, Winding Stair Campground & Ouachita Trail, Oklahoma
We walked the trail from the campground S about 2½ miles and back. I started off with Acmaeodera tubulus on Krigia sp. flowers, eventually finding a lot of them on this plant at higher elevations along with a single Acmaeodera ornata, and I beat a few Agrilus cepahlicus off of Cornus drummondii. This had me thinking it would be a good buprestid day, but it wasn’t, the only other species collected being some Chrysobothris cribraria off of small dead Pinus echinata saplings and Pachyschelus laevigatus on Desmodium sp. Also beat a few miscellaneous insects off of Cercis canadensis and Vaccineum arborea and swept some from grasses and other herbaceous plants. Back at the campground I collected Chrysobothris dentipes on the sunny trunks of large, live Pinus echinata trees.

Emerald Vista, along the Talihema Scenic Drive.
Acmaeodera tubulus on flower of Rosa carolina.
Acmaeodera tubulus on flower of Rosa carolina.
Canthon sp.
Apheloria virginiensis reducta (ID by Derek Hennen).
The biggest cairn I’ve ever seen.

Ouachita National Forest, Talimena Scenic Dr at Big Cedar Vista, Oklahoma
There were lots of native wildflowers like Coreopsis tinctoria and Ratibida columnifera in bloom, so we stopped to check them out. There were lots of butterflies, however, I found only a single Typocerus zebra on Coreopsis lanceolata.

View south from Talimena Scenic Drive at Big Cedar Vista.
Coreopsis tinctoria (plains coreopsis).
Ratibida columnifera (upright prairie coneflower).

Ouachita National Forest, Winding Stair Campground, Oklahoma
We returned to the campground in the evening to do some blacklighting. I had high hopes, but only five cerambycids came to the lights, all represented by a single individual: Monochamus carolinensis, Acanthocinus obsoletus, Amniscus sexguttatus, Eutrichillus biguttatus, and Leptostylus tranversus (the first four are pine-associates). I also picked up a few other miscellaneous insects.

Rich processes the day’s catch as the blacklight hums in the background.

Day 3 – Medicine Park Primitive Campground, Oklahoma
There wasn’t much insect activity going on in eastern Oklahoma, so we drove out west to the Wichita Mountains for hopefully better luck. We found a small park with a primitive campground in the city of Medicine Park—my first thought was to beat the post oaks dotting the campground, but when I went into the native prairie between the campground and the creek I never came out! Right away I found what must be Acmaeodera ornatoides on flowers of Opuntia sp., then I found more on flowers of Gallardia pulchella along with Acmaeodera mixta. The latter were also on flowers of Thelesperma filifolium along with Acmaeodera neglecta—took a nice series of each, and I also got a few of the latter on flowers of Coreopsis grandiflora. Strangalia sexnotata were on flowers of C. tinctoria and Torilis arvensis, and then on the latter plant I saw a male Strangalia virilis—a Texas/Oklahoma specialty that I’ve never collected before! I spent the next hour looking for these guys and ended up with 3 males and 2 females along with a few Trichiotinus texanus—another Texas/Oklahoma specialty—and a single Agrilaxia sp. nr. flavimana (could be A. texana). One single Typocerus octonotatus was on flowers of Achillea millefolium. I think we may come back here tomorrow—I’d like to look for more S. virilis and beat the post oaks (the reason we stopped here to begin with).

A cacophony of native wildflowers!
An orgy of Euphoria kernii (Kern’s flower scarab) in Opuntia sp. flower. Multiple color forms exist for this species.
At first I thought this was a type of hover fly (family Syrphidae), but eventually I determined it to be Esenbeckia incisuralis, a horse fly (family Tabanidae)—incredible emerald green eyes!
Papilio polyxenes asterius (black swallowtail) caterpillar.
Echinocereus reichenbachii baileyi (lace hedgehog cactus).

Lake Lawtonka nr. Ma Ballou Point, Oklahoma
We stumbled into this area while looking for stands of Sapindus drummondii (soapberry)—found a small stand along the road, but it was too inaccessible. The same diversity of blooms were present as at the previous spot, so I picked a few longhorns off flowers of Coreopsis grandiflora and Gaillardia pulchella. Super windy, so we didn’t stay long.

View across Lake Lawtonka from Ma Ballou Point.
Neochlamisus sp. (case-bearing leaf beetle) larvae inside their “casas de caca” on Monarda fistulosa (bee balm).
I believe this is Harrisina coracina, a leaf skeletonizer in the family Zygaenidae. Both BugGuide and the Moth Photographers Group show records only from Texas.

Day 4 – Wichita Mountains National Wildlife Refuge, Oklahoma
Before starting the day’s collecting, we wanted to go into the Wichita Mountains National Wildlife Refuge to have a look around. On the way into the refuge we some American bison near the road and had to stop, take photos, and simply admire these massive, majestic beasts. We then went to the Cedar Plantation, where I had visited before back in 2012 and photographed black individuals of Cicindelidia obsoleta vulturina (prairie tiger beetle). No tiger beetles were out now (they come out in the fall), but I’d hoped to maybe see Cylindera celeripes (swift tiger beetle) along the 2-tracks in the area. No such luck—nevertheless, we saw a myriad of interesting insects, including several more Esenbeckia incisuralis (green-eyed horse flies) and a beautiful Trichodes bibalteatus (checkered beetle), the latter of which I photographed on flowers of Ratibida columnifera and Achillea millefolium with the big camera. Afterwards we visited the “prairie dog town” and got marvelous views and photographs of black-tailed prairie dogs.

Native American wildlife on a native American landscape.
American bison (Bison bison bison).
Wichita Mountains from Cedar Plantation.
Acmaeodera mixta on flower of Rudbeckia hirta (black-eyed Susan).
Typocerus octonotatus on inflorescence of Achillea millefolium (yarrow).
Strangalia sexnotatus on flower of Ratibida pinnata (gray coneflower)
Trichodes bibalteatus on flower of Rudbeckia hirta (black-eyed Susan).
Trichodes bibalteatus on flower of Ratibida pinnata (gray coneflower)
The author walks a bison trail through the Cedar Plantation.
Black-tailed prairie dog at its burrow entrance.
“Watch you lookin’ at, Willis?!”
Black-tailed prairie dog (Cynomys ludovicianus).

Medicine Park, Jack Laughter Park, Oklahoma
We’d noticed this spot yesterday because of the old post oaks and wealth of wildflowers blooming up the mountainside. There wasn’t much going on today, however—just a few Acmaeodera mixta on flowers of Gaillardia pulchella. I did find an Anthaxia (Haplanthaxia) sp. on my arm! Otherwise I spent some time photographing the landscape and some geometrid larvae on flowers of Gaillardia pulchella.

Opuntia sp. (prickly-pear cactus) amidst rocky exposures.
Small Oncoptus fasciatus (large milkweed bugs) nymphs colonize seed pods of Asclepias asperula (antelope-horns).

Medicine Park Primitive Campground, Oklahoma
We returned to this spot since we had so much luck yesterday. I was hoping to collect more Acmaeodera ornatoides and Strangalia virilis, but there was much less going on today than yesterday—basically didn’t see anything for the first hour and a half. I didn’t give up, however, and kept checking the area where we saw most of the S. virilis yesterday, and eventually I saw another male in the same area as yesterday on the same stand of Torilis arvensis. I found two more males in the same area over the next hour, so three males on the day was a good reward for the time spent looking for them. I also collected Trichodes apivorus and Trichiotinus texanus on flowers of Allium sp. Interestingly, beating the post oaks—the reason why I originally wanted to stop here—produced nothing. So, not very many specimens on the day, but happy with those I did get.

Thelesperma filifolium (stiff greenthread).
Coreopsis grandiflora (large-flowered tickseed).
Gaillardia pulchella (firewheel).
Coreopsis tinctoria (plains coreopsis).
Torilis arvensis (erect hedge parsley), introduced.
Opuntia sp. (prickly-pear cactus).
Opuntia sp. (prickly-pear cactus).
Allium sp. (wild onion).

Medicine Park, Jack Laughter Park, Oklahoma
I was pretty much done for the day after spending all morning at the refuge and most of the afternoon at the previous spot, but Rich wanted to take another look at Jack Laughter Park because he’d found some interesting grasshoppers there. As with earlier in the day there were few beetles of interest to me, but I did collect a couple of Trichiotinus texanus on flowers of Cirsium undulatum. I checked out some large post oaks with large dead branches thinking that might be what Strangalia virilis was breeding in but never saw any, and eventually I turned my attention to photographing a few interesting native plants that I found along the way.

Krameria lanceolata (trailing krameria).
Cirsium undulatum (wavyleaf thistle or gray thistle) inflorescence.

Cirsium undulatum (wavyleaf thistle or gray thistle) can be distinguished by its wavy leaves that are gray-green on both upper and lower surfaces.

Day 5 – Epilogue
We were tempted to do one last little bit of collecting on the way back to St. Louis, but since had pretty good luck during the last couple of days and the drive alone would take more than nine hours we decided to leave well enough alone and get home at a reasonable hour. A walk with Beauregard when I got home to stretch the post-drive legs was the perfect way to end the mini-vacation.


©️ Ted C. MacRae 2019

“62nd” Annual “Season-Opener-Birthday-Bug-Collecting-Trip”

This past Tuesday was my birthday, and as I have done for my entire adult life I took the day off and went on my traditional “Season-Opener-Bug-Collecting-Trip”. In the past I’ve usually just had one friend go with me, but today I had three—Rich, Chris, and Bill!

We started off by heading south about 3½ hours to Tingler Prairie Natural Area in Howell Co.. I wanted to come here because: 1) it was a place I’d never visited before, and 2) there is a population here of the endemic Ozark trillium (Trillium viridescens) that I thought might be in bloom. Rain in the forecast and a patch of the real thing on the way down had us a little worried, but the day turned out spectacular with temps climbing to 80°F and skies partly to mostly sunny.

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Tingler Lake – the centerpiece of Tingler Prairie Natural Area.

We never did find the trillium—even calling another friend (Casey), who had seen them here before to find out exactly where he saw them. Turns out we hit the right spot, but apparently we were too early, especially given how late spring has been moving along this year. Nevertheless, we still found much to capture our interest, including a yellow-bellied racer (Coluber constrictor flaviventris)—too quick for photos—and a black rat snake (Pantherophis obsoletus) that coiled up nicely under the end of a fallen log and tolerated my prodding and “cleaning” of the scene around him to get some good photos (albeit, only with the iPhone).

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Black rat snake (Pantherophis obsoletus).

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The “cloudy” eye is actually an artifact of the flash lighting reflected by the retina.

Unfortunately, insect activity was very light. We saw no tiger beetles and only the earliest spring forms of most other insects. I took about 10 specimens each of the early-spring jewel beetles Acmaeodera tubulus and Pachyschelus purpureus, which I found on flowers and foliage (respectively) of wild geranium, Geranium maculatum. It was the first time that I’d found the latter as abundant as this, so the series of specimens will be a welcome addition to my cabinet. I also took a couple of A. tubulus on a flower of violet wood sorrel, Oxalis violacea and showed the others the characteristic end of an oak twig pruned by a twig pruner (Anelaphus parallelus) larvae (and revealed the culprit for them to see). Despite the paucity of insects, it was a beautiful and high-quality spot, and I look forward to collecting here again sometime when the season is more advanced.

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Several adult Acmaeodera tubulus feeding on petals of wild geranium (Geranium maculatum) flowers.

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Twig pruner, Anelaphus parallelus, larva cut from its gallery near the base of a pruned oak twig.

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Revealing the culprit while Chris takes notes on some of the birds he’s been hearing.

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Redspotted purple butterfly, Limnetis arthemis astyanax (family Nymphalidae), caterpillar on black cherry (Prunus serotina), one of its favorite host plants. The caterpillar mimics bird droppings for protection.

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Fire pink, Silene virginica (family Caryophyllaceae) thrives on a dry, cherty hillside.

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Mayapple, Podophyllum peltatum (family Berberidaceae) prefers the rich, more mesic hillsides (this is where we expected Trillium viridescens to occur).

After finishing up at Tingler Prairie, we drove east an hour and bushwhacked over rough Forest Service 2-track to Bald Hill Glade Natural Area in Ripley Co. It had been close to 10 years since my previous visit, but I remembered the roads well. We had to park, however, about a mile from the entrance to the glade due to fallen trees across the final stretch 2-track. The landscape had changed considerably since my last visit—gone was the dense, close forest lining the 2-track, and in its place stood open woodland brought to this condition by active management that included the use of prescribed burning apparently as recently as the past season.

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Dwarf iris (Iris cristata).

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These diminutive plants are distinguished from other membe