Art and Bob and Ted’s Excellent Adventure

Back in mid-July, I was fortunate to have the chance to host a couple of beetle colleagues from out-of-state and to show them a few of Missouri’s premier habitats. Art Evans is a scarab specialist from Virginia who has published several books on beetles—most recently “Beetles of Eastern North America” and “Beetles of Western North America.” Bob Anderson is a weevil specialist from the Canadian National Collection in Ottawa. I’ve been in the field with Art a couple of times, both out west in Arizona, while this is my first time in the field with Bob. They are passing through as they work their way west for a 4-week collecting trip, and since neither has ever collected in Missouri it was a perfect opportunity to spend a few days together.


Sand Prairie Conservation Area

Photo by Art Evans.

We met up at Sand Prairie Conservation Area in southeastern Missouri, which contains a high quality sand prairie remnant—one of Missouri’s rarest and most endangered natural communities. This is the place where a few years ago I discovered two scarab beetles not previously known to occur in Missouri—including Strategus antaeus (smooth ox beetle), the second largest beetle in eastern North America.

L-R: Me, Art Evans, and Bob Anderson (representing 195 years of entomological expertise!).

I was hoping there might still be jewel beetles (family Buprestidae) on the Quercus spp. (oaks) and Diospyros virginiana (persimmon) trees ringing the perimeter of the sand blowout area, and my hopes increased when I swept two Dicerca obscura off high branches of the latter. Those would be the last buprestids I saw there. Sweeping the high branches of Quercus marilandica (blackjack oak) and southern Q. falcata (red oak) produced only epitragine tenebrionids.

Me using a long-handled net to shake loose buprestids and cerambycids from Diospyros virginiana (persimmon). Photo by Art Evans.

I looked for live individuals of S. antaeus but, as is typical, only saw bits and pieces of carcasses near the base of certain oaks. Under one I did collect an almost intact female carcass (missing only the head), and the head and pronotum of a super-major male.

Strategus antaeus carcass remnants found under Quercus marilandica (post oak) at sand blow perimeter in xeric sand prairie remnant. Photo by Art Evans.

Continued sweeping of high branches finally produced something besides epitragines—an Enoclerus sp. Returning to the vehicles, I exchanged my long-handled net for a sweep net and swept the Polygonellum americanum (American jointweed), collecting only a single Cryptocephalus sp. For their part, Bob and Art collected a small variety of beetles, including some weevils that Bob found interesting.

Stylisma pickeringii (Pickering’s dawnflower) in dry sand prairie remnant.

Otter Slough Conservation Area

After Art and Bob had seen enough of Sand Prairie, I took them to Otter Slough Conservation Area. This area features sloughs and wet bottomland forests where I’ve collected many good species, most notably Agrilus concinnus—formerly considered very rare until I determined it to be a later-season species associated with species of Hibiscus (rose mallow)—and an undescribed species of Taphrocerus that I’ve found on Carex hyalinolepis here but nowhere else in the state (despite finding the host plant). I alerted Art and Bob to these possibilities and started down the 2-track where I’ve seen these species before.

My beating sheet has seen better days. Photo by Art Evans.

Unfortunately, the 2-track was completely overgrown to the point of being impassable (it’s been many years since I last walked it), so I opted instead to walk the perimeter of Mudsnake Marsh where I’ve also seen the two species. The marsh was dry—first time I’ve seen it like that, allowing me to check Hibiscus plants in the marsh interior as well as along the edge. No A. concinnus were seen on any of the plants—just a few Paragrilus tenuis and good numbers of Chaetocnema quadricollis (hibiscus flea beetle).

Chaetocnema quadricollis on Hibiscus lasiocarpus (rose mallow) in lowland wetland.

Failing to find A. concinnus, I swept the Carex along the perimeter on the edge of the wet bottomland forest side and even in a spot where I had collected Taphrocerus abundantly in previous years, but to no avail and finding instead only one chrysomelid. Collecting was slim at Sand Prairie, and it was even slimmer here at Otter Slough, so it appears seems we have definitely entered the “summer doldrums” stage of the collecting season—at least for beating and sweeping.

Owl-fly (family Myrmeleontidae), probably genus Ululodes, sweeping sedges in lowland wetland.

Holly Ridge Natural Area

After going into bearby Dexter to get a motel room and eat some dinner (Dexter Bar-B-Q pulled pork—pretty good!), I took Art and Bob to Holly Ridge Natural Area for an evening of blacklighting. I wanted to come here to 1) avoid the hoards of aquatic insects that would come to our sheets if we blacklighted at Otter Slough and 2) give myself a chance (however outside) of getting Saperda obliqua—known from only a single specimen collected at Hawn State Park but likely here as well due to the stands of Alnus serrulata (hazel alder).

Photo by Art Evans.

Bob and Art agreed it looked like a good spot to blacklight when we arrived, so Art placed his light setup in the parking lot, while I placed mine about 100 meters into the mesic lowland deciduous forest. I had high hopes for the evening—it was warm (89°F) and humid, and we were two days past the full moon so moonlight would not be an issue for at least two hours after sunset.

Art Evans collects beetles at a mercury-vapor light.

A lot of insects ended up coming to the lights, but not a lot of cerambycids (and certainly not S. obliqua). Nevertheless, I picked up one Enaphalodes atomarius and a few Lepturges confluens, along with some telamonine treehoppers and other miscellaneous beetles—all at my light setup.

Enaphalodes atomarius (robust oak borer) at UV/MV light in mesic lowland deciduous forest.

We also saw Elytrimitatrix undata and several other miscellaneous beetles on the trunk of a large standing Quercus sp., and E. atomarius and another E. undata at Art’s lights, all of which I let him collect. He was also happy to see the many Lucanus capreolus and Neocicada hieroglyphica that came to both our lights. By 10:30 pm few additional insects were coming to the lights, so we broke them down and heads back to Dexter.

Elytrimitatrix undata (family Disteniidae) nocturnally on trunk of large standing Quercus sp. in mesic lowland deciduous forest.
Neocicada hieroglyphica (hieroglyphic cicada) at UV/MV light in mesic lowland deciduous forest.
Callosamia angulifera (tuliptree silkmoth) at UV/MV light in mesic lowland deciduous forest.
Phobetron pithecium (hag moth) at UV/MV light in mesic lowland deciduous forest.
Dolomedes albineus (whitebanded fishing spider) nocturnally on trunk of large standing Quercus sp. in mesic lowland deciduous forest.

Long Bald Glade Natural Area

It took us all morning to drive across southern Missouri to reach this near the easternmost limits of the White River Hills, a fantastic region in the extreme southwest of Missouri featuring dry oak/juniper woodland surrounding extensive dolomite glades atop rounded knobs. It is my favorite region of the state for collecting insects, and I wanted Art and Bob to see the area before they continue on to western Texas. I also have jug traps and a Lindgren funnel trap placed here, so coming here would give me a chance to service them before checking the rest during the next two days on the way back to St. Louis.

By the time we arrived, temperatures were already soaring, and I was disappointed to see that conditions were very dry. Nevertheless, you never know how collecting will be until you try. I started out by checking the Lindgren funnel trap, which I had placed in actively-restored dry post oak woodland and was pleased to see a variety of beetles, including a series of Neoclytus scutellatus. In addition to re-baiting with ethanol-only, I added a pheromone lure (Fuscumol Lure MR, #P655-MR, Chemtica International) to the trap to increase the attractiveness of the trap to cerambycids.

Bob Anderson watches as I service a Lindgren funnel trap.

The ethanol/red wine trap had a nice variety of beetles that caught the attention of Art and Bob and included Plinthocoelium suaveolens, Stenelytrana emarginata, Eburia quadrigeminata, undetermined elaphidiines, Euphoria fulgida, E. sepulchralis, Cnephus mutilatus, miscellaneous beetles, and a cicada. The ethanol-only trap, however, was much less productive, yielding no cerambycids and just one E. sepulchralis, a few C. mutilatus, and some miscellaneous beetles. This continues the trend noticed four weeks ago during the first round of trap collections, when it appeared that the ethanol/red wine traps yielded higher numbers and diversity of not only beetles but also bycatch of other insects (primarily moths, flies, and wasps), suggesting that red wine possesses additional components attractive to beetles that more than make up for the reduction in ethanol content resulting from mixing the two.

Servicing a “jug” trap.
“Jug trap” catch.

The condition of the glade vegetation was extremely dry, and as I walked between the traps I didn’t see a single plant in bloom. I encountered Bob on the way back to the car and mentioned this to him, and I suggested to both Art and Bob that instead of spending time here we should look at locations further west that may have gotten more rain. They agreed, so we cut our visit short and headed further west.


Mincy Conservation Area

As we traveled west, we passed two locations where I had placed traps, but the entire area still looked exceedingly dry so we didn’t stop. By the time we reached Branson, however, conditions looked much better, and I suggested to Art and Bob that we visit Mincy Conservation Area just south of town since we were now in an area that looked like it had received some rain. Mincy is another of my favorite localities in this area due to the presence of high-quality dolomite glades, and in fact it is one of the localities where I have placed jug traps. Beetle numbers and diversity were much higher in the ethanol/red wine trap, which had Plinthocoelium suaveolens, Eburia quadrigeminata, elaphidiines, Neoclytus scutellatus, Strangalia luteicornis, Acmaeodera texana, Cotinis nitida, Euphoria fulgida, E. sepulchralis, elaterids, Cnephus mutilatus, mordellids, and a cicada, while the ethanol-only trap yielded most of the same but in lower numbers. While servicing the traps, I noticed an Acmaeodera on the flower of Liatris hirsuta (hairy blazingstar). I at first assumed it was A. pulchella, a common summertime species here, but looking more closely at it I realized it was A. texana, a very uncommon species in Missouri that I have not seen for many years. I looked at other Liatris flowers but didn’t see any, so I swept the surrounding herbaceous vegetation of this xeric dolomite prairie and found one more (plus a few chrysomelids). We had originally planned to go further west to Roaring River State Park for an evening of blacklighting. Hiwever, seeing that the vegetation looked good here and that the area had obviously gotten some rain, I suggested that we stay here instead of taking a chance on moving to an area where we were not sure what conditions were like. This also would allow us to spend more time collecting—we could go into Branson for dinner and return here quickly rather than driving another hour to Roaring River. Art and Bob agreed this was a good idea, so we headed into town and enjoyed Mexican cuisine at Los Poblanos (I had crispy tacos and, in a true rarity, resisted the siren call of the beans, rice, chips and salsa that would have totally derailed my 3-week stretch of healthy eating).

Acmaeodera texana on flower of Liatris hirsuta (hairy blazingstar) in xeric dolomite prairie remnant.

We returned with still about an hour before it would start getting dark, so I decided to continue sweeping the glade vegetation to look for more Acmaeodera texana. I paid particular attention to any flowers (primarily Liatris hirsuta and Rudbeckia missouriensis), since those were the plants most likely to have the beetles on them. Most sweeps, while not yielding A. texana, did produce hispine leaf beetles in the genera Anisostena and Microrhopala—certainly interesting enough to collect and motivate me to continue sweeping. After going through one particular area and looking at the sweep contents on the net, I saw the unmistakable shape of an Agrilus. This was not, however, just any Agrilus, but rather A. impexus—one of North America’s rarest Buprestidae! I recognized it because some years ago I received two specimens from another person who collected them sweeping prairie vegetation at Ha Ha Tonka State Park in west-central Missouri. I identified them as this species but noted they were much larger than specimens collected commonly in the southwestern U.S. I sent the specimens to Henry Hespenheide, who not only confirmed their identity but also determined they were not conspecific with a more common but as yet undescribed southwestern species going under the same name. In reality, true A. impexus is very rare, known from only a handful of specimens—most many decades old—collected in the tallgrass as prairie region of the central U.S., and the common but unnamed southwestern species was described as A. paraimpexus. I have swept tallgrass prairies abundantly ever since but failed to find the species—until now. I alerted Art and Bob to the find and worked up to the brink of darkness sweeping the area to look for more. I never did, but Bob, on his last sweep of vegetation before closing darkness, found another (right in the same area where I had collected mine)!

Acanthepeira stellata (starbellied orbweaver) swept from herbaceous vegetation in xeric dolomite prairie remnant.

As exciting as this find was, darkness prevented continuing to look for it, and I had to take advantage of what little light remained to setup my lights. I felt a few sprinkles as I did this but didn’t think much about it, assuming it would pass, and fired up the generator. The sprinkles continued, however, and gradually increased to the point where I worried about the mercury-vapor bulb. Art had also set up his lights but waited on turning on the mercury-vapor light, and together we hoped against hope that it would blow over and we would be able to proceed with blacklighting. It was all in vain though, as temps continued to drop and light sprinkles turned to steady drizzle. With darkness well developed and absolutely no insects flying around the ultraviolet lights (which we had left on, it was clear that it would be pointless to continue. With that, we took our lights down and said our goodbyes, as their plans were to head towards Texas in the morning while I went to Roaring River to begin checking the rest of my jug traps as I worked my way back to St. Louis. It seemed an inappropriately inauspicious end to an otherwise successful stop and fun trip with two exceptional coleopterists. I’m already looking forward to the next chance I get to spend time in the field with each of them!

Asclepias viridiflora (green milkweed, green comet milkweed, green-flowered milkweed) in xeric dolomite prairie remnant.

Roaring River State Park—Chute Ridge Glade

First stop of the day to service jug traps, and the area has apparently gotten more rain as the glade vegetation looked reasonably lush. I was hoping to see more Glycobius speciosus here after getting a single individual in the ethanol/red wine trap last time, but that was not the case. The trap was, however, still full of beetles, including numerous Plinthocoelium suaveolens and a variety of other longhorns such as Eburia quadrigeminata, Elytrimitatrix undatus, and elaphidiines, scarabs like Cotinis nitida, Euphoria fulgida, and E. sepulchralis, and other beetles such ad elaterids, one Chrysobothris chlorocephala?, one Chariessa pilosa, mordellids, and Cnestus mutilatus. The ethanol-only trap, as before, had fewer individuals and lower diversity. Between checking the traps, I swept the herbaceous glade vegetation and didn’t get much, but after checking the second trap I swept along the roadside and got five more Agrilus impexus, effectively doubling the known series of this species! I was very happy to have collected more of this very rare species after the two that Bob and I collected yesterday at Mincy Conservation Area.

Asclepias verticillata (whirled milkweed) in xeric dolomite prairie remnant.

Hercules Glades Wilderness

The jug traps here didn’t produce much last time, and I wasn’t expecting much this time either. The ethanol/red wine trap had Plinthocoelium suaveolens, Eburia quadrigeminata, a few elaphidiines, one Cotinis nitida, several Euphoria fulgida, a couple of elaterids, and a clerid, while the ethanol-only trap had one cicada, one elaterid, and one miscellaneous beetle. I also collected Cicindela sexguttata and Cicindelidia rufiventris on the rocky-clay path through dry oak-juniper woodland; however, sweeping the herbaceous glade vegetation produced nothing. The area has been quite dry, but I believe the low trap numbers are also due to the lack of vegetational diversity here due to the lack of any management via prescribed burns.

Cicindelidia rufiventris (eastern red-bellied tiger beetle) on rocky-clay path through dry oak-juniper woodland.

Blackjack Knob

There is lots of Sideroxylon lanuginosum (gum bumelia) at this Mark Twain National Forest site along Hwy 160 in Taney Co., and last time I checked the jug traps here there was already Plinthocoelium suaveolens in them. As a result, I expected to see lots of them this time, and such was the case. The ethanol/red wine trap had nearly three dozen P. suaveolens along with Stenelytrana emarginata, Eburia quadrigeminata, a few elaphidiines, Cotinis nitida, Euphoria fulgida, E. sepulchralis, and elaterids m. The ethanol-only trap had far fewer P. suaveolens (only 2) but similar numbers and diversity of other beetles. The area looks quite dry, but I swept the herbaceous glade vegetation anyway in hopes that I would find more Agrilus impexus. I did not, but I did collect about a dozen Taphrocerushowardi” (the quotation marks are a story for another day), which was surprising to me given how crispy brown the vegetation looked.

View from Blackjack Knob.

Bald Hill Glade Natural Area

This was my best locality last time in terms of trap numbers (at least the ethanol/red wine trap, not so much the ethanol-only trap). This is also one of two localities where I have a Lindgren funnel trap, which had a nice diversity of cerambycids (Neoclytus scutellatus, Eburia quadrigeminata, Aegomorphus modestus, Xylotrechus colonus, an elaphidiine, and an undet. cerambycid), scarabs (Cotinis nitida, Anomala sp.), elaterids, (scolytines (C. mutilatus), and other miscellaneous beetles. The area looked quite dry—both in the glade proper and in the surrounding dry upland deciduous forest, but trap numbers (again, only in the former) were incredible. In fact, there were so many beetles in the ethanol/red wine trap that I skipped sorting them in the field and brought the whole unsorted catch back to the car to sort out later. Part of this decision was due to the dark line of clouds I noticed to the north and that seemed to be moving closer. I hustled to the ethanol-only trap and quickly sorted its much sparser contents (only three Plinthocoelium suaveolens and a few E. quadrigeminata, E. fulgida, and elaterids), all the time keeping an eye on the approaching front. As I started the half-mile hike back, the winds really started picking up and the temps began to drop, and it became clear it would be a race to reach the car before the skies opened up. There isn’t much that phases me when I’m out in the field, but bushwhacking a half-mile stretch of abandoned, overgrown 2-track under dark, ominous clouds and with increasingly gusty winds had me feeling a bit nervous. I almost made it to the car before the rain started but did get wet in the final stretch. Still, I was able to get out of the area and back onto asphalt before the real deluge started. All traps were re-baited with the same bait, and a pheromone lure (Fuscumol Lure MR, #P655-MR, Chemtica International) was added to the Lindgren funnel. Sweeping herbaceous glade vegetation leading to the ethanol-red wine trap produced only a single Exema sp.

Threatening clouds.

That evening in the hotel room, I sorted the contents of the ethanol/red wine trap and recovered a nice diversity of cerambycids (Plinthocoelium suaveolens, Stenelytrana emarginata, Enaphalodes atomarius, Eburia quadrigeminata, Neoclytus scutellatus, Lepturges confluens, and elaphidiines), one Chrysobothris sp., numerous cetoniine scarabs (Cotinis nitida, Euphoria fulgida, E. herbacea, E. sepulchralis), elaterids, and Cnestus mutilatus, a mordellid, and two miscellaneous beetles.

Sorted contents of an ethanol-red wine-baited “jug trap.”
Euphoria fulgida (emerald euphoria) in ethanol/red wine-baited jug trap in xeric dolomite prairie remnant. The pink morph (right) and pink/green morph (center) were the only ones out of many normal green morphs (left) trapped.

Peck Ranch Conservation Area

I wasn’t optimistic about the jug traps here, since it was rather unremarkable dry-mesic upland deciduous forest—a last minute replacement locality for a dolomite glade that I could not access due to a closed gate. The area got a good rain last night and looked lush anyway, and as it turned out there were huge numbers of beetles in the ethanol/red wine trap. These included one Plinthocoelium suaveolens—unexpected, since I didn’t think there was any gum bumelia in the area, a nice series of Stenelytrana emarginata, lots of Eburia quadrigeminata, Cotinis nitida, and Euphoria fulgida, single specimens of Neoclytus scutellatus, Strangalia luteicornis, and Batyle suturalis, Euphoria sepulchralis, Trigonopeltastes delta, and Chrysobothris sp., and a handful of elaterids, Cnestus mutilatus, and other miscellaneous beetles. As at most previous spots, the ethanol-only trap had fewer numbers and lower diversity of beetles, although this did include one Lepturges sp. not caught yet in ethanol/red wine. Given the success with sweeping at previous spots (Agrilus impexus as two locations and Taphrocerushowardi” at the last one), I decided sweep the herbaceous roadside vegetation in this dry mesic upland deciduous forest and got singletons of Taphrocerus nicolayi, Acmaeodera pulchella, and Exema sp.—the Taphrocerus definitely making the effort worthwhile.

Helenium amarum (bitterweed) along roadside in dry mesic upland deciduous forest.

Stegall Mountain Natural Area

I was not a fan of this spot when I set the traps in mid-May given the fact that the spot and a large surrounding area had been recently burned. Despite that, I did get Purpuricenus in one of the traps last time, so I was more optimistic about it this time. As with the previous spot the area looked lush and got good rain yesterday, and as with the previous spot there were huge numbers of beetles in the ethanol/red wine trap, the most exciting of which were Stenelytrana emarginata, Purpuricenus humeralis, and Knullianaspinifera” (I think this may actually be an undescribed species). Other longhorns in the trap included lots of Eburia quadrigeminata, Neoclytus scutellatus (may include a few N. mucronatus), and elaterids and small numbers of Batyle suturalis, Enaphalodes atomarius, Lepturges sp., Chrysobothris/Actenodes, Euphoria fulgida, E. sepulchralis, Cnestus mutilatus and other miscellaneous beetles. I was especially happy to see Purpuricenus once again—the first of this trip. Again, the ethanol-only trap had fewer numbers and lower diversity of beetles, but this did include a single specimen of Phaenops aeneola—only the second known Missouri specimen (I collected the first many years ago at this very location)! Sweeping herbaceous vegetation in the xeric rhyolite prairie around Trap A and dry oak/pine woodland around Trap B produced nothing, but doing so along the roadside in the oak/pibe woodland produced Microrhopala vittata and Exema sp.

Liatris hirsuta (hairy blazingstar) in post-burn understory regrowth of dry post oak woodland.

Russell Mountain, Jakk’s Glade

This area also looked lush and got more good rains last night. The ethanol/red wine trap was down last time (not vandalized, I just failed to tie the knot on the carabiner securely), so I was anxious to see what it might produce. Like the previous spots on this side of the state, there were huge numbers of beetles in it, including Stenelytrana emarginata, Purpuricenus humeralis, Eburia quadrigeminata, Knullianaspinifera”, Neoclytus scutellatus (may include a few N. mucronatus), Enaphalodes atomarius, Strangalia luteicornis, undetermined elaphidiines, Chrysobothris sp., Cotinis nitida, Euphoria fulgida, E. sepulchralis, and undetermined elaterids (including many very tiny ones that I didn’t collect). The large series of Purpuricenus was unexpected and quite nice and means I’ve now collected species in this genus in all three of the igneous glades where I placed traps (but none in any of the dolomite glades—the opposite of what I expected given the number of individuals of this genus I’ve bait-trapped over the years at dolomite glades in Jefferson Co.). I was also surprised and happy to see the Knulliana, which I think is actually an undescribed species—they looked very similar coming out of the trap as the much more abundant Eburia, so I’ll have to check the collected specimens (both from here and other locations) more closely to see if there are other specimens that I’ve counted as Eburia. As expected, the ethanol-only trap again had far fewer numbers and lower diversity of the same species plus a single Lepturges sp. (same as the previous location). Both traps were re-baited with the same bait, though I started to doubt the value of continuing the bait comparison when ethanol/red wine already seemed far superior to ethanol-only. In the end, I decided to continue the continue the comparisons to allow for the possibility of other species later in the season that might show a different preference (and to allow publication of the study data even if this is not the case). Disappointingly, sweeping herbaceous vegetation in the xeric igneous prairie around both traps produced nothing.

Trimerotropis saxatilis (lichen grasshopper) on lichen-covered rhyolite exposures in xeric igneous prairie.

Hughes Mountain Natural Area

Again, like the previous spots today, the area looks lush and got more rain last night, but the numbers of beetles in the ethanol-red wine trap was not quite as overwhelming as in other locations. Nevertheless, there still a lot and two very good species : Sphenostethus taslei and Purpuricenus paraxillaris! I’ve only seen the former once before in a bait trap at Victoria Glades, and I’m always happy to see more individuals of the latter (the first new species I ever discovered!). Other beetles in the trap were Eburia quadrigeminata, an undetermined elaphidiine, Strangalia luteicornis, Typocerus velutinus, Xylotrechus colonus, Cotinis nitida, Euphoria fulgida, and various elaterids (including many tiny ones that I didn’t collect). In what is becoming a familiar refrain, the ethanol-only trap again had lower numbers and diversity, with Cnestus mutilatus being the only species not also caught in the ethanol/red wine-baited trap. Sweeping herbaceous vegetation, both in xeric igneous prairie around Trap A and dry post oak woodland around Trap B, produced nothing. This was the last location I was able to visit this day because of the unexpectedly large amount of time it took to sort the large numbers of beetles in many of the traps, so I saved the last two locations for the next day.

Sunset at Hughes Mountain.

St. Joe State Park

Like other areas on this side of the state, the vegetation was lush and got even more rain two nights ago. I’d been very anxious to see this spot, as there is lots of Sideroxylon lanuginosum (gum bumelia) in the small xeric dolomite glades remnants and extensive surrounding areas of restored dry post oak woodland, which promise to yield Plinthocoelium suaveolens. Unfortunately, the ethanol/red wine trap was completely missing—not fallen or pulled down by raccoons, but more likely stolen by humans (I probably should have placed this trap further away from the nearby paved bicycle trail). This is first case of trap theft I’ve experienced in a long time. If they had to steal one of the traps, I wish it would have been the ethanol-only trap rather than the ethanol/red wine trap—the former haven’t produced nearly as well as the latter, thus the loss of data would have been less. The ethanol-only trap had only a few beetles representing E. quadrigeminata, Batyle suturalis, Dichelonyx sp., Elateridae, and Cnestus mutilatus. I did not replace the ethanol/red wine trap, but instead re-baited the ethanol-only trap with ethanol/red wine. This means I will need to drop this location from the bait comparison analysis, but I wanted to give myself the best chance for trapping Plinthocoelium, and ethanol/red wine is clearly the better bait for accomplishing this. Sweeping herbaceous vegetation in the xeric dolomite prairie near Trap A and dry post oak woodland around Trap B produced nothing, but I did see an impressive Mydas tibialis (golden legged mydas fly) visiting flowers of Eryngium yuccifolium (rattlesnake master) in the dry post oak woodland.

Mydas tibialis (golden legged mydas fly) visiting flowers of Eryngium yuccifolium (rattlesnake master) in dry post oak woodland

Salt Lick Point Land & Water Preserve

Pflasterer’s Glade at Salt Lick Point Land & Water Preserve.

This is the only locality in Illinois that I placed traps—specifically because last fall I saw good numbers of Sideroxylon lanuginosum (gum bumelia), host for Plinthocoelium suaveolens (bumelia borer), and one in particular that showed signs of active infestation. Plinthocoelium has not been recorded from Illinois, so a voucher specimen will be important for publishing the record. The area apparently had gotten decent rains to this point, with vegetation in the woodlands looking lush and that in more exposed situations showing a tinge of wilt. Anticipation turned to disappointment when I looked inside the ethanol/red wine trap (placed right next to the infested bumelia tree) and saw lots of Eburia quadrigeminata and elaphidiines but only one Neoclytus acuminatus and no Plinthocoelium or other cerambycids. The infested teee was again churning out frass at the base of the trunk, so the infestation remains active—I will consider bringing screen with me next time so I can place a skirt around the base of the tree and check periodically for the adult once it emerges. In addition to the cerambycids noted above, the ethanol/red wine had Cotinis nitida, Enoclerus sp., Elateridae, Cnestus mutilatus, and other miscellaneous beetles. The ethanol-only trap had lower numbers of beetles, with Neoclytus scutellatus and Lepturges sp. being the only longhorns not represented in the ethanol/red wine trap. Interestingly, the trap also contained a few Buprestidae, which have been scarce in traps to this point (Actenodes sp.—prob. A. acornis or A. simi, Anthaxia sp.—prob. A. cyanella or A. dichroa, and Agrilus sp. Both traps were re-baited with ethanol-red wine to increase the likelihood of picking up P. suaveolens during the remainder of this season, and I will accordingly drop this locality from the bait comparison analysis (along with St. Joe State Park due to the ethanol-red wine trap being stolen) as well, thus limiting the analysis to the remaining 10 locations in Missouri.

Freshly-ejected frass at the base of a small living Sideroxylon lanuginosum (gum bumelia) tree indicates active larval infestation by Plinthocoelium suaveolens (bumelia borer)—a species not yet formally recorded from Illinois.

Sweeping herbaceous vegetation in the dry hilltop prairie near the traps produced nothing, but I did find a dead Lucanus capreolus (reddish brown stag beetle) dead on the trail though the dry-mesic upland deciduous forest leading up the them.

Lucanus capreolus (reddish brown stag beetle) female. Found dead on trail through dry upland deciduous forest.
Silphium integrifolium (prairie rosinweed) in dry upland deciduous forest.

©️ Ted C. MacRae 2022

2021 Texas/New Mexico/Arizona Insect Collecting Trip iReport

This is the 10th “Collecting Trip iReport”; this one covering 13 days of collecting in western Texas, southern New Mexico, and southeastern Arizona July 19–31, 2021. This trip was a “two-parter”—the first week with frequent field mate Jeff Huether (our seventh joint collecting trip) as we made our way from western Texas through southern New Mexico and into southeastern Arizona on our way to a memorial celebration for Jim Wappes at the home of Steve Lingafelter and Norm Woodley (for which I took the day off from collecting), and the second week visiting various locations in the “Sky Islands” of southeastern Arizona with several other entomologists.

As with all previous “iReports” in this series, this report is illustrated exclusively with iPhone photographs (thus the term “iReport”). Previous iReports in this series include:
2013 Oklahoma
2013 Great Basin
2014 Great Plains
2015 Texas
2018 New Mexico/Texas
2018 Arizona
2019 Arkansas/Oklahoma
2019 Arizona/California
2021 West Texas


Day 1 – Monahans State Park, Texas
This was my first stop on the previous trip back in late April and early May, and what a difference a couple of months with good rains makes—dry as a bone then but bursting with a great variety of wildflowers now. Like last time we stopped at the Shin Oak Picnic Area first, and almost immediately Jeff got an Acmaeodera gibbula on living Prosopis glandulosa (mesquite). I started beating the trees and got a good series of that species along with a good series of an Actenodes sp. (prob. A. mendax). There were also some mesquites that had been killed (apparently by herbicide), and when I started beating them I got several more A. gibbula and one Paratyndaris sp. It went from blazing hot when we arrived to raining about an hour later, and for a while after the rain moved through it stayed cloudy and quite comfortable. Eventually we decided to look for another spot with more mesquite to beat.

Monahans Sandhills State Park with rain moving in!
Poecilanthrax effrenus (family Bombyliidae) on flower of Thelosperma megapotamicum (rayless greenthread).
Acanthochalcis nigricans (family Chalicididae) female looking to oviposit on dead Prosopis glandulosa (mesquite).
Palafoxia sphacelata (othake).
Palafoxia sphacelata (othake).
Asclepias arenaria (sand milkweed).
Asclepias arenaria (sand milkweed).
Penstemon ambiguus (gilia beardtongue).

We found an area closer to the entrance (Equestrian Area) with lots of mesquite and also sunflowers, which Jeff was interested in looking at to search for meloids (blister beetles). Before I even reached the first mesquite I saw an Acmaeodera sp. (maybe A. obtusa) sitting on the flower of Thelosperma megapotamicum (rayless greenthread) and later found another plus one A. immaculata. Off the mesquite I beat just one Actenodes sp. prob. mendax and a few treehoppers, while another and a Chrysobothris sp. got away (it was by now quite hot and they bolted!). I continued beating mesquite but just wasn’t seeing anything, so we decided to take a look at the area around the main dunes and another picnic area.

Helianthus petiolaris (prairie sunflower).

At the Pump Jack Picnic Area, we saw a lot of Thelosperma megapotamicum (rayless greenthread) in bloom and decided to check the flowers for Acmaeodera. We each got a nice series of what appear to be A. obtusa and A. immaculata. Also, I finally found a single Acmaeodera immaculata on the flower of Hymenoppapus flavescens (collegeflower), which I’d been looking at all day thinking it must be a good Acmaeodera flower host. On the way back to the vehicle, I scared up a cicada that had been singing on a nearby plant—I’d been hearing them all day but assumed they were grasshoppers or katydids. I listened for another and saw it perched on the stem of Mentzelia nuda (bractless blazingstar), and netted it—a fine male, smallish and with a very white venter. I wanted to find one more—catching another A. gibbula in flight, and saw one singing in a mesquite tree. This time I took some photos of it (working carefully not to alarm it) and then hand-caught it (later identified as Diceroprocta texana). A nice end to the visit.

Male Diceroprocta texana singing on Prosopis glandulosa (mesquite).
Mentzelia nuda (bractless blazingstar).
Mentzelia nuda (bractless blazingstar).

After dinner, we returned to Shin Oak Picnic Area for night collecting. I setup my new Mercury-vapor (MV) station (first time using the gas-powered generator and tripod—slick setup) and my two ultraviolet (UV) stations. Jeff set out three prionic acid lures. Large numbers of Polyphylla monahanensis (which came mostly to the UV stations) and P. pottsorum (which came mostly to the MV station), but otherwise few beetles showed up and not a single longhorn. Other insects were also limited mostly to large numbers of ground-nesting bees and several big grasshoppers. No Prionus came to the lures—not surprising since we are in the tail end of the season for P. arenarius (April to July) and too early for P. spinipennis (mostly August). I’ve only gotten a few P. monahanensis and P. pottsorum before now, so it’s nice to have good series of each, but I would have preferred to collect some longhorns.

Blacklights humming as dusk settles over the dunes.
Polyphylla pottsorum at the MV sheet.

Day 2 – Toyahvale, Texas
We’re on our way to the Davis Mountains, and along the way I decided to stop at the “Agrilus cochisei” spot we found (on a tip from Jason) back during the April trip. I swept the roadsides—not just the host plant (Artemisia occidentalis, western ragweed) but a variety of other plants in bloom but did not find any A. cochisei. I did collect a few meloids (which were on Solanum eleagnifolium) and Zygogramma leaf beetles but nothing else.

Proboscidea parviflora (family Martyniaceae)—doubleclaw or red devil’s-claw.

A bit further down the road from the last stop, we noticed this memorial to the many horses that have been transported along this highway on their way to slaughter.

Davis Mountains, Ft. Davis, Texas
Another roadside stop for one of the places where I collected during last April’s trip. I was hoping we would not be too late for Acmaeodera—there were plenty of plants in bloom but we did not see any. Senegalia greggii (formerly Acacia greggii, commonly called cat-claw acacia) and Vachellia constricta (formerly Acacia constricta, commonly called whitethorn acacia) were both in bloom, and off the former I got the obligatory Stenaspis solitaria male/female pair as well as a Lampetis drummondii, but I collected nothing off the latter. Tried for a couple of cicadas and missed ‘em both!

Stenaspis solitaria male on flowering Senegalia greggii (cat-claw acacia).
Lampetis drummundii captured on flowering Senegalia greggii (cat-claw acacia).
Eurema nicippe (sleepy oranges) on yellow asteraceous flower.

Davis Mountains, Boy Scout Rd, Texas
This looked like a good spot, with water in the creek and lots of butterflies flying around. Beating, however, yielded nothing but lots of lep larvae. It seems we are in the mid-summer lull—too late for spring things, but too early for late summer-fall species. I think we’ll try some higher elevations and see what it is like.

Argemone aenea (golden prickly poppy).
Argemone aenea (golden prickly poppy).
Argemone albiflora (white prickly poppy).
Argemone albiflora (white prickly poppy).

We were headed back towards the highway when I spotted a stand of Thelosperma megapotamicum along the road. We got out so I could take a look at them and immediately encountered this Texas horned lizard who seems to be saying “WTF?!” I swept through the Thelosperma and picked up two Batyle sp., one Enocleris sp., and a couple of species of meloids. There were lots of other plants in bloom, too, including several that are typically attractive to beetles such as Sphaeralcea and Ratibida. However, nothing was seen on them, further reinforcing our desire to go to higher elevations to see if that would improve the collecting.

Phrynosoma cornutum (Texas horned lizard).

Davis Mountains, Madera Canyon, Texas
We wanted to get up to higher elevation to see if that might improve the insect collecting. It is strange—the Davis Mountains are greener than I’ve ever seen them, yet there are almost no insects, no flowers. Jeff and I were wondering if the deep freeze Texas experienced this past winter might have knocked out insect populations. We beat along the way but just we’re not seeing anything on the beating sheets. We hiked our way up to the overlook, and up there I ran into a few species of tenebrionids running along the trail and doing their famous “headstands” when we disturbed them. On the way back down I saw a few large, red and black clytrine chrysomelids on what I take to be a fall-flowering helianthoid aster (old flowering stalks were 5–6 feet tall), so I picked up a few for Shawn (my scope of insects that I’ll collect expands greatly when I’m not finding anything in the groups that I study). I think we’ve had it with the Davis Mountains, and tomorrow we’ll travel further west and try our luck around Fabens.

Lower part of the Madera Canyon Trail.
The trail at upper elevations.
A small Eleodes sp. doing the headstand.
The larger Eleodes carbonaria doing the headstand.
Same individual as in the previous photo.

Day 3 – Van Horn, Texas
Just a quick stop along the highway when we saw a variety of plants in bloom. Things are different at these lowers elevations compared to the Davis Mountains. Sweeping yielded a number of Agrilus sp. (vittate) and two Agaeocera gentilis—I suspect they were on the Sphaeralcea angustifolia (narrow-leaved globemallow), along with an Acmaeodera sp. and assorted other beetles. On a much taller globemallow I found two Tylosis jiminezi (male/female) perched on the foliage—a first for me! I did a little more sweeping further to the south but came up with only a few melyrids and a blister beetle (Epicauta segmenta). Nice stop!

Tylosis jiminezi on Sphaeralcea angustifolia (narrowleaf globemallow).
Sphaeralcea angustifolia (narrowleaf globemallow).
Sphaeralcea angustifolia (narrowleaf globemallow).

San Felipe Park, El Paso Co., Texas
When we first arrived, we were not at all optimistic—it looked like it hadn’t rained in years. However, the Larrea tridentata (creosote) and Prosopis glandulosa (mesquite) were both in full bloom, so we began looking about. Almost immediately I saw Stenaspis solitaria in the mesquites, which in itself is not exciting, but two of the first three I saw were the form that has the distinctly reddish-brown pronotum, which I’ve never seen before. I saw a couple more and tried to get photographs, but they were too skittish. As I searched for them, I caught one Aethecerinus latecinctus (a second got away), one Plionoma sp. (not sure if it’s suturalis or rubens), one Chrysobothris sp. (prob. C. octocola) and one Acmaeodera gibbula. The real fun began, though, when I walked by a creosote and saw a Gyascutus planicosta (should be subsp. obliteratus in this area) take flight. I tracked it to see where it landed, caught it, and then put most of my effort into getting a decent series of individuals. I succeeded, but it took more than four hours with the heat maxing out at 96°F! In addition to the Gyascutus, Jeff was quite excited to see the bright green and orange blister beetle Eupompha fissiceps abundant on the creosote in mating pairs and feeding on the petals of the flowers. I finished off the blister beetle fun by finding Cysteodemus wislizeni (black bladder-bodied meloid) crawling in the sand, which, despite the common name, was decidedly bluish.

The author looks out over the vast creosote/mesquite scrub.
Eupompha fissiceps on Larrea tridentata (creosote).
Eupompha fissiceps feeds on Larrea tridentata (creosote) as “Mr. Meloid” (Jeff Huether) looks on.

After going into town to restock on supplies and catch some dinner, we returned to the park to do some lighting and night collecting. The moon is almost full, which generally puts the kibosh on longhorns coming to the lights, but we decided to try anyway because of the high amount of activity during the day in a rarely-visited location. I set up the Mercury-vapor (MV) light only and skipped going through the trouble to put up the ultraviolet lights also. I also wanted to beat the mesquite since I didn’t have much chance during the day, spending most of my time hunting Gyascutus with an aerial net, and once I got the MV setup going I started whacking the mesquite. Almost immediately, I got three Aethecerinus latecinctus, which came off the first two plants I beat. This motivated me further and caused me to commit to beating for the next hour or so—never even getting a single beetle of any kind! By then the lights had been going for awhile, and I was pleased to see several Derobrachus hovorei (palo verde root borer) crawling on the sand near the light. Even though it is a common species, I’ve not seen many myself, so I was happy to have a nice series to take up beaucoup room in one of my prionid drawers. Otherwise, very few beetles came to the lights, or most other insects as well—the sheet being covered primarily by dozens of white-lined Sphinx moths and lots of wasp/bee-type things. We did enjoy the evening, however, as we sat in our chairs and drank a cold brew between checkings of the light.

Mercury-vapor light setup at dusk.
Mercury-vapor light just turned on at sunset.
Mercury-vapor light going strong as darkened settles.
Derobrachus hovorei (palo verde root borer).

Day 4 – Chaparral, New Mexico
We had planned to take NM-213 north to White Sands, but public access was blocked at Ft. Bliss. While backtracking, I spotted Sphaeralcea angustifolia (narrow-leaved globemallow) growing along the roadside and stopped to check it out. There was nothing on any of the plants, despite good growth and appearing to be coming into flowering. I did a little beating on Prosopis glandulosa (mesquite) as well—again very little going on, just a few treehoppers, tiny blister beetles, and one weevil. Larrea tridentata (creosote) was in full bloom, just like at Fabens, and here too there were many Eupompha fissiceps on the flowers and in mating pairs. I couldn’t resist collecting just a few and even made a short video of a mating pair engaged in some interesting behavior. I did see one Gyascutus planicosta as it flew by, but I could not track it to see where it landed. Otherwise all I picked up was another Cysteodemus wislizeni (black bladder-bodied meloid)—along with a photo, and a male cicada (maybe Diceroprocta texana) singing on a dry yucca stalk (hand-collected!).

Cysteodemus wislizeni (black bladder-bodied meloid).
Eupompha fissiceps mating pair engaged in some interesting behavior.

Point of Sands, New Mexico
Jeff wanted to stop here to look for Pleurospasta mirabilis, a really cool-looking blister beetle that looks unlike anything else. He found just one by disturbing the host plant (small purple blooms), and I found none—seems we are right at the tail end of their activity period. I looked around for other things also, but there was not much out. I did catch a couple more cicadas (males singing), and near an Ephedra sp. bush I found a mostly-intact carcass of Sphaerobothris ulkei. The most interesting find, however, was a couple of apparently lost pitfall traps—the barriers had fallen over, and the cops were filled with sand and the carcasses of numerous tenebrionids that had fallen into the traps and never been retrieved. I pulled up the cups and filled in the holes to prevent further loss, finding a few live tenebrionids and trogids and one Pasimachus sp. ground beetle that had not yet succumbed in the process.

Point of Sands, New Mexico.
“Lost” pitfall trap.

Hatch, New Mexico
While passing through the town of Hatch, I couldn’t resist taking the opportunity to stop and photograph some interesting town characters.

An interesting take on the Sinclair dinosaur.
Chili Pepper Man!
Cast of characters.

Deming, New Mexico
Another roadside stop in an area that was green with flowers and also had Yucca (to look for Tragidion) and Ephedra (to look for Sphaerobothris). Neither of those insects were found, and no buprestids or cerambycids were seen on or swept from any of the many composite flowers about including Thelosperma megapotamicum. I did find a couple of the meloid Lytta biguttata, one on flowers of Cirsium sp. and another on an unidentified yellow composite flower, and Jeff found a huge aggregation of another meloid, Epicauta costata, on herbicide-treated Kali tragus (prickly Russian thistle). I finished off the stop by finding a spectacular ridged tenebrionid beetle walking about after the sinking sun went behind some clouds.

Lytta biguttata on yellow composite flower.
Epicauta costata on herbicide-treated Kali tragus (prickly Russian thistle).
Epicauta costata on herbicide-treated Kali tragus (prickly Russian thistle).

Day 5 – Sunshine, New Mexico
We saw a nice stand of what proved to be Picradeniopsis absinthifolia (formerly Bahia absinthifolia, hairyseed bahia) and stopped to check them for buprestids/meloids. None were seen, just a few bees and lots of bee flies (genus Geron?). We did find a fair number of Cysteodemus wislizeni (black bladder-bodied meloid) crawling on the roadsides in a patch of Solanum elaeagnifolium (silverleaf nightshade).

Geron? sp. (family Bombyliidae) on flower of Picradeniopsis absinthifolia (hairyseed bahia).
Geron? sp. (family Bombyliidae) on flower of Picradeniopsis absinthifolia (hairyseed bahia).

Columbus, New Mexico
We found a moist drainage along the roadside with plants blooming in abundance, including Isocoma tenuisecta (burroweed) and Sphaeralcea angustifolia (narrow-leaved globemallow); however, insects were very scarce. I didn’t see any buprestids or cerambycids at all on any of the plants, only picking up one Lytta biguttata and one Cotinis mutabilis on the Isocoma and sweeping single examples of a small black/red Cleridae from the latter and a yellow-flowered composite.

Pepsis thisbe (Thisbe’s tarantula-hawk wasp) on flowers of Isocoma tenuisecta (burroweed).

Animas, New Mexico
We drove a fair distance west hoping to get into a different rainfall system in hope that insects would be present and saw a roadside in good bloom with the surrounding creosote scrub also green and blooming. As soon as we got out of the car we saw big beetles flying overhead and tracked them back to several creosote bushes very near the car with an aggregation of yet another blister beetle, Pyrota postica, which were mating and feeding on the flowers and leaves. After taking a few photos (and collecting my small series), I started sweeping through the variety of plants in bloom along the roadsides. I did not see anything on the flowers themselves (including Baileya multiradiata, pretty good buprestid flower) but collected a series of clytrine chrysomelids and one Dectes sp. While sweeping through Sphaeralcea hastulata (spear globemallow), I got one Agaeocera gentilis, and sweeping though a mix of S. hastulata and S. angustifolia (narrow-leaved globemallow) I got three Agrilus sp. (perhaps the same as I collected south of Van Horn, Texas). Finally, on the latter, I found one more A. gentilis perched on the leaf.

Pyrota postica female on flowering Larrea tridentata (creosote).
Pyrota postica female feeding on the flowers of Larrea tridentata (creosote).
Sphaeralcea hastulata (spear globemallow).

Portal, Arizona
Finally made it to Arizona, and for the first stop I wanted to try a spot below Portal where I’ve had limited success finding Sphaerobothris ulkei on Ephedra. Last time I was I here I found a few, but not until after being distracted by Gyascutus caelatus and Hippomelas sphenicus in the acacias and mesquites. I vowed not to let that happen this time and weaved a zigzag pattern looking at every Ephedra I could find. While I was doing that. I did see one G. caelatus take flight and then caught another that I saw sitting on Vachellia constricta (white-thorn acacia) but remained focused on looking at the Ephedra. Eventually, after not seeing any S. ulkei, I started looking for other buprestids. The acacias were just beginning to flower—only a few plants had open flowers, and Jeff noticed the tiny silhouette of an insect in flight approaching the flowers. He netted it and showed it to me, and to my surprise it was one of the species in the A. stigmata group (black with two red apical spots)—none of which I have ever collected before but which I believe could be A. davidsoni. We spent the next hour watching for and netting the silhouettes on the few trees we could find with flowers, and then I got the beating sheet out and beat more off of the trees (whether in flower or not) to end up with a nice little series. I’ll be anxious to confirm whether these are A. davidsoni.

Mozena arizonensis on Vachellia constricta (white-thorn acacia).
Vachellia constricta (white-thorn acacia).

South of Willcox, Arizona
We got to Willcox around dusk, and the cool, breezy conditions told my gut that it would be pointless to set up the lights. Still, I couldn’t let myself not try, and without much opportunity to look for a good place to set up I just went down Blu Sky Rd to E Moonlight Rd and hoped for the best. My optimism waned rapidly, as conditions continued getting colder and breezier, and not a single insect came to the light—I should’ve listened to my gut!

Moonrise. A twilight zone moment occurred when I realized that I had taken this photo on Moonlight Rd!😮

Day 6 – Jim Wappes Celebration, Hereford, Arizona
Fun day with lots of fellow coleopterists at the home of Steven Lingafelter and Norm Woodley in memory of Jim Wappes. We got there in the early afternoon and enjoyed eats, conversation, and war stories from the field.

Min, Sangmi Lee, Lisa Lee, Candy Kuckartz, the author, Margarethe Brummermann, Jason Botz, & Andrew Johnston (photo by Steven Lingafelter).
The author, Andrew Johnston, Norm Woodley, & Ed Riley (photo by Margarethe Brummermann).

It was a good day for a party, as rain made insect collecting a no-go. As dusk settled, I admired the incredible view from Steve’s and Norm’s back patio!

Low clouds hang over the Huachuca Mountains.

The fun extended well into the evening hours. My thanks to Steve and Norm for hosting the celebration—what fun to see and talk to so many entomologists in one place.

Steve Lingafelter, Gino Nearns, “Kira Brummermann,” & the author (photo by Margarethe Brummermann).
Bill Warner, Norm Woodley, Paul Skelley, Jeff Huether, Steve Lingafelter, Andrew Johnston, & the author (photo by Margarethe Brummermann)

Day 7 – Superstition Mountains, Weaver’s Needle Vista Viewpoint, Arizona
We awoke to rain yesterday morning in Willcox, and it has stayed with us since—first on our way to Hereford for the Wappes Celebration, then up to Phoenix this morning—our efforts to escape the rain by coming north thus proving futile. I came to this spot on a tip that I might find Agrilus cavifrons on Celtis pallida (spiny hackberry) (although maybe a bit early), and I’d hoped despite the light rain I would still be able to find it. I did not—though I found the plants, but I did get a Chrysobothris sp. that I don’t recognize while beating Senegalia greggii (formerly Acacia greggii, cat-claw acacia). Of course, it was on the first plant that I beat, so I ended up beating for another hour with nothing to show for it! While walking the short paved trail, I found Cercidium sp. (palo verde) tree that had been cut up and showed evidence of buprestid infestation in one of the larger branches, so I retrieved and cut it up for rearing.

Water… water… everywhere!
Cylindropuntia fulgida (chain-fruit cholla, jumping cholla, hanging chain cholla).

Lost Dutchman State Park, Arizona
We made a quick stop here to admire its incredible scenery before heading back south.

Superstition Mountains as seen from Lost Dutchman State Park.
Stands of Cylindropuntia fulgida (chain-fruit cholla) frame this view of the Superstition Mountains from Lost Dutchman State Park.
Carnegiea gigantea (saguaro) in front of the Superstition Mountains.
Carnegiea gigantea (saguaro).

Upper Tanque Verde Falls Trailhead, Tucson, Arizona
The rain finally moved out and it was sunny for the drive back to Tucson, but with still-cool temps we weren’t sure if lighting would be worthwhile. We decided to try this spot—not too high (based on a tip by Bill Warner), and I setup the lights using a technique recommended by Roy Morris that involved placing one ultraviolet (UV) light on each side of the sheet, extending the Mercury-vapor lamp above to the top of the sheet, and periodically shutting off the latter to allow the UV lights the pull in the “shyer” insects before turning it back on. While I waited for the lights to started pulling things in, I did some beating around the area. I only collected one specimen, but it was a Cleridae that I don’t recognize, which was beaten off of Prosopis glandulosa (mesquite). Temps ended up in the lower 70s to upper 60s, but at the lights we still got a few longhorned beetles (Aneflomorpha sp.), a few melolonthine scarabs, and a nice series of two species of Pachybrachis (that I will send to Bob Barney).

Jeff looks for beetles at the light sheet.
Lethocerus medius at ultraviolet/mercury-vapor light in mesquite/acacia scrub.

Day 8 – Atascosa Mountains, Ruby Rd near Atascosa Lookout Trailhead, Arizona
The primary quarry here was Acmaeodera chuckbellamyi, a species I described in 2014 from a single specimen collected by my friend and hymenopterist, Mike Arduser, at this location on flowers of Aloysia sp. Several people have tried to find it since—without success, and in June 2011 the area was severely burned by the 27,550-hectare Murphy Fire. I was hoping enough time has passed to allow the area (and beetle population) to recover in this, my first attempt, at finding the species myself. I knew it was a long-shot, and long sorry short I did not find either the beetle or it’s Aloysia host flowers. I did collect a number of other beetles, however, including Lycus lorises, a few longhorned beetles and pachybrachine leaf beetles, and one Acmaeodera quadrivittatoides on flowers of Eysenhardtia orthocarpa (desert kidneywood), several Aneflomorpha sp. and a few pachybrachines on Quercus oblongifolia (Mexican blue oak), and more pachybrachines on Senegalia greggii (catclaw acacia), Mimosa dysocarpa (velvetpod mimosa)—the latter also yielding a few tiny Chrysobothris spp. (one looking like C. lucanus), and Propopis glandulosa (mesquite). Jeff also collected and gave to me a couple of Acmaeodera parkeri on flowers of Talinum aurantiacum (orange flameflower). I hiked 0.7 mi E on Ruby Rd to a spot where I swept a few beetles from low vegetation, and on the underside of a large, fallen branch of Q. oblingifolium I found a large female Polycesta arizonica. Finally, about halfway back to the trailhead I encountered a few tiger beetles on the road near standing puddles of water from the recent rains.

View from Ruby Rd looking south toward Mexico.
View from Ruby Rd looking southwest toward Mexico.
Lycus loripes on flowers of Eysenhardtia orthocarpa (desert kidneywood).
Lycus loripes on flowers of Eysenhardtia orthocarpa (desert kidneywood).
Talinum aurantiacum (orange flameflower).

Atascosa Mountains, Peña Blanca Lake, Arizona
We came here to look for Deltaspis tumacacorii, which like many rare beetles the odds are against finding it despite it having been taken in the area on several occasions. Again, this would not be one of those occasions, but I was happy to find a tiny tiger beetle (Cylindera viridisticta arizonensis) along the creek and even happier to beat a series of two species of Paratyndaris, one large Lampetis webbii (only my fifth specimen). and one Aneflomorpha sp. from mostly dead Senegalia greggii (catclaw acacia).

Leptinotarsa lineata on Ambrosia monogyra (cheeseweed burrobrush)—adult.
Leptinotarsa lineata on Ambrosia monogyra (cheeseweed burrobrush)—larvae.

Huachuca Mountains, lower Carr Canyon, Arizona
After dropping Jeff off at his hotel in Tucson (he flies home in the morning), I high-tailed it down to Carr Canyon to do some light collecting. It has been a long time since I’ve seen my sheet covered so quickly and thoroughly with insects! My quarry was longhorned beetles—of which I got a nice variety, but who can resist also the variety of scarabs, ground beetles, tenebrionids, blister beetles, and even planthoppers that flock to the lights in the mountains of southeastern Arizona? I’ll have to control myself a little better in the coming nights! Walking about along the roadsides (hoping to see Amblycheila giant tiger beetles), I encountered a gorgeous male Aphonopelma chalcodes (Arizona blonde tarantula).

Mercury-vapor/ultraviolet light setup.
Enaphalodes cortiphagus at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Parabyrsopolis chihuahuae at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Phileurus truncatus (triceratops beetle) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Chrysina beyeri (Beyer’s jewel scarab) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Chrysina gloriosa (glorious jewel scarab) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Pachysphinx occidentalis (western poplar sphinx) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Manduca florestan (Florestan sphinx) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Aphonopelma chalcodes (Arizona blonde tarantula) on ground in in oak/juniper woodland.
Aphonopelma chalcodes (Arizona blonde tarantula) on ground in in oak/juniper woodland.
Aphonopelma chalcodes (Arizona blonde tarantula) on ground in in oak/juniper woodland.

Day 9 – Huachuca Mountains, Copper Canyon, Arizona
I met Steve Lingafelter and Norm Woodley at their house, and together we drove to the famed Copper Canyon on the south side of the Huachucas. On the way I got a nice primer about the species that have been collected there and the plants they have been collected on. Steve and I started out walking the trail up the canyon while Norm swept the area down below. I pretty much beat every oak along the way, for a while only getting a smattering of beetles—Agrilaxia sp. on Quercus emoryi (Emory oak) and also on Q. arizonica (Arizona white oak) along with Sternidius decorus?. About a half-mile up the trail I beat dead branches of Q. hypoleucoides (silverleaf oak) and, not seeing anything at first, said “I’m going back; I’m literally not getting anything.” Right then, a small black beetle on the sheet caught my eye. I looked at it closely and realized it was a Mastogenius (prob. M. robustus)! I popped it in the bottle and beat more dead branches from the same tree and got not only another Mastogenius but also Tigrinestola tigrina. Freshly motivated, I spent the next half-hour working all the oaks in the area—and, as often happens, did not see another beetle! Nevertheless, it was hard not to be happy with the beetles that I’d gotten.

An unidentified treehopper nymph, likely something in the subfamily Smiliinae, tribe Telamonini, on twig of Quercus arizonicus (Arizona white oak).

I walked back down the trail and met Norm, who was just starting up. He filled me in on the results from below, which included sweeping a few Agrilaxia hespenheidei—one of my target species—and an Agrilus sp. on Bouvardia ternifolia (firecrackerbush). I decided to work the slope under the road, reasoning that Norm had likely already worked the flat ground below. On the first B. ternifolia I approached, I saw an A. hespenheidei on the flower, gave the plant a sweep, and caught not only the A. hespenheidei but also an Agrilus sp. (maybe A. latifrons). Over the next hour I would sweep a nice series of A. hespenheidei from B. ternifolia (but not another Agrilus sp.). While I was doing that, I also swept the numerous stands of Acaciella angustissima (formerly Acacia angustissima, prairie acacia) looking for the large, spectacular Agrilus cavatus. I would find two, and considering that I swept perhaps 50 or more stands they were well earned. Also, in the meantime, I noticed Acmaeodera parkeri on small blue flowers that I eventually identified as Evolvulus arizonicus (Arizona blue-eyes). The flowers were few in number and the beetles difficult to catch, so I only ended up with two specimens. Further down the slope the flowers much more abundant, but there was not a beetle to be found on them. In the waning moments of my visit, I encountered two Trichodes peninsularis horni on flowers of Lasianthaea podocephala (San Pedro daisy).

Bouvardia ternifolia (firecrackerbush), flower host for Agrilaxia hespenheidei.
Acaciella angustissima (prairie acacia), host for Agrilus cavatus.
Evolvulus arizonicus (Arizona blue-eyes), flower host for Acmaeodera parkeri.
Lasianthaea podocephala, flower host for Trichodes peninsularis horni.

Eventually we all met up at the car, compared our catches (not surprisingly, Norm did very well with buprestids and Steve did very well with cerambycids), and I did okay in both counts. We headed back over Montezuma Pass and were greeted with stunning views looking down into the Coronado National Monument!

Coronado National Monument from near Montezuma Pass.

Huachuca Mountains, Miller Canyon, Arizona
After dropping Steve off, Norm and I went to nearby Miller Canyon to look at a spot where he has collected three species of Taphrocerus (I’ve only collected two, but only once way back in 1987 at a spot in the lower canyon). We thought it might be too late in the season, but it was at least worth a shot. The sedges were lush and green, but the “sedgy wedgies” were absent. Looks like I’ll have to just come back out in April or May!

After striking out with Taphrocerus, I went back down to the lower elevations of the canyon to look for Tragidion on the stand of Baccharis sarothroides (desert broom) that occurs there. Once again, I was likely too late to find them, but as with Taphrocerus it was certainly worth a shot, and again I would not find any despite looking and most of the large plants in the area. I did find a few Euphoria leucographa feeding on the sap flows and a very large red/black clytrine, so it wasn’t all for naught. Just another reason to come out earlier in the season.

Euphoria leucographa feeding on a sap flow on stem of Baccharis sarothroides (desert broom).

Day 10 – Santa Rita Mountains, upper Box Canyon, Arizona
After getting reports of buprestid activity near Madera Canyon, I decided to head to the Santa Rita Mountains today instead of continuing in the Huachucas. The shortest route to this most famous of canyons in the Santa Ritas goes down Box Canyon, a less-well-known but still-fantastic canyon in its own right and where I’ve had good luck collecting the two previous times I’ve been there (August 2018 w/ Art Evans, Steve Lingafelter, and Norm Woodley; and September 2019 w/ Jeff Huether). I stopped at the “dry falls” and worked my way back up the road to a point where I’ve collected the majority of my insects there. Along the way, I beat the flowering Eysenhardtia orthocarpa (desert kidneywood)—insects were not numerous on the plants, but over the course of the trips up and back I got Acmaeodera gibbula, A. cazieri, two Aneflomorpha sp., and a few Lycus sp. I also swept the just-beginning-to-flower Mimosa dysocarpa (velvetpod mimosa) but got just a single Sphaenothecus bivittatus. When I reached the top of the canyon, I looked for a small patch of Allionia incarnata (creeping four-o’clock) in the steep road bank, off the flowers of which I have previously collected Acmaeodera cazieri, A. parkeri, and A. yuccavora. I found the patch, but the plants were not yet in flower. What was in flower on the flats above the bank, however, was Talinum aurantiacum (orange flameflower), off which I collected all three species (A. cazieri being the most abundant and only a single A. cazieri). As at previous stops this week, it seems that insect activity is beginning but is still a bit shy of coming into full peak.

Larva of Euscirrhopterus gloveri (purslane moth), which was present in outbreak numbers on a portulacaceous plant growing amongst Talinum aurantiacum.

While I was collecting, a caravan of cars came by. They turned out to be filled with entomologists attending the Invertebrates in Education Conference, one of whom I knew—Tad Yankoski of the Sophia M. Sachs Butterfly House in my hometown of St. Louis. He handed me a vial containing a large, live individual of Polycesta aruensis. I was excited to see this, especially when he told me he found it and saw many more in the flats below Madera Canyon, where I had planned to go next!

Santa Rita Mountains, flats below Madera Canyon, Arizona
It was a frustrating afternoon on several fronts. Starting off, I had trouble finding the Polycesta aruensis locality, and when I finally did find it there was nary a Polycesta to be seen. Perhaps they sleep during the heat of the day.🤷 After that, there was little time to go anywhere but Madera Canyon, where I spent a half-hour beating Quercus oblongifolia (Mexican blue oak) hoping to see Chrysobothris chalcophoroides (I didn’t) and another two hours checking out Baccharis sarothroides (desert broom) in the area where I collected Stenaspis verticalis and Tragidion deceptus two years ago (also fruitless). Of course, the dreadfully common Stenaspis solitaria was everywhere, but all I ended up collecting was a tenebrionid on the 2-track, a clytrine beaten from Cercidium aculeata (retama), and one Euphoria leucographa along with a nondescript cerambycid (Heaperophanini maybe) on B. sarothroides. On the way back to the vehicle, I encountered a dead, mostly skeletonized deer, and while I rarely collect from carcasses, I noticed a little green beetle crawling on the jaw bone. I figured it must be the cosmopolitan clerid, Necrobia rufipes—something I’d not seen before, so I collected it and tried to collect but missed another one.

Rain clouds gather over Madera Canyon.
Another view of the mouth of Madera Canyon.
Magusa sp. (one of the narrow-wings) caterpillar feeding on Sarcomphalus obtusifolius (lotebush).

Santa Rita Mountains, lower Florida Canyon, Arizona
I suppose I can credit my frustrating afternoon for one of my best nights of lighting ever. I stumbled upon this spot at the bottom of Florida Canyon during this afternoon’s Polycesta wild-goose chase and immediately thought, “Wow, what a perfect spot to set up a light!” A nice place to pull off the road with a small, level clearing embedded within low-elevation oak woodland. There was even a babbling creek in the background! It was close to dark by the time I returned and set up the lights (would’ve been even later if I’d gone into town for a “real” dinner). Ironically, there were neither the diversity nor quantity of beetles as two nights ago in lower Carr Canyon. But the cerambycids brought it… and kept bringing it! It seemed like every time I got up to check the sheet there were another 4–5 individuals. I ended up leaving the sheet up for four full hours and collected perhaps 40–50 specimens representing a dozen or more species. A few I don’t recognize, and most of those that I do recognize have resided in my cabinet in precious few numbers until now. It’s been years literally since I’ve had a night like this, and it’s a nice shot of motivation leading into the last few days of what is starting to feel like a long trip.

Sometimes circumstances dictate an unusual dinner.
Enaphalodes niveitectus at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Tigrinestola tigrinus at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Coenopoeus palmeri at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Syssphinx hubbardi (Hubbard’s silk moth) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Eacles oslari (Oslar’s imperial moth) at Mercury-vapor/ultraviolet light in oak/juniper woodland.

Day 11 – Madera Canyon Rd, Continental, Arizona
I came back to the spot where Tad Yankoski had seen Polycesta aruensis so abundantly yesterday morning but which was completely absent by the time I got there in the afternoon. I did not see any adults on the trees this morning either and was about to give up when I spotted a few partially dead trees with very large, apparently fresh emergence holes in the main trunks that were the perfect size for P. aruensis—good thing I brought my chainsaw! I cut a trunk with its branches and segregated the cut up wood into age (fresh dead versus older) and size (twigs, medium branches, and main trunk) classes. While I was doing this, a Polycesta adult dropped off one of the fresh-dead, medium-sized branches! I beat the remaining branches on the tree and on nearby trees but did not see any more, so whatever Tad witnessed yesterday morning must have been an ephemeral event, perhaps related to synchronized emergence from the very trees among which I collected the wood. NOTE: don’t let anyone tell you that cutting up wood for rearing beetles is anything but a sweaty, exhausting endeavor, even with temps still in the mid-80s and decent cloud cover!

This camouflaged tanker marks the spot—trees on the W side of the tanker.
Job half-done—wood cut up.
Job complete—wood segregated and bundled.

Santa Rita Mountains, lower Florida Canyon, Arizona
I had such good night collecting here last night that I thought I’d come back and see what I could find during the day in this ribbon of riparian oak/hackberry woodland. Almost immediately I beat a Paratyndaris sp. off of dead Celtis reticulata (netleaf hackberry). The tree was very dead, but I knew Paratyndaris spp. like old, dead wood, so I split open some of the branches and found larvae inside and also a dead and unemerged but perfectly intact adult of a very tiny Chrysobothris sp. inside one of the smaller branches. Beating on other plants in the area was, in general, fruitless, but occasionally (and just often enough) I encountered something of interest that motivated me to continue working: Paratyndaris sp. and Agrilus sp. on Quercus oblongifolia (Arizona white oak), a small red/brown elaterid on Prosopis glandulosa (mesquite), and Acmaeoderopsis sp., Paratyndaris sp., and a few clytrines on Senegalia greggii (cat-claw acacia).

Lower Florida Canyon.
Celtis reticulata (netleaf hackberry).

Santa Rita Mountains, Gardner Canyon, Arizona
Last night, I got a message from fellow buprestophile Robert Velten, who told me that he would be in Gardner Canyon the next day, so after finishing at Florida Canyon I drove over to Gardner Canyon to meet up with him. Despite being longtime correspondents, Rob and I had never actually met face-to-face, so I was thrilled to have the chance to do so and spend some time with him in the field. Joining him were his mothing buddies Steve McElfresh and Paul Tuskes, and a little later our mutual friend and Arizona coleopterist-extraordinaire, Margarethe Brummermann, also joined us for a night of lighting. It was great to spend time at the lights with so many like-minded folks! There were three light stations between us, but the weather was less than cooperative—a persistent cool breeze accompanied constant lighting and thunder in the mountains above. Eventually, the threat was realized when the skies opened up, prompting a hasty dismantling and storage of all my lighting equipment safely inside the vehicle. Nevertheless, in the time that I was able to collect, I got a small number of longhorned beetles (half of which came to my light in the moments I was taking it down—longhorns typically become very active right before a storm) along with a variety of showy scarabs and clerids. The rain ended as quickly as it began, so the socializing continued. The entire evening I was continuously taunted, however, by a large prionid sitting inside its emergence hole on the trunk of a large Quercus emoryi (Emory oak). It only showed its jaws and antennae, and if I even touched the tree to boost myself up for a closer look it withdrew deep into the hole. I’m convinced it was Nothopleura madericus—a species I’ve never collected. I can still hear it laughing at me! My attempt to find one out and about by scanning the trunks and branches of of the other oaks in the area with my headlight was not successful, although I did collect another elaphidiine longhorn in such manner.

Polyphylla decemlineata (or perhaps a new species) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Lucanus mazama (cottonwood stag beetle) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Xyloryctes thestalus (western rhinoceros beetle) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Hemiphileurus illatus at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Cymatodera horni at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Manduca rustica (rustic sphinx) at Mercury-vapor/ultraviolet light in oak/juniper woodland.

Day 12 – Santa Rita Mountains, Gardner Canyon, Arizona
Rob had noticed a stand of Anisacanthus thurberi (desert honeysuckle)—a host for Spectralia cuprescens—along the road into Gardner Canyon, so together (after morning coffee!) we checked the spot on the way out. The plants were in the early stages of leafing out, and after visually inspecting them for a while and not seeing anything I decided to get out the beating sheet to sample the stand more thoroughly. My eyes did not deceive me—neither one of us found any. Too early? Low population? Who knows! I did beat one clytrine off the plants and collected a few weevils by beating Prosopis glandulosa (mesquite) before finding Acmaeodera parkeri and A. cazieri on the flowers of Allionia incarnata (creeping four o’clock). The flowers were common in the area around the road, but no Acmaeodera were seen until I started scaling the steep hillside nearby—a similar situation in which I’ve found these species on this flower in Box Canyon.

Rob Velten and the author enjoying morning coffee (photo by Margarethe Brummermann).
Dense stand of Anisacanthus thurberi (desert honeysuckle) in Gardner Canyon.
Gardner Creek running full after last night’s rain.
Yours truly with buprestophile extraordinaire Rob Velten.

Huachuca Mountains, upper Carr Canyon, Reef Township Campground, Arizona
I’ve been wanting to explore the higher reaches of Carr Canyon ever since I arrived in Arizona a full week ago. It is the only high canyon in the Huachucas that has a road leading all the way into its upper reaches. At these high elevations the forest is Pinus ponderosa (ponderosa pine) and Quercus hypoleucoides (silverleaf oak). There are many species of woodboring beetles at these high elevations that simply aren’t found down below. Another reason is the recent discovery up here of one of the rarest and most enigmatic of North American longhorned beetles, Placoschema dimorpha. Not even know to science until it was described from Mexico in 2007, it has since popped up here and a few other places in southeast Arizona—some of which have been heavily studied by coleopterists or many decades. Now, I didn’t actually <u>expect</u> to find such a rare thing, but maybe I could get lucky or at least find some other unusual species. The specimen at this location was photographed on a burned pine tree; however, I do not think that is the host (as far as I am aware, no species in the tribe utilizes gymnosperms as larval hosts). I think the host must be oak, as is the case for many trachderines. So, while I kept an eye out for burned pine trees, I also looked for oak, and especially recently dead oaks showing signs of woodboring beetle infestation. I did some of beating on Q. hypoleucoides and had collected just a clerid (Enoclerus bimaculatus) when I came upon a recently fallen Q. hypoleucoides that showed a few buprestid(ish) emergence holes and looked to be “the right age.” Cutting away the bark of the trunk revealed galleries, and chopping into sapwood revealed buprestid larvae in their galleries. I tagged it for retrieval, eventually cutting it up and segregating the trunk sections from the branches. Very nearby, I found another dead Q. hypoleucoides, this one much smaller and apparently cut rather than fallen. Unlike the previous one, however, this one showed the round holes with ejecting frass that indicated infestation by cerambycid rather than buprestid larvae. Cutting into the wood confirmed the presence of such, and so this one also was later cut up and bundled for bringing back. I saw no beetles on the trunks of any of the many fire-scarred pines lofting overhead, but at one point I spotted in the distance the telltale brown flagging of a recently died pine up the slope. Hiking up to it took some effort, but when I reached it the first thing I saw was a giant click beetle—Chalcolepidius apacheanus (Apache click beetle)—nestled against the ground at the base of the trunk (apparently ovipositing?). Inspecting the trunk of the tree itself, I noted just a few buprestid emergence holes that seemed fresh. Once again, chopping away the bark revealed the frass-packed galleries, and chopping into the heartwood revealed a large pre-pupal buprestid larvae. This was put into a vial, and I noted the location so I could return with the chainsaw and “bring ‘er down.” As I was cutting up the oaks, I found some small, recently cut pine branches near where I had parked the car. I found (and accidentally killed) a woodboring beetle larva of some type (I mangled it pretty good, but I think it was a buprestid) in one of them, so that was good enough to earn a spot in the rearing tubs. Unfortunately, I was not able to retrieve the dead pine tree—cutting up the oaks took a fair bit of time, during which darkening clouds gathered over the nearby peaks. Eventually cracks of thunder began piercing the air. It was all I could do to get the oaks into the car and all of the equipment put away before heavy rain drops began pelting the car. I had no idea what the storm would bring, but the last thing I wanted to be was stranded on the top of a mountain on my last full day of field collecting. As it turned out, the storm was more bark than bite (although the sharp drop in temperatures would kill lighting later in the evening). Perhaps I’ll be able to get back up the mountain in the morning and retrieve the pine tree.

Chalcolepidius apacheanus (Apache click beetle) on trunk base of dead Pinus ponderosa (ponderosa pine).

Huachuca Mountains, lower Carr Canyon, Arizona
It seems like forever ago that I began this trip, and now the last night of collecting has arrived. I decided to come back to this spot where I’d had such good luck earlier this week, but I wasn’t optimistic given how rain chased me out of the canyon earlier. My pessimism was warranted, and while I did picked up a variety of things, it included only two longhorns—both rather pedestrian species. No, the real charm of the night came not from collecting insects, but after the lights were down and my mind was free to wander as I leaned back in my chair and gazed into the crisp, dark, starry Arizona sky—its perimeter along the horizon bound by a craggy silhouette of nearby oak trees and distant peaks; from listening to the sounds of the night, alternately focusing on the individual cricket or distant coyote versus the chorus as a whole. Only to the north could I see the faint glow of city lights—the only sign that anything beyond me and this moment exist. These moments happen only once on a trip (maybe twice), and they are to be savored; indelibly stamped into the memory banks for future enjoyment; one of those experiences that, when recalled, is guaranteed to trigger euphoric recall.

Campground set for a final night of lighting.
Manduca sexta (Carolina sphinx) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Sphinx dollii (Doll’s sphinx) at Mercury-vapor/ultraviolet light in oak/juniper woodland.
Sphinx dollii (Doll’s sphinx) at Mercury-vapor/ultraviolet light in oak/juniper woodland.

Day 13 – Huachuca Mountains, upper Carr Canyon, Reef Township Campground, Arizona
I went back up to the top of Carr Canyon to retrieve the dead Pinus ponderosa that I found yesterday. Good thing I did, as I also found my favorite hatchet (which I’d inadvertently left behind yesterday). The 9” diameter trunk was almost too big a job for my Stihl MiniBoss chainsaw, but I kept at it and finally felled the the 25’ tall tree. It took three trips up and down the steep slope—each round trip almost a half-mile—to haul out the upper 10 ft of trunk and associated branches, which I segregated into three batches for rearing: trunk, 1–2” día. branches, and <1” dia. twigs. Now let’s hope the effort was worth it and I get some good species out of the wood.

Pinus ponderosa (ponderosa pine) dominates the high elevations in the Huachuca Mountains.

Huachuca Mountains, near Carr Canyon, Waterfall, Arizona
One last stop to take in the terrifyingly magnificent views from atop Carr Canyon! You can see the road that I traveled up this morning snaking back down the right side of the mountain. The massif to the left is the highest point that you can see from the valley below, but there are much higher peaks behind me.

Terrifyingly steep, magnificently endless drop!
Carr Canyon Rd snakes up the right side of the mountain.
This massif is the highest point visible from the mouth of the canyon.

Epilogue
Sadly, I could squeeze no more stops into the trip—I’d allowed myself two days to make the 24-hour drive back to St. Louis, and it was already almost noon on the first of the two days. I left, however, with a rack full of vials filled with insects and a renewed love for Arizona and the desert southwest that first captured my heart some 37 years ago!

©️ Ted C. MacRae 2021

Stag and “stagette” beetle

Lucanus capreolus, female (L) and male (R) | Fort Defiance Park, Illinois

Lucanus capreolus, female (L) and male (R) | Fort Defiance Park, Illinois

Last month I posted some photos of the very “stag beetle-ish” looking longhorned beetle, Parandra polita. Chestnut brown in color with large, forward projecting mandibles, this member of the longhorned beetle subfamily Parandrinae looks almost nothing like longhorned beetles in other subfamilies but very much like a small species of stag beetle (family Lucanidae). If it weren’t for the straight rather than elbowed antennae, even experienced coleopterists might be fooled by its appearance. The beetle had been attracted to an ultraviolet light setup in wet bottomland forest at the southern tip of Illinois where the Mississippi and Ohio Rivers meet. Perhaps not coincidentally, several true stag beetles representing both males and females of the species Lucanus capreolus were attracted to the lights that night as well.

Males are distinctive by their large, sickle-shaped mandibles.

Males are distinctive by their large, sickle-shaped mandibles.

The genus Lucanus contains the largest stag beetles in North America—the most desirable of the handful of species it contains being L. elaphus (North America’s largest stag beetle) due to the male’s outrageously enlarged mandibles and the species’ general scarcity. Lucanus capreolus nearly matches L. elaphus in size and has an equally broad distribution across eastern North America, but it seems to be a more common species and has the male mandibles only moderately (though still distinctly) larger than the female. Despite its more routine occurrence, I rarely see more than a few individuals at a time, and they are almost always all males. This night, however, I was fortunate to encounter not only males but several females as well. I’ve previously photographed the female of this species (Diminishing Stag Beetle), but this was my first chance to photograph both male and female together.

Females have much smaller mandibles (but are still capable of delivering a painful 'nip').

Females have much smaller mandibles (but are still capable of delivering a painful ‘nip’).

While male L. elaphus are undeniably distinct, I frequently see confusion about how to distinguish male L. capreolus from L. placidus (the third eastern North American species of the genus, occurring more sporadically than L. capreolus), and separating females of all three species can be even more confusing. Male L. elaphus are readily identified by their greatly elongated and multi-toothed mandibles, but a suite of characters may need to be employed for females and non-elaphus males. The best character to use for L. capreolus are the distinctly bicolored femora that are yellowish at the base; however, color can be variable and some individuals will exhibit the more uniform chestnut-brown color typical of L. elaphusLucanus placidus, on the other hand, is usually distinctly darker in color than either of the other two species. Surface sculpture of the elytra and pronotum also offer useful characters. The elytra of L. capreolus and L. elaphus are rather smooth, while in L. placidus they are more distinctly punctate/rugose. The pronotum of both L. capreolus and L. placidus, however, is usually distinctly punctate compared to the relatively smooth pronotum of L. elaphus. The shape of the labrum (projection between the mandibles) is also usually distinctive and is not influenced by gender like the mandibles. In L. elaphus the labrum is rather pointed, while in L. capreolus and L. placidus it is more blunt (indeed, in L. placidus the labrum can almost be described as quadrate, or “squared”). Lastly, the number of teeth on the inner margin of the mandibles is usually diagnostic for females of the three species—L. capreolus possessing one tooth, L. placidus possessing two, and L. elaphus possessing more than two.

Unlike in most insect groups, males rather than females.

Unlike most insect groups, male stag beetles rather than females are generally larger.

© Ted C. MacRae 2014

Diminishing Stag Beetle

This past June I made a couple of trips to north-central Arkansas. They were my first real efforts to collect insects in Arkansas, despite hundreds (literally) of trips to various localities throughout the Ozark Highlands in adjacent southern Missouri. The similarities between the two areas were obvious, yet there was also the feeling of a brand new area just waiting for exploration. On the second trip, I found a campground that looked good for blacklighting to see what wood-boring beetles I might be able to attract amongst the surrounding pine/oak-hickory forest. The evening was warm (very warm!) and humid with no moon—typically ideal for blacklighting, but beetles were sparse at the sheets for some reason (perhaps deterred by the obnoxiously unrelenting yells of drunk Arkansans and their out-of-control offspring?!). The evening, however, was not a total loss—at one point an enormous stag beetle landed on the top of the sheet.  It was so big that I couldn’t even fit it into the viewfinder of my camera:

I fiddled with the camera and changed some settings.  I got a little more of the beetle in the viewfinder this time, but it was still just too big:

Additional fiddling with the camera allowed even more of the beetle to be seen:

As I took the photographs, I even began wondering if the beetle itself was actually shrinking:

Eventually, it turned out to be a normal-sized beetle after all:

This is a female of the common eastern North American species Lucanus capreolus.¹  I don’t seem to encounter female stag beetles as often as the males, so this was still a nice find on an otherwise frustrating night.

¹ Two bonus point in the current BitB Challenge session to the first person who correctly explains how I know this.  Overall contenders: here’s your chance to score an advantage as we enter the final stretch in the current Challenge session.

Copyright © Ted C. MacRae 2011

North America’s largest stag beetle

 

Here is the full-sized photo that provided the image for yesterday’s Super Crop Challenge #3.  The insect in the photo is, of course, a fine example of a male Lucanus elaphus – the giant stag beetle (family Lucanidae).  This striking insect is easily among North America’s most distinctive and recognizable species by virtue of the enormously super-sized mandibles sported by the males.  Its fearsome appearance belies the true nature of this harmless beetle, which spends its days feeding on sap that flows from wounds on the trunks and roots of trees.  Males use their massive mandibles in combat with other males, not for “biting,” but rather as tools to pry and lift their adversaries before dropping them to the ground.  Some marvelous photos of this behavior in a related European species can be seen at Stag Beetles Lucanus cervus Mating Behaviour.

I collected this specimen many years ago at an ultraviolet light (“blacklight”) that I had setup in the pine/oak forests at Pinewoods Lake, Carter Co., in the southeastern Ozarks – one of my favorite 1980’s beetle collecting spots.  This was in my early days of studying beetles, during which time I was actively collecting material as part of my statewide surveys for the families Buprestidae (MacRae 1991) and Cerambycidae (MacRae 1994).  Lucanus elaphus is not a commonly encountered species, especially in the western reaches of its distribution here in Missouri, and I’ll never forget my rabid excitement when I encountered this fine major male at my blacklight sheet.  For many years afterward it remained the only individual that I had ever encountered, until a few years ago when I came across a group of two males and one female feeding on a sap flow in a wet bottomland forest along the Mississippi River in the lowlands of southeastern Missouri.  I encountered another male the following year at a nearby location “rafting” on debris in floodwaters from the nearby river, and two weeks later at that same site I picked up several males and females in a fermenting bait trap.¹  Like most “uncommon” species with broad distribution across the eastern U.S., I suspect that its apparent rarity is an artifact due to habits that make it infrequently encountered rather than being truely scarce.

¹ I have used fermenting bait traps to collect a wide variety of beetles, but especially longhorned beetles.  My recipe is based on that described by Champlain and Knull (1932) – bring 12 oz. dark molasses and 12 oz. beer up to 1 gal. with water, mix well and add a packet of dry baker’s yeast to get the fermentation started.  Hang a 1/2-gallon milk jug with big holes cut in the sides in a tree along the edge of a woods and add ~1 quart of fresh liquid.  It generally takes 2-3 days for the liquid to really start fermenting and become attractive, and it will remain so for about another week or so.  Check traps every 2-3 days by pouring the liquid through a kitchen strainer into another container – reuse or replace as necesssary. Place the collected specimens in vials of water to wash off the molasses residues, and either pin immediately afterward or transfer to 70% ethanol for longer term storage.  Some of the more desireable species I’ve collected in this manner, besides L. elaphus, are Plinthocoelium suaveolens, Purpuricenus axillaris, P. humeralis, P. paraxillaris, Stenocorus cylindricollis, S. shaumii, Sarosesthes fulminans, Stenelytrana emarginata [= Leptura emarginata], and S. gigas [= Leptura gigas].


Congratulations to Ben Coulter and Janet Creamer, both of whom correctly identified the species and most of the mouthparts.  Each earned 14 pts and, thus, tied for the win, while JasonC. earned 5 pts. to take the final podium spot.  The pointed structure is the labrum (its shape distinguishing it from other North American species of the genus), and it is flanked on each side by the fuzzy yellow galeae (derived from the maxillae) and the labial palps.  Nobody correctly named the galeae, which seem to be greatly elongated and hairy in stag beetles as a function of their sap feeding behavior.  A portion of the left maxillary palpus can also be seen in the corner of the photograph, but nobody scored those points either.  Brady Richards just missed the podium, but his witty reference to Gene Shalit (if not immediately picked up on by me) earns him an honorable mention.

With points being formally awarded now beginning with the previous competition (ID Challenge #1), I’ll start keeping an overall leaders board, and with wins in both competitions Ben takes a commanding lead in the overalls with 23 pts, followed by Janet Creamer at 14 pts and TGIQ at 8 pts.  I guess I should start thinking of some sort of tangible prize for winners periodically – suggestions welcome.  Stay tuned for another issue of Super Crop Challenge or ID Challenge in the near future.

REFERENCES:

Champlain, A. B. and J. N. Knull.  1932. Fermenting bait traps for trapping Elateridae and Cerambycidae (Coleop.).  Entomological News 43(10):253–257.

MacRae, T. C. 1991. The Buprestidae (Coleoptera) of Missouri.  Insecta Mundi 5(2):101–126.

MacRae, T. C. 1994. Annotated checklist of the longhorned beetles (Coleoptera: Cerambycidae and Disteniidae) known to occur in Missouri. Insecta Mundi 7(4) (1993):223–252.

Copyright © Ted C. MacRae 2010