2025 Southwestern U.S. Insect Collecting Trip iReport: Act 2 (Finale)

This—the 18th “Collecting Trip iReport”—covers the second of two insect collecting trips to the southwestern U.S. this season—this first one occurring from June 4–13, during which I placed “bottle traps” and “jug traps” at several locations, and this one from September 3–14 to retrieve the traps and take advantage of any late-season collecting opportunities. I was fortunate on this trip to have longtime collecting buddy and melittologist Mike Arduser joining me, during which we visited the same 15 localities that I visited on my precious trip back in June (one in northwestern Oklahoma, six in northern Arizona, six in southern Utah, and two in southern Nevada) plus two additional localities (one each in northwestern Oklahoma and northeastern New Mexico).

Like previous trip reports, this one assembles field notes largely as they were generated during the trip. They have been lightly “polished” but not substantially rewritten or changed based on subsequent examination of collected specimens (unless expressly indicated by “[Edit…]” in square brackets). While I did photograph a few insects in the field with my “macro” rig to feature individually in future posts, this post contains only iPhone photographs (thus the title “iReport”). Previous collecting trip iReports are:
2013 Oklahoma
2013 Great Basin
2014 Great Plains
2015 Texas
2018 New Mexico/Texas
2018 Arizona
2019 Arkansas/Oklahoma
2019 Arizona/California
2021 West Texas
2021 Southwestern U.S.
2022 Oklahoma
2022 Southwestern U.S.
2023 Southwestern U.S.
2024 New Mexico: Act 1
2024 New Mexico: Act 2
– 2024 New Mexico: Finale
– 2025 Southwestern U.S.: Act 1


Day 1

I’m on my way out west with fellow collector Mike—today being only a travel day but with a quick stop in Texas Co., Oklahoma to retrieve bottle traps set back in June and then a night of camping at Black Mesa State Park before continuing the drive west tomorrow. After 7 hours, we needed to stretch our legs and stopped at Salt Plain National Wildlife Refuge’s Sandpiper Trail—a spot we both have visited several times and know well. Recent rains had the alkaline flats filled with more water than I’ve seen during any previous visit, …

Salt Plain National Wildlife Refuge, Alfalfa Co., Oklahoma.

… and only a few Ellipsoptera nevadica knausi (Knaus’ tiger beetle) and Eunota circumpicta johnsoni (Johnson’s tiger beetle) were seen on the drier margins of the alkaline flats. Several Epicauta conferta (red-cornered blister beetle) were seen lumbering across the path, and a Diogmites angustipennis (prairie robber fly) posed nicely on the trail for pictures as well.

Diogmites angustipennis (prairie robber fly—family Asilidae).

Showy Eustoma russellianum (prairie gentian)—a plant I’ve never seen before—were blooming spectacularly and prolifically in the vegetated areas bordering the alkaline flats.

Eustoma russellianum (prairie gentian—family Gentianaceae).

It was a quick but interesting stop that rejuvenated the legs before we continued our journey westward.

The first “real” stop of the trip was ~5 miles north of Goodwell in Texas Co., Oklahoma where I placed three white bottle traps in early June hoping to capture the very rare Acmaeodera robigo, which had been photographed here on flowers of Melampodium leucanthemum (blackfoot daisy) and the photos posted on BugGuide. I was never able to contact the photographer, but since bottle traps are so effective at sampling species of Acmaeodera I reasoned placing three traps here (one at the precise spot and two more several hundred yards to the north and to the south) would give me the best chance of collecting it. It had already been dark for an hour before we reached the spot, but despite the darkness and late hour I had no trouble finding each of three traps. I was happy to see all three traps still in place, undisturbed, and filled to the brim with insects. I tried to pick out the larger insects (mostly crickets and grasshoppers) and stir through the remaining contents of each trap a bit to see if I could detect any Acmaeodera, but the majority of insects appeared to be small blister beetles, followed by bees. The darkness made further sorting impossible, so I bagged the contents of each trap and saved for later sorting. [Edit: later sorting found only a couple of Acmaeodera mixta? in just one of the traps and no A. robigo.]

We arrived at our first overnight spot—Black Mesa State Park—quite late (~11:30 pm) and quickly setup camp before retiring for the evening. In the middle of the night I got up, came out of the tent, and was greeted by an incredible amazing starscape that is normally only seen during winter. Taurus was already high in the sky, and Orion was well above the horizon with a brightly shining Jupiter not too far to its left.

Night sky at Black Mesa State Park. Orion has risen, with Taurus above and Jupiter just rising in the lower left.

Day 2

In the morning, a canyon towhee (Melozone fusca) kept us company as we prepared breakfast and then broke camp for another mostly travel day

Canyon towhee (Melozone fusca).

Our destination this evening is Devils Canyon Campground near Monticello, Utah, to which we will travel by way of northeastern New Mexico and then southwestern Colorado.

Abandoned homestead outside of Clayton, New Mexico.

Shortly after crossing into Colorado we made a pit stop for ice and I began searching the pavement around the gas station looking for beetles that may have come into the previous night’s lights. I didn’t find any cerambycids but I did find a small tenebrionid beetle that didn’t look like familiar to me.

Crossing the Sangre de Cristo and then the San Juan ranges were as spectacular a mountain crossings as any that Colorado has to offer, and a coffee stop in Pagosa Springs (at Faire Society Cafe and Patisserie) provided not only good coffee and pastries to fuel me for the rest of the drive to Devils Canyon Campground near Monticello, Utah, but interesting and creatively framed art work to treat the eyes while waiting for our orders.

“Art wall” in Faire Society Cafe and Patisserie, Pagosa Springs, Colorado.
“Butterflies”—Faire Society Cafe and Patisserie, Pagosa Springs, Colorado.
“Gentleman Frog”—Faire Society Cafe and Patisserie, Pagosa Springs, Colorado.
“Cow”—Faire Society Cafe and Patisserie, Pagosa Springs, Colorado.

Weather during the drive had been good all day, but on the final approach to Devils Canyon Campground the skies began looking worryingly threatening. The last time I came here (early June) I had intended to camp here but got rained out and took a motel in town. I did not want a repeat of that, so we kept our fingers crossed and made the final drive to the campground. Although still threatening, it was not actually raining when we arrived (unlike last time), so we took our chances and set up camp. No sooner than that did the rain start! Fortunately, the tent was already up, so it just meant that instead of cooking dinner at the site, we would instead go to town and have dinner (Dave’s BBQ). When we got back to camp, the rain had stopped (although we could tell that it had rained hard), so I decided to walk the roads looking for night-active beetles. I knew this was probably a fruitless exercise—by then the post-rain temps at this 7000’ site were already down to the mid-50s, but it would give me a chance to stretch my legs after two straight days of driving, and I could also take that opportunity to retrieve the bottle and jug traps that I had set back in June. The bottle trap was disappointing, especially after seeing the ones I’d set near Goodwell, Oklahoma filled to the brim with insects—just a handful of bees (for Mike) and no beetles of any kind. This was surprising given the many Acmaeodera I have collected in alpine habitats just like this (Ponderosa pine and Gambel oak). The SRW-baited jug trap, on the other hand, was nicely (if not overwhelmingly) productive (Tragosoma sp., Enaphalodes sp., small elaphidiines, small acanthocine with very long antennae, Xestoleptura?)—enough to make it worth the effort. I was also pleased to see that the jug trap was in still place and intact with the catch in good shape despite three months in the field. The bait bottle was still about half full of red wine, but since the propylene glycol had dried the trap was no longer trapping insects. As I’d expected, no night-active insects were seen in the way to the traps or on the way back.


Day 3

It was a chilly morning, and though it had not rained since our arrival last night the skies remained overcast. The day’s plan was to continue west to the Ponderosa Grove Campground in southwestern Utah (north of Kanab), but with only five hours of driving required to get there we would have time to make a few stops along the way. Before leaving I started checking the Ericameria nauseosa (rubber rabbitbrush), several of which had begun blooming in the campground, and found a few Crossidius coralinus and Acmaeodera amabilis on the flowers. I was tempted to suggest staying put—at least for a short time—and exploring the area a little more fully, but my real objectives were further west and I elected to stick with the plan.

Crossidius coralinus (family Cerambycidae) on flowers of Ericameria nauseosa (rubber rabbitbrush).
Acmaeodera amabilis (family Buprestidae) on flowers of Ericameria nauseosa (rubber rabbitbrush).

One of the stops I’d made along this way last June was 4 miles east of Bluff in San Juan Co., Utah, where the famous Mont Cazier had collected what would later be described as Agrilus utahensis. I did not find it in June (nor did I fully expect to, since the record was from late July), and I was equally skeptical about my chances this time given how much later it was in the season. On the way here, we got caught behind an oversized load on the highway that was so big it required three highway patrol chaperone vehicles to clear the road ahead. Going at about half the speed limit, I worried we might have to follow it the entire way to our turnoff, which would nearly double our travel time to the first spot. Fortunately, the caravan pulled over at one point to let the long train of trsffic that had accumulated behind it pass, and we were on our way (the convoy would later pass us at the very spot where we had stopped to collect).

Oversized convoy!

The location was disappointing dry and crispy, although Gutierrezia sarothroides (broom snakeweed) was coming into bloom. Sweeping it eventually produced about a dozen Crossidius pulchellus and two other beetles (a clytrine leaf beetle and a weevil), and Mike collected a handsome series and diversity of bees off of flowering Eriogonum sp. (buckwheat).

Crossidius pulchellus (family Cerambycidae) on flowers of Gutierrezia sarothrae (broom snakeweed).

Our drive afterwards through southeastern Utah and northeastern Arizona took us through some of the most amazing scenery that the American West has to offer—red sand/siltstones sculpted through the eons by wind and rain have created a landscape that can only be described as “planetary.”

Monument Valley, southeastern Utah.

Eventually, the dramatic landscape gave way to a more monotonous series of desert plateaus periodically interrupted  by dramatic descents and canyons as we got deeper into north-central Arizona. Though pleasing to the eye, there were few signs of greenery of flowers to tempt the passing entomologist except occasional stands of Ericameria nauseosa (rubber rabbitbrush) beginning to bloom in higher elevation spots. We kept our nose to the grindstone, hoping to see better things once we passed through Page and crossed back into south-central Utah, but the landscape became even crispier, with a stop about 20 miles northwest of Page to look for Nanularia brunnea on Eriogonum inflatum (which I had found two years ago in late June) being a total bust. We expected/hoped that the higher elevations around Ponderosa Grove Campground (~6000’) would provide better collecting and continued there without haste. Our expectations/hopes proved well-founded, as blooming rabbitbrush was seen with greater frequency as we traveled north of Kanab and even more so along Hancock Rd approaching the campground. We took a few moments to scout out a good campsite and setup camp before spending the rest of the available daylight hours exploring. For me the rabbitbrush was most tempting, and scouting plants in the campground and the vicinity east produced small but nice series of Crossidius coralinus and Typocerus balteatus. As I was doing so, a sinking sun and virga to the east produced an impressive rainbow that became irresistible for photography—not only as a subject itself but as a backdrop for the beetles I was finding.

Rainbow at Ponderosa Grove Campground.
Crossidius coralinus (family Cerambycidae) on flowers of Ericameria nauseosa (rubber rabbitbrush).
Typocerus balteatus (family Cerambycidae) on flowers of Ericameria nauseosa (rubber rabbitbrush).

After some downtime back at camp (and grilled sirloin steaks for dinner), I did my customary nighttime patrol to check for night-active insects. This campground was especially productive when I did this back in June and found several Zopherus utahensis and other tenebrionids on the trunks of the massive Ponderosa pines that are the namesake of this campground.

A near-full moon rises over our campsite at Ponderosa Grove Campground.

This time was no different—while I found only a single Z. utahensis, I did also find a few specimens of Coelocnemis sulcata, including a mating pair, on the trunks of the trees …

Coelocnemis sulcata (family Tenebrionidae) mating pair on trunk of Pinus ponderosa (Ponderosa pine) at night.

… and a single Embaphion sp. on the ground at the base of another. There are several massively-trunked Juniperus osteosperma (Utah juniper) in the campground as well, on which another C. sulcata was found. Despite this success and the relatively early hour, I was exhausted and called it quits for the night and retire. We will spend the entirety of the day here tomorrow, so I’ll have another chance to check the tree trunks again tomorrow night.


Day 4

After a relaxing morning at the campsite (during which time I caught up on my field notes while enjoying double-pour-through coffee), I walked over to the sand dune-adjacent woodlands to retrieve the traps that I’d set there back in June and brought them back to the campsite for sorting. I was happy to see Ericameria nauseosa (rubber rabbitbrush) and many other plants in bloom and looked forward to checking them more closely after servicing the trap catches.

Ericameria nauseosa (rubber rabbitbrush) blooming in sand dune habitat at Moquith Mountain Wilderness Study Area.

The yellow bottle trap had ~15-20 beetles, including several Acmaeodera spp., a lepturine cerambycid, and a tiny Dichelonyx-like scarab (a relief after getting skunked with the bottle trap I’d set in Devils Canyon). There were also a fair number of bees in the trap, which I gave to Mike. The SRW-baited jug trap also did well, containing Tragosoma sp., Enaphalodes sp., several Psyrassa sp., and another colorful little lepturine along with several Euphoria inda, several small clerids, a mantispid, and numerous small beetles I take to be oedemerids. After processing the trap catch, I went back over to the woodlands and dunes, spending more than three hours collecting off the flowers of E. nauseosa and other flowers.

Typocerus balteatus was found not uncommonly on the flowers in most of the areas that I covered, while Crossidius coralinus and C. suturalis were found a bit more sparingly.

Crossidius coralinus (family Cerambycidae) on flowers of Ericameria nauseosa (rubber rabbitbrush).
Crossidius suturalis (family Cerambycidae) on flowers of Ericameria nauseosa (rubber rabbitbrush).

I also found Acmaeodera rubronotata on the flowers of Grindelia squarrosa (curlycup gumweed), Dieteria canescens (hoary tansyaster), and E. nauseosa, but they were limited to the woodlands and not seen in the dunes.

Acmaeodera rubronotata (family Buprestidae) on flowers of Dieteria canescens (hoary tansyaster).

I also found several individuals of an interesting little weevil on the rabbitbrush flowers, its gray/black longitudinally striped body making a good subject for photography (for which I also brought back a live A. rubronotata and a mating pair of ambush bugs), and spent a bit of time photographing some of the other blooming plants in the area.

Castilleja linariifolia (Wyoming paintbrush—family Orobanchaceae).

By the time I feel like I’d gotten a good enough look at the area, nearly four hours had gone by and I was famished. Sardines and Triscuits did the trick, after which we did a quick ice run into town and back—the highlights being an authentic Sinclair dinosaur (he’s smiling!) and an real (though non-functioning) pay phone.

The famous “Sinclair dinosaur.”
He’s smiling!
An authentic (though non-functional) pay phone.

Returning to the campground, I walked with Mike back into the dunes to retrieve his bowl traps. I had hoped that some of them would pick up Acmaeodera (as is often the case with bowl traps and why I have started utilizing them on my own collecting trips), but the only species that would be out in this area at this time of season would be A. rubronotata, which I had already collected earlier in the day (there were none).

Mike services a “bowl trap.”

After a period relaxing (with a cold beer and burgers hot off the grill), I began my customary night walk. I have yet to find a cerambycid on tree trunks at night here, but still I enjoy night walks here as much as anywhere due to the consistent presence of ironclad beetles and other tenebrionoids on the trunks of the massive Ponderosa pine trees that give the campground its name.

Another near-full moon rises over the campsite.

Tonight would be no different—I found Zopherus uteanus on just the second tree that I examined (right in our campsite), and I would also find two species of tenebrionids in the vicinity on the trunks of ponderosas (Coelocnemis sulcata and Eleodes obscura sulcipennis) and a third species (ID unknown) on the ground at the base of one of them.

Zopherus uteanus (family Zopheridae) on trunk of Pinus ponderosa (Ponderosa pine) at night.
Coelocnemus sulcata (family Tenebrionidae) on trunk of Pinus ponderosa (Ponderosa pine) at night.
Eleodes obscura sulcipennis (family Tenebrionidae) on trunk of Pinus ponderosa (Ponderosa pine) at night.

At that point, I decided to go outside of the loop towards a couple of large P. ponderosa at the entrance, and on the way I found another E. obscura sulcipennis on the trunk of a massively old Juniperus osteosperma (Utah juniper). There was nothing on the P. ponderosa trees that I had targeted, but nearby was another large one, and high up on the trunk (as far as my fully extended net could reach) was another Z. uteanus, and several C. sulcata—including a mating pair—were found on the ground at its base.

At that point I decided to limit my tenebrionoid collecting to only Zopherus unless I saw species that I hadn’t seen before, so I passed by some additional tenebrionids of the same species as the previous on a large P. ponderosa in the RV loop, then saw a large, multi-branched Pinus edulis (Colorado pinyon pine) that I thought might be interesting to check. I did not find any beetles on it, but I did locate at eye level—with considerable difficulty!—a male Oecanthus californicus (western tree cricket). I was not only able to take a photograph of it with its wings fanned but also record an audio track up close (posted on iNaturalist).

Oecanthus californicus (western tree cricket—family Oecanthidae) male singing on trunk of Pinus edulis (Colorado pinyon pine) at night (song).

Returning to the tent loop, I checked all of the large P. ponderosa trees and junipers that had—during the past visit and last night—been so productive, but the only beetles I found was a very small tenebrionid that, fortunately, was yet another species I had not previously seen here (ID unknown). After two hours of searching tree trunks, I called an end to the night, which also meant a close to the collecting at this spot—tomorrow we will drive to Kyle Canyon in southern Nevada!


Day 5

Our exit from Ponderosa Grove took us through more of the spectacular canyonland that southern Utah is famous for and past the incredible Coral Pink Sand Dunes State Park (so spectacular it is that I think it could—despite its small size—be a national park). A dramatic descent down into the Virgin River valley and the town of Hurricane was only the first such descent—the second one being even more dramatic as I-15 dropped off the edge of the Colorado Plateau along the Virgin River Gorge. The remainder of the drive to Kyle Canyon northwest of Las Vegas was mindnumbing in its contrast—an endless stretch of interstate highway through a vast expanse of low, hot, featureless desert punctuated at regular intervals by palm tree dotted oases, each with a gaudy, glittering casino at its center. Driving up Kyle Canyon Rd put an end to this, however, as each thousand foot gain in elevation brought with it an increasingly interesting landscape. At about 4500’ elevation, I had set a couple of bottle traps—one yellow, one blue—hoping to catch the recently described Acmaeodera raschkoi (whose namesake—Mike Raschko—I had happened to meet at Ponderosa Grove Campground last June a few days before I set the traps!). Mike R. had also placed a bottle trap (white) at the site and was kind enough to reset it for me so I would have three colors sampling the area. Both of my traps were still in place, intact, and filled with numerous Acmaeodera that I take to be A. quadrivittata (along with many bees for Mike A.) and the yellow also containing a larger species that I didn’t immediately recognize (not unusual since I have never collected this area). Unfortunately, I did not see any specimens that appeared to be A. raschkoi, and even more unfortunately the white bottle trap had been pulled from the ground (although I was able to recover a few A. quadrivittata that were still inside the trap). Not much else was going on at the site—only a few things in sparse bloom but no beetles visiting the flowers, nor was there any rabbitbrush around on which to look for Crossidius.

We got a scare when we arrived at the campground and saw a sign at the entrance saying “Campground Full.” This was bad—if this campground was full, then surely the much more heavily used one down below was also full, and I didn’t look forward to spending the rest of the afternoon scrambling for a campsite somewhere in the Spring Mountains. We drove through the campground anyway, and, in fact, there were many campsites available! Looks like somebody forgot to do their job!

“Campground Full” sign at a not-full Hilltop Campground above Kyle Canyon, Nevada.

Crisis averted, we selected a nice spot overlooking the desert below and set up camp. Cool evening temps come early at this high elevation (~8300’), so with the remaining afternoon hours I retrieved and sorted my SRW-baited jug trap, finding several Tragosoma sp. but, curiously, not a single other longhorned beetle (or any beetle for that matter). Searching around the area afterwards, I extracted a dead Dicerca tenebrosa partial carcass in its emergence hole in a stump of Pinus monophylla (single-leaf pinyon pine), then went back down to the area I had collected last time, focusing especially on the two large half-dead Juniperus osteosperma (Utah juniper) trees on which I had seen damage from the rarely collected Semanotus juniperi (on one of which I collected three adults later that night). There was nothing on the first (the one on which I found the beetles), but at the second one I saw a large wind-thrown branch that I had not noticed last time. It exhibited S. juniperi damage and emergence holes on the lower part, and chopping into it I quickly recovered a dead but intact carcass of yet another S. juniperi adult. Further chopping turned up nothing, and I was about to walk away when I thought maybe I should cut into the upper part of the branch as well to look for evidence smaller woodboring species. Doing so, I quickly encountered a Chrysobothris sp. larva, and with that I decided to bring the entire upper part of the branch back for rearing. There wasn’t much else going on—few plants were in flower and nothing was seen on the trunks of trees or various pieces of downed wood that were laying about, so I went back to camp.

Alpine pinyon/juniper woodland at Hilltop Campground above Kyle Canyon, Nevada.
An elevation of 8300’ affords spectacular views into the desert below.

Later in the evening, we watched a spectacular moonrise, then enjoyed “surf ‘n’ turf” (grilled sirloin steak and salmon) before I started up my customary night walk to check tree trunks for night-active beetles.

Angel Peak Radar Station at sunset.
“Corn Moon” rising.
Peeking through the trees.

Last time here when I did this, I found not only S. juniperi but also a few Zopherus uteanus, so I was hopeful for my chance tonight despite the lateness of the season. It started out well—on the first P. monophylla tree that I checked (right at our campsite), I found Oeme costata and a weevil.

Oeme costata (family Cerambycidae) on trunk of P. monophylla (single-leaf pinyon pine) at night.

Those would be the last live beetles I would see (other than an occasional tenebrionid beetle, none of which I collected). However, I would still find success—back at the J. osteosperma on which I had found three S. juniperi back in June, I found three more. They were not alive, however, but dead carcasses at the base of the tree—two nearly completely intact and the other partially so.

Western deer mouse (Peromyscus sonoriensis) at night.

Day 6

We had planned to visit few localities at middle and lower elevations but stopped to check out the profusely-blooming Ericameria nauseosa (rubber rabbitbrush) right outside the campground entrance for bees and beetles. Curiously, hardly anything was seen on the flowers despite the by then late-morning hour, sunny skies, and temps above 70°F. It seemed odd to me that there were no Crossidius beetles on the blooms, and the thought occurred to me that maybe the occurrence of E. nauseosa in the area itself could be a relatively recent phenomenon since it is only seen—albeit profusely—along the roads and highways in the area but nowhere further within the native habitats. With nothing going on, we pushed down to the lowest elevation point that caught our eye on the way in yesterday—the Step Ladder Trailhead at ~6700’.

Again, E. nauseosa was blooming profusely around the parking lot, but a quick perusal made it clear the situation would be similar here as well. Mike, on the other hand, was having good success collecting bees off of E. nauseosa and especially Gutierrezia sarothrae (broom snakeweed), so I was content to stay and refocus by searching for infested wood. I noticed a lot of Quercus gambelii (Gambel oak) and quickly found one with a dead but still attached, fully-barked branch. I broke the branch off the tree, and there in the broken butt of the branch was a large chrysobothroid larvae! This quickly prompted a decision to collect the infested branch and put it up for rearing. A second larvae was found in a small dead (but still fully-barked) tree nearby, which was added to the bundle. While this was going on, Mike found what seemed to be Agrilus blandus? in his net while sweeping bees from the flowers of G. sarothrae. I recall collecting this species in southern California in flowers of Eriogonum (wild buckwheat), so finding it on other flowers—particularly if Eriogonum is in the area (but not seen because it was not blooming) did not seem out of the question. My much more thorough sweeping of the plants around the area where he found it, however, produced no additional specimens. I was also interrupted in my sweeping attempts by a couple of curious bystanders—one a woman from Ukraine who wondered what the plant was that I was sweeping (I told her “broom snakeweed”) and what it was good for (“brooms” I wryly replied and then quickly clarified its role in the ecosystem), and then offered me a beetle collecting tip by telling me about large beetles they call “bombers” and that bite people sitting in spas in Southern California (I presume these are diving beetles in the family Dytiscidae); and the other a young man who was pleased to hear I was from St. Louis because he used to live there when he was married to his ex-wife. This all happened while I was in the middle of my sweeps, so I held the net bag firmly to keep insects from escaping my net until I could resume my sweeping. Eventually, I gave up the ghost and resumed my search for dead, infested wood, eventually finding a Cercocarpus ledifolius (curl-leaf mountain mahogany) tree with one recently-dead and one older dead branch, the former buprestid-infested (verified by cutting into the wood and finding young buprestid larvae) and which I collected for rearing.

View of Fletcher Peak from Step Ladder Trailhead.

After lunch back at the campground and some time spent processing specimens (as well as enjoying the antics of our resident golden-mantled ground squirrel [Callospermophilus lateralis certus]), …

Mount Charleston golden-mantled ground squirrel (Callospermophilus lateralis certus).

… I wanted to check out the nearby Deer Creek Picnic Area where I’d seen a lot of iNaturalist observations (suggesting it might be an interesting place). At first all I saw was the massive parking lot below an equally massive road-cut slope—the only thing that looked like a trail was a steep drop down to the creek below. I checked it out, only to find it dead-ending at the creek and clambered back up.

A massive Ponderosa pine looms above.

Then I saw a gravel trail behind the guardrail on the opposite side of the highway and found it leading to a paved path up the creek. Much of the trail was covered with a deep layer of gravel from flooding (and indeed some of the picnic tables were also nearly completely buried). I hiked the trail as it ascended alongside the creek under massive ponderosa pines until it dead-ended at a gravel road and turned around. The only plants in flower was Ericameria nauseosa (rubber rabbitbrush), and it was only near the highway, and while I saw no insects that I wished to collect, I did see a large Adejeania vexatrix (orange bristle fly) that frustrated my attempts to photograph it until I finally “pre-set” the focus, exposure, and zoom and quickly fired off a few shots at the distance I’d set it for as soon as the fly landed. The virtual lack of insect activity here confirmed what we’ve been seeing in the area as a whole, so I’ll be anxious to leave tomorrow and head for (hopefully) greener pastures at Leeds Canyon back in southwest Utah.

Adejeania vexatrix (orange bristle fly—family Tachinidae) on flowers of Ericameria nauseosa (rubber rabbitbrush).

Day 7

The drive from Kyle Canyon to Leeds Canyon was essentially a straight shot on I-15—normally a recipe for extreme boredom; however, coming back up through the Virgin River Gorge was a different, even more awe inspiring experience than the descent two days earlier. Ascending such a steep narrow canyon has the breathtakingly tall canyon bluffs looming high overhead, dwarfing the traffic, even the largest semi tractor trailers, snaking up below, whereas descending into a seemingly bottomless chasm feels a little more “dangerous.” We arrived at Leeds Canyon relatively early thanks to the “only” 3-hours drive. The area looked very dry, but a variety of blooming plants kept us optimistic as we made our way up the canyon road towards Oak Grove Campground at the top.

Leeds Canyon in southwestern Utah.

Sadly, optimism turned to dismay in an instant when we encountered a “Road Closed” sign about halfway up—a result of the ongoing fire risk that has bplagued the area this season. We checked to see if the campground on the other side of the mountain range was available, only to learn that it was closed due to fire damage. At that point, our decision was made for us—we would need to continue another two hours to the Kaibab Plateau where I had my last sets of traps to retrieve and where we could camp at Jacob Lake. While we were here, however, we took the opportunity to stop at a spot along Leeds Creek and see what we could find.

Oak Grove Rd crosses Leeds Creek in the lower part of Leeds Canyon.

Several different plants were in bloom, on which I’d hoped to find either Acmaeodera or longhorned beetles, the first that I looked at being Dieteria canescens (hoary tansyaster), but I only saw small dasytines (a few of which I collected). Nothing was seen on Solidago velutinus (velvety goldenrod) or Sphaeralcea grossulariifolia (gooseberry leaf globemallow) flowers, but then Mike came up with a Crossidius discoideus on flowers of Gutierrezia sarothrae (broom snakeweed).

Crossidius discoideus (family Cerambycidae) on flowers of Gutierrezia sarothrae (broom snakeweed).

Careful searching of the plants in the surrounding area atop a small hill turned up an additional half-dozen individuals, but none were seen on any of the plants further up or down along the road. By this time, we’d spent about an hour and decided to finish the additional 2-hours drive needed to get to Jacob Lake.

Megaphorus sp. (family Asilidae).
Historic home in nearby Harrisburg, Utah.

Some of the western U.S.’s worst fires this season occurred on Arizona’s Kaibab Plateau. The Dragon Bravo Fire destroyed over 100 structures on the Grand Canyon’s North Rim—including the historic Grand Canyon Lodge and the North Rim Visitor Center, while the White Sage Fire simultaneously burned significant areas of Kaibab National Forest north and east of Jacob Lake. It was the latter that, unfortunately, swept across both of the sites where I had placed traps a month earlier in June, so I was not optimistic about the likelihood that they had survived. Fortunately, the fires did not reach the immediate vicinity of Jacob Lake, so the campground was unaffected and—unusual in my experience—nearly devoid of people. After setting up camp, I went back north into the burn zone to see if my traps 1) had survived and 2) could be retrieved. The area around the site was almost completely destroyed, with charred black skeletons of trees dotting blackened soils devoid of any vegetation.

Aftermath of the White Sage Fire in the Kaibab National Forest.

I had low expectations for the traps at this site even before the fires, as the area had already burned several years early and was in the early stages of recovery (I had decided to place traps here anyway because I wanted to see what the woodboring beetle fauna in a recovering area might look like).

Blackened trunks punctuate a stark landscape.

I continued walking the 2-track toward the trap location surveying the damage, came around a bend, and saw it—a lone, still-green pinyon pine with my jug trap hanging from a branch and a bottle trap, its yellow funnel only slightly heat deformed, still planted in the soil beneath the tree!

A lone green tree amongst the destruction—with my traps in the tree and below it both intact!

At first I was elated, but then I saw the jug trap reservoir was dry and almost completely empty save for a few dried beetle carcasses—the trap had survived the fires, but the associated winds had blown the trap and dumped the contents (none of which could be detected on the ground beneath). The bottle trap, on the other had, looked to be full of insects with plenty of liquid still in the reservoir, so I was hopeful that I would retrieve some good specimens from it. This proved to be the case (sort of!) as I pulled a few Anthaxia sp., a meloid, and lots of bees (for Mike) from the trap. The dried carcasses in the SRW-baited jug trap turned out to be an elaphidiine & several silphids.

After leaving the first trap site and driving towards the second (a few miles east of Jacob Lake), I saw little to no fire impacts as I continued east of Jacob Lake. However, as I got closer to the site I began to see impacts—first along the ridge above, then down the slope and engulfing the area where I had placed my traps. Fortunately, the fire did not seem to have been as severe in the immediate area, so I remained hopeful.

Intact jug trap in burned woodland east of Jacob Lake, Arizona.

The bottle trap was found first and was in much the same condition as the bottle trap at the previous location—it’s blue funnel slightly heat-deformed, but the reservoir was filled with liquid and insects. Later sorting yielded an Acmaeodera diffusa?, a Melanophila sp., a couple of Anthaxia sp., a clerid, and lots of bees (for Mike). The Melanophila sp. was especially welcome—known collectively as “fire beetles” for their attraction to active fires, its presence in the trap may have been been a direct result of the fire. The SRW-baited jug trap was quickly found next, and much to my relief the trap was not only intact and undamaged but also filled with insects (in fact, the propylene glycol had not even completely dried). Later sorting would yield only a single longhorned beetle (plus a silphid and a Euphoria inda), but that longhorned beetle would prove to be the catch of the trip—Calloides nobilis mormonus! I have reared a single individual of the nominate subspecies from fire-damaged oak collected in Missouri, so I suspect the presence of this beetle is also a direct result of the fires that swept through the area—a satisfying irony.

Calloides nobilis mormonus (family Cerambycidae) attracted to sweet red wine-baited jug trap.

Back at camp and after another “surf & turf” dinner of sirloin steak and salmon, I did my customary nighttime walk to look for night-active beetles on the ground and in tree trunks. I had good luck with this here back in June, finding a Zopherus uteanus and several other beetles, but tonight’s catch consisted of just a single Temnochila sp. (family Trogosittidae) and a single weevil (superfamily Curculionoidea) crawling on the large trunks of Pinus ponderosa.

Unidentified weevil (superfamily Curculionoidea) on trunk of Pinus ponderosa (Ponderosa pine) at night.

Day 8

I’d seen a fair amount of Ericameria nauseosa (rubber rabbitbrush) in bloom at the second site the previous day when I retrieved my traps but didn’t see any insects (or, at least, beetles or bees) on them. I figured this was due to the early evening hour, so we decided to come back during the day to try again. The E. nauseosa flowers were still, puzzlingly, devoid of insects (save for honey bees and enormous numbers of a large, black, bristly tachinine fly—possibly Archytas metallicus or Juriniopsis adusta).

Tachinine fly (possibly Archytas metallicus or Juriniopsis adusta—family Tachindae) on flowers of Ericameria nauseosa (rubber rabbitbrush).

Like at Kyle Canyon, the absence of Crossidius spp. on E. nauseosa flowers was surprising, as I have seen them on this plant at almost every other location I have ever checked. You can’t make things appear no matter how hard you look, however, so we continued into some openings farther up the slop and encountered a few other plants in bloom. One was Dieteria canescens (hoary tansyaster), on the flowers of which I found a few small beetles of an unknown family. Nearby I saw several dead main branches in a clump of Quercus gambelli (Gambel oak)—cutting into them revealed a very small woodboring beetle larvae, so I collected several of the stems for rearing. Coming back down the slope I found a single cryptocephaline on the flower of Eriogonum racemosa (redroot buckwheat) and a single Acmaeidera rubronotata on flowers of Gutierrezia sarothrae (broom snakeweed)—the latter on which Mike had also found a single Crossidius discoideus. (This seems to be the first record of any species of Crossidius from the Kaibab Plateau! Maybe the other species are here as well but are not quite out yet at this relatively early date in fall.)

Crossidius discoideus (family Cerambycidae) collected on flowers of Gutierrezia sarothrae (broom snakeweed).

We were getting ready to leave when I spotted a large, fallen Pinus ponderosa (Ponderosa pine) with the twigs brown but still attached (indicating it might be the right “amount” of dead to host woodboring beetles. Damage by such could be seen on the smaller branches, and cutting into them confirmed the presence of larvae and led to a second wood-cutting/bundling session to bring the beetle-infested twigs and branches back for rearing.

Chasing more floriferous pastures, we went back up to higher elevations and stopped at a spot close to the campground where we 1) saw a great diversity of plants in bloom and 2) could safely pull off the highway. A huge diversity of blooming plants were seen (from which Mike collected ~20 species of bees), but the only beetles of interest that I saw and collected were numerous small black/red cryptocephalines on the flowers of Eriogonum racemosum (redroot buckwheat). After Mike was satisfied he’d sufficiently sampled the diversity of bees at the site, we looked for another place to collect.

Eriogonum racemosa (redroot buckwheat—family Polygonaceae).

We drove south towards DeMotte Campground on Hwy 67, but much of the landscape was complete devastation due to the fires and no access was allowed beyond Kaibab Lodge. It was depressing to see the immense scale of destruction and loss of natural resources, but as one forest worker that Mike talked to put it, “It’s just trees, and he didn’t have to call anybody’s family [i.e., there was no loss of life].” We then drove back down past my second trap location where we had collected earlier in the day to see if we could find better stands of Gutierrezia sarothrae (broom snakeweed) on which to look for more Crossidius discoideus (still represented in the area by the single individual Mike had collected earlier). We drove through even more complete and utter destruction but eventually found undamaged areas at about 6600’ elevation. Not only was there G. sarothrae in bloom, but also Chrysothamnus visicidiorus (green rabbitbrush) and Ericameria nauseosa (rubber rabbitbrush)—all three species serving as hosts for various Crossidius and providing a perfect scenario for comparing and contrasting the sometimes-tricky-to-identify plants! The promise of Crossidius, however, would not be realized, and after an hour of searching—finding only a lone weevil on the flowers of E. nauseosa—did I finally admit defeat and concede that Crossidius from this after would have to come some other time.

Inflorescences of Gutierrezia sarothrae (left), Chrysothamnus viscidiflorus (middle), and Ericameria nauseosa (right). Note the distinct ray florets of G. sarothrae and blue-green leaves of E. nauseosa.

I hadn’t planned to do my customary nighttime walk this evening—my motivation waning after the less than meager success of the previous night, continuing relative absence of beetle life I’d seen during the day, and temps now dropping towards the 50s. A cute but shy Uinta chipmunk in our campsite captured my photographic interest as we relaxed, and the setting sun turned the clouds a stunning pink!

An Uinta chipmunk (Neotamias umbrinus) climbs a Ponderosa pine to stay as far from my camera as possible!

However, as night fell the siren song of the nighttime walk began singing its tune and I was unable to resist.

The setting sun turns the clouds an exceptional pink.

It turned out to be a more successful night than I’d expected—I found a small species of tenebrionid [Edit 10/1/25: Eleodes pimelioides] on the rocks surrounding the campsite, and there turned out to be dozens of the little buggers crawling though the pine duff in the campground.


Eleodes pimelioides (family Tenebrionidae) in alpine coniferous forest at night.

Nothing else was seen, however, and since tomorrow would be a long travel day I called it an early night.


Day 9

On a mostly travel day, we tried to take a big bite out of the many miles that still separate us from St. Louis, where we planned to be in three days time. We got another look at the devastation east of Jacob Lake before reaching the dramatic drop off the Kaibab Plateau and down into the Vermilion Cliffs National Monument—their massive red bluffs accompanying us all the way to Mojave Canyon. Just south of there as the highway climbed up and out of the valley, we made a quick stop to remove excess clothing (having gone from high elevation to low), and alongside the road I spotted a Crossidius pulchellus on the flowers of Gutierrezia sarothrae (broom snakeweed). This is the furthest southwest I have collected this species, and I was tempted to continue looking to see if I could find the other G. sarothrae associate—C. discoideus—as well. We still had a long drive ahead of us, however (destination Mills Rim Campground in northeastern New Mexico), so I resisted temptation and we continued the drive.

The Vermilion Cliffs, stunning as they were, still weren’t the highlight of the drive. That honor would come from the moonscape formations along Hwy 89 as it followed the Echo Cliffs and then turned east onto Hwy 160 towards Tuba City. We then passed through a series of stunning plateaus and drops on Hwy 264 as it passed through the Navajo and Hopi Reservations. One abandoned house as we dropped down off the Ganado Mesa was especially picture-worthy.

Abandoned home on the Ganado Plateau near Kykotsmovi Village, Arizona.

Eventually we crossed the state line at Picture Rocks into New Mexico, and, suddenly, the landscape seemed more “tame” and less hostile. I don’t normally like interstates, and the stretch of I-40 to Albuquerque did nothing to change my opinion of them, but I must admit that I-25 north towards Santa Fe was among the most picturesque I have ever seen. Eventually, we left the mountains and found ourselves once again on the western edge of the Great Plains—its vast featureless expanse a true contrast to the landscape we had witnessed throughout most of the day. This apparent homogeneity, however, is misleading—tucked away in places unknown to most are some remarkable natural areas, and Mills Rim is one such place. We arrived after dark, so the explorations of its hidden charms would have to wait until the next day, but after getting camp set up I did a short nighttime walk to see what was out and about.

Night sky at Mills Rim Campground, Harding Co., northeastern New Mexico.

Only one beetle, Stenomorpha sp. (family Tenebrionidae) ambling across the ground, was seen.

Many plants in bloom were also seen however, so I went to bed optimistic about my prospects for finding beetles the next day.


Day 10

Mosquitoes were bad during the previous night, and they were bad again the following morning, prompting liberal use of repellent to a much greater degree than I am used to. At the same time, the presence of mosquitoes indicates abundant moisture in an area, and it was with that optimism that I set about searching for jewel beetles, longhorned beetles, tiger beetles, and whatever other insects could catch my eye in this hidden jewel of a place. Surrounded by treeless grasslands (and preserved as the Kiowa National Grasslands), Mills Rim Campground sits at the edge of Mills Canyon—a chasm in the landscape at the edge of a plateau bordering the foothills of the Rocky Mountains. Firmly embedded in the Great Plains, the juniper/pine/oak woodland at the edge of and down in the canyon features plants and animals at their easternmost extent—residents of the Rocky Mountains that have found an isolated home in the middle of the grasslands. Pinus edulis (Colorado pinyon pine), Pinus ponderosa (Ponderosa pine), Juniperus monosperma (one-seed juniper), Juniperus scopulorum (Rocky Mountain juniper), Quercus grisea (gray oak), and Quercus × undulata (wavy leaf oak) all make their homes here, hosting innumerable insect species that are normally more at home in the Rocky Mountains.

View into Mills Canyon from Mills Rim Campground.

I’ve collected here several times and recorded many different western species of beetles, but the height of the season seems to be in June and July. Mid-September, in contrast, seems to be near the tail end of the season, the numbers and diversity of beetles and other insects dropping from their highs earlier in the season. The flowers of Gutierrezia sarothrae, however, were hosting lots of insects. In addition to Crossidius pulchellus, Chauliognathus basalis, Bothrotes canalicularis, and Collops sp., Mike found numerous bee species, mostly females, collecting pollen from the flowers.

Crossidius pulchellus (family Cerambycidae) on flowers of Gutierrezia sarothrae (broom snakeweed).
Chauliognathus basalis (family Cantharidae) on flowers of Gutierrezia sarothrae (broom snakeweed).
Bothrotes canalicularis (family Tenebrionidae) on flowers of Gutierrezia sarothrae (broom snakeweed).

A couple of species of robber flies—Ospricerus sp. and Efferia sp.—were also taken in flight, presumably patrolling the flowers of G. sarothrae for bee prey.

Ospricerus sp. (family Asilidae) patrolling flowers of Gutierrezia sarothrae (broom snakeweed) for bee prey.
Nysius sp. (false chinch bugs—family Lygaeidae) on flower of Erigeron divergens (spreading fleabane).
Mirabilis linearis (narrowleaf four o’clock—family Nyctaginaceae).

After a rest and rehydration break, I followed the road down Mills Canyon as it approaches the Canadian River to see if I could find Ericameria nauseosa—should I be able to, it would surely be at or near the easternmost limit of occurrence for the species in this part of New Mexico.

Mills Canyon above the Canadian River.
A bit further down Mills Canyon Road.

About half  a mile down the road I began seeing Acmaeodera rubronotata on the flowers of G. sarothrae, and I eventually secured a series of about a half dozen specimens. This is a nice record, as I found a single specimen a couple of years ago at Black Mesa in northwestern Oklahoma—a new state record and northeastern range extension, and this record helps bridge the gap between that record and the species’ more normal range of distribution across New Mexico and Arizona. Finally, nearly a mile into the canyon, I found one large blooming E. nauseosa and a smaller pre-bloom plant, but there were no Crossidius beetles on them, nor were any additional plants were seen a hundred yards or so further down the road, so I turned around.

Ericameria nauseosa (rubber rabbitbrush—family Asteraceae) along Mills Canyon Road.

On the way back out of the canyon, I collected a Calopteron sp. (family Lycidae) on senescing Melilotus alba (white clover) and photographed a female Stagmomantis limbata (Arizona mantis—family Mantidae).

Stagmomantis limbata (Arizona mantis—family Mantidae) on flowers of Gutierrezia sarothrae (broom snakeweed).

Back at camp, a couple of scarabaeoid beetles flew to the light of the lamp while we were relaxing with refreshments in hand and waiting for the coals to be ready. This suggested that maybe conditions were favorable for setting up the ultraviolet lights to attract other insects.

A final night of relaxing at camp with refreshments in hand.

After finishing dinner, we did exactly that and saw a few interesting insects show up, but shortly after setting up the lights the wind began to kick up, the temps began to drop, and increasingly frequent gusts making further lighting impossible!

Stagmomantis limbata (Arizona mantis—family Mantidae) male attracted to ultraviolet lights.
Schistocerca nitens (gray bird grasshopper—family Acrididae) attracted to ultraviolet lights.
Ophion sp. (short-tailed ichneumon wasp—family Ichneumoidae) attracted to ultraviolet lights.

Day 11

It was a rather sleepless final night in the tent—winds whipped as we turned in for the night, and shortly afterwards we awoke to rain splattering our faces through the fly-less tent roof. We quick got up and put on the rain fly, then listened to light but steady rain for most of the rest of the night. By the time we got up it had mostly stopped, but cool conditions with low-hanging, fast-moving clouds caused us to quickly break camp and save coffee and breakfast for Mocks Coffee Shop in Clayton, New Mexico near the Oklahoma state line (let me tell you how difficult it was for me to drive two hours first thing in the morning without coffee!). We had wanted to make our final collecting stop at a a lot near Kenton in the Black Mesa area of extreme northwestern Oklahoma, but the forecast for the area showed only slightly warmer temperatures and very gusty winds. This would make collecting there pointless, so we instead traveled four more hours east to get in front of the cold front at another of our favorite collecting spots, Gloss Mountain State Park in Major Co.

Gloss Mountain State Park features gypsum-capped mesas atop Permian red siltstones.

I was hoping to see an attractive late-season jewel beetle—Acmaeodera macra, which I had collected here and at nearby Alabaster Caverns State Park in previous years during late September. Temps were good (well over 80°F) when we arrived but the hour was already late (near 4:00 p.m.), so we had limited time for collecting before insects would start bedding down for the evening (usually around 5:00 to 6:00 p.m. at this point in the season). Megatibicen dorsata (bush cicada) and Neotibicen aurífera (prairie cicada) males were still singing abundantly, filling the air with their distinctive songs (video of M. dorsata male singing here).

Megatibicen dorsata (bush cicada—family Cicadidae) male singing on my hand after being taken from the stem of Helianthus annuus (annual sunflower).

Helianthus annuus (annual sunflower) and Grindelia ciliata (wax goldenweed) were blooming prolifically, off the flowers of which Mike collected a fair diversity of bees. I’d hoped to find beetles on the flowers as well, but they were limited almost exclusively to Chauliognathus limbicollis.

Chauliognathus limbicollis (family Cantharidae) on flower of Grindelia ciliata (wax goldenweed).

I did collect a male/female pair of Tetraopes femoratus (red-femured milkweed borer) on the seed pod of Asclepias engelmanniana (Engelmann’s milkweed) and photographed the striking and beautiful caterpillar of Schinia gaurae (clouded crimson moth) on the stem of Oenethera glaucifolia (false gaura).

Tetraopes femoratus (red-femured milkweed borer—family Cerambycidae) on seed pod of Asclepias engelmanniana (Engelmann’s milkweed).
Schinia gaurae (clouded crimson moth—family Noctuidae) on the stem of Oenethera glaucifolia (false gaura).

After that, I went up on top of the main mesa where I expected A. macra to occur. Heterotheca stenophylla (stiffleaf false goldenaster), on the flowers of which I collected A. macra in previous years, was blooming abundantly, but intense searching their flowers produced no beetles. I also noticed that Gutierrezia sarothrae (broom snakeweed), abundant in the area as well, was in only the earliest stages of bloom, suggesting to me that it might still be a bit too early for the jewel beetles to be out. By the time the 6:00 hour arrived, insect activity was noticeably diminished, and we wrapped up this, our final, collecting stop of the trip.

No camping is available at Gloss Mountain State Park, so we knocked out another hour and a half of travel by driving to Ponca City in north-central Oklahoma and taking a hotel there. For the first time since we left, we enjoyed dinner at a restaurant (fried catfish for me!), a hot shower, and a real bed!


Day 12

The following morning, we were surprised to learn that the only coffee shops in town were Starbuck’s and drive-throughs. This just wasn’t going to cut it for us on our final travel day, so we drove 15 minutes north to the small town of Newkirk and enjoyed great coffee, breakfast sandwiches, and scones at “Savvy Cactus” (Newkirk Mercantile Boutique & Espresso Bar). (The coffee was good enough that I bought a bag of their coffee!)

“Savvy Cactus” Newkirk Mercantile Boutique & Espresso Bar in Newkirk, Oklahoma.

The rest of the drive back to St. Louis was spent reflecting on the many experiences we’d just had and synthesizing the new knowledge while enjoying the landscape as it skirted the southern edge of the Flint Hills of Kansas and traversed the familiar hills and dales of our beloved Missouri Ozarks—the end of a 3,931-mile trip!

62.3 mph average for the trip.

©️ Ted C. MacRae 2025

2025 Southwestern U.S. Insect Collecting Trip iReport: Act 1

Welcome to the 17th “Collecting Trip iReport” covering the first of two planned insect collecting trips to the southwestern U.S. this season—this first one occurring during June 4–13. This was another solo trip, and while it was not as long as I’d hoped, I still managed to visit 15 different localities—one in northwestern Oklahoma, six in northern Arizona, six in southern Utah, and two in southern Nevada.

Per usual, this report assembles field notes largely as they were generated during the trip. They have been lightly “polished” but not substantially modified based on subsequent examination of collected specimens unless expressly indicated by “[Edit…]” in square brackets. Again, I brought my “big” camera and took macro photographs of certain insects in the field that will be featured in future individual posts. However, this “iReport”, as always, features iPhone photographs exclusively. Previous collecting trip “iReports” are:
2013 Oklahoma
2013 Great Basin
2014 Great Plains
2015 Texas
2018 New Mexico/Texas
2018 Arizona
2019 Arkansas/Oklahoma
2019 Arizona/California
2021 West Texas
2021 Southwestern U.S.
2022 Oklahoma
2022 Southwestern U.S.
2023 Southwestern U.S.
2024 New Mexico: Act 1
2024 New Mexico: Act 2
– 2024 New Mexico: Finale


Day 1

I left Kansas City in the morning with two goals: 1) look for Acmaeodera robigo (family Buprestidae) in Texas Co., Oklahoma (based on a 2011 observation posted to BugGuide), and 2) make it to Black Mesa State Park in the far northwestern corner of the Oklahoma panhandle before sunset (to avoid having to set up camp in the dark). It was a long day of driving (mostly through heavy rain), but by the time I arrived at the A. robigo locality, skies were clear and the area was dry. Texas Co. is the largest (and seemingly flattest!) of the three Oklahoma panhandle counties, but I have not collected in the county previously because most of it has been converted to cultivated wheat. This particular location, however, lies within the Beaver River drainage and, thus, features native shortgrass prairie vegetation.

Beaver River drainage where Acmaeodera robigo was observed in 2011.

The host plant mentioned in the A. robigo observation, Melampodium lecanthemum (blackfoot daisy—family Asteraceae), was blooming abundantly along the roadsides, so I checked the flowers carefully up one side of the highway and down the other side.

Melampodium lecanthemum (blackfoot daisy).

Nothing was seen on the flowers, but given the lateness of the hour and cool temps I did not expect to see anything. Still, I set three white bottle traps in the area within patches of M. leucanthemum and will plan on retrieving them up later this season. With luck, the traps will have attracted adults if the species as they visit the nearby flowers.

Bottle trap, antifreeze, and a spade are all that is needed to trap flower-visiting insects.

Moving west I quickly entered Cimarron Co.—I always enjoy seeing this double-line of utility poles stretching to a seemingly endless horizon at the border between Texas and Cimmaron Counties. The latter is not only the westernmost county in the state, it is also the only county in the U.S. that borders four other states (Texas, New Mexico, Colorado, and Kansas).

Entering Cimmaron Co., the westernmost county in Oklahoma.

I made it to Black Mesa State Park with plenty of daylight left to set up camp. Per usual, after getting camp setup I enjoyed “refreshments” and waited for nightfall to walk the roads looking for nocturnal beetles (or any other critters that might be out and about). I did see an Eleodes sp. (darkling beetle—family Tenebrionidae) early in, but the night was chilly, and the onset of light drizzle put an end to the walk.

Ready for bed!

This would not be the end of my nighttime hiking, however—at 3am I was awakened by the drone of a motor, and it was loud enough that sleep would be impossible. Thinking it was an inconsiderate camper down at the RV campground, I decided to investigate and discovered it was a water pump within the campground. I decided if I couldn’t sleep, then somebody else would also have to be awakened, namely the campground host. It took the elderly woman a few minutes to come to the door after I rapped a few times on her RV, and I apologized for waking her before explaining the situation. She said she wouldn’t know anything about how to turn it off, to which I replied that I didn’t expect her to know that but I’ll bet somebody does and maybe she could make a call and figure out who that is or this wouldn’t be the last time somebody wakes her at 3am. I figured that would be the last of it, that I’d be awake until morning, and that I would have another nine hours of driving the next day on only three hours of sleep. But, to my relief, about a half hour later a park maintenance vehicle pulled up to the pump house, and a young man turned the motor off (I thanked him profusely). I did eventually get back to sleep and awoke to birdsong and clearing skies.


Day 2

There is a low water bridge at the south entrance to the park that crosses a normally dry creek. However, this time the creek was filled with water with lush growth of sedges and grasses along the water’s edge. This is a perfect situation for Taphrocerus species (family Buprestidae), and at this literal transition point from the eastern to the western fauna I’m not sure what species I might encounter there. Unfortunately, thorough sweeping of the sedges produced no beetles, so the question will for now remain unanswered.

Creek at south entrance to Black Mesa State Park, Oklahoma.

Heading south and then west from the park takes you shortly into the northeastern reaches of New Mexico and the town of Clayton. Whenever I pass through Clayton I like to stop at Mock’s Crossroads Coffee Mill for a bag of freshly ground coffee.

Mock’s Crossroads Coffee Mill, Clayton, New Mexico.

Skimming across the northern reaches of New Mexico for the next several hours was uneventful with no indication of a repeat of the heavy rain that pounded me for much of my drive yesterday across Kansas and Oklahoma. The situation changed, however, as I crossed the Continental Divide on the approach to Farmington.

I think I’m going to get wet.
I’m sure I’m going to get wet.

Sure enough, I once again had the pleasure of driving through driving rain. In addition, I arrived at my next planned spot just north of town (Brown Springs Campground) to find the entrance and main road under several inches of water. No point in trying to collect there, so I continued on towards the next stop (Devils Canyon Campground) in southeast Utah (San Juan Co.) in hopes of arriving early enough to set up camp before dark. Things looked promising as I crossed through the southwestern corner of Colorado towards Monticello, Utah, as patchy clouds filled a sun-filled sky.

Near Dove Creek, Colorado.

The presence of large oval emergence holes in the lower part of the limb confirmed that this tree was, indeed, under attack by such, and stripping the bark around the holes revealed fresh workings from the larvae indicating that some of them may still be inside the tree.

By the time I reached the campground, however, the rain had returned—not hard, but steady. I didn’t relish the thought of setting up camp in the rain and did something I almost never do—heading back into town and spending the night in a motel! At least I was able to enjoy a burger cooked by someone else for a change.


Day 3

Day 3 started off gray and overcast with the threat of more rain, but looking at radar and the areas I was going further west had me optimistic that I would get out of the rain once and for all. My ultimate stop for the day would be Jacob Lake Campground in north-central Arizona (Cocconino Co.), and my plan was to stop at a number of places along the way. Devils Canyon Campground itself, however, was still under clouds, meaning that collecting there would likely not be very productive. Nevertheless, despite last night’s rainout and the continuing cool, wet conditions, I decided to place some traps that I could pick up later in the season and, thus, get at least a sample of the area’s beetle fauna.

Devils Canyon Campground, Utah.

The campground lies at ~7000 ft, and I found a ridge with ponderosa pine/gamble oak forest on the slope to one side and pinyon/juniper/oak woodland on the crest of the ridge to the other. On the ponderosa side I set a white bottle trap, and on the pinyon/juniper side I set a sweet red wine-baited jug trap. There wasn’t much in bloom at the time other than a few yellow composites and the occasional Oenothera cespitosa (tufted evening primrose—family Onagraceae) with blossoms still wide open despite the daylight hour due to the overcast, cool conditions.

Oenothera cespitosa (tufted evening primrose).

Coming down off the plateau towards Bluff, Utah, thick low clouds gave way to higher broken clouds with warmer temps (mid-60s). I stopped a few miles west of town at a location where Mont Cazier once collected what was later described as Agrilus utahensis (family Buprestidae) and began sweeping roadside vegetation. While I was sweeping, I encountered three Amannus vittiger (family Cerambycidae) on the flowers of Sphaeralcea coccinea (scarlet globemallow—family Malvaceae) (along with a few meloids and bees for others). The record of A. utahensis was from quite a bit later in the season (late July), so I didn’t really expect to find it (although I was hopeful). However, I did sweep what appears to be Agrilus malvastri.

Amannus vittiger collected on flowers of Sphaeralcea coccinea (scarlet globemallow).
Sphaeralcea coccinea (scarlet globemallow).

A short distance west of the stop had me entering northeastern Arizona, and traveling west of Dennehotso on Hwy 160 I encountered patchy, low lying clouds though which beams of bright sunlight lit up the red rocks below and cast a reddish hue on the bottoms of the clouds.

West of Dennehotso, Arizona on Hwy 160.

A bit further west on Hwy 160 is a formation apparently known as “Baby Rocks.”

Baby Rocks, west of Dennehotso, Arizona,

As I drove, I kept my eye out for areas along the highway with stands of plants in bloom, and I found just such at a roadcut on Hwy 98 near mm 325 as I approached Page, Arizona (Cocconino Co.). There was a fair amount of Sphaeralcea coccinea in bloom, off which I collected a handful of Acmaeodera pubiventris lanata and one Amannus vittiger from the flowers. I swept the grasses and other roadside vegetation thoroughly, but that produced nothing. I returned my attention to the Sphaeralcea and swept it thoroughly as well to be sure I didn’t miss anything. Good thin I did, as that turned up one more each of A. p. lanata and A. vittiger as well as a few Agrilus sp. prob. malvastri.

Acmaeodera pubiventris lanata on flower of Sphaeralcea coccinea (scarlet globemallow).

After passing through Page, Arizona, rather than continuing straight west to Jacob Lake Campground, I veered north into Kane Co., Utah to revisit a locality I’d visited a couple of years ago about 23 mi northwest of Page on Hwy 89. The last time I stopped here (late June 2023), I found a few Nanularia brunneata (family Buprestidae) on the stems of Eriogonum inflatum (desert trumpet—family Polygonaceae). That visit was about three weeks later in the season than this time, and not only was the area dry (despite all the rain east of here), but there was very little new growth on any of the E. inflatum plants—just the dried stalks of last year’s growth. Still, there was fresh growth present at the spot, in the form of Ericameria nauseosa (gray rabbitbrush—family Asteraceae), on which I collected a single cryptocephaline leaf beetle, and along the roadside rain shadow in the form of Sphaeralcea parvifolia (small-leaved globemallow), on the flowers of which I collected several more Agrilus sp. prob. malvastri. Eventually, I did find a single E. inflatum plant with small sprouts of new growth on which I found several clytrine leaf beetles. Sweeping the S. parvifolia produced one more Agrilus sp. prob. malvastri, a single Acmaeodera navajoi—a species I collected here abundantly two years earlier, and a series of a very small chrysomelid leaf beetle species that I observed feeding on the foliage.

Once I got a bit further west, the landscape turned green again, indicating recent rains, so I checked iNaturalist for nearby records of E. inflatum that I could check for N. brunnea. There was one a few miles east of Kanab (Kane Co.) and on the way to Jacob Lake Campground, so I stopped by to see if I could find the plants. Unfortunately, the record turned out to be on private property, so I couldn’t go to the precise location. I looked around the area outside the property but didn’t see any plants; however, S. parviflora was blooming abundantly along the roadsides, and sweeping it produced a few more Agrilus sp. prob. malvastri and a couple of small bees (for Mike).

11 mi E of Kanab, Utah.

There were also several blooming Opuntia polyacantha var. erinacea (Mojave pricklypear—family Cactaceae), which I checked for Acmaeodera, but I found only one large bee (again, for Mike). With the sun by then dropping close to the horizon, I called it a day and headed for the campground.

Opuntia polyacantha var. erinacea (Mojave pricklypear)—pink-flowered form.

I arrived at Jacob Lake Campground (Cocconino Co., Arizona) and got camp setup before dark, which settled in as I was cooking dinner. It was chilly at this relatively high elevation (7900 ft)—temps were already in the mid-50s and were forecast to drop down into the 40s by morning. Despite the chill, I found a Zopherus uteanus (family Zopheridae) and a smaller tenebrionid on a cut stump of Pinus ponderosa (ponderosa pine).

Zopherus uteanus on cut stump of Pinus ponderosa (ponderosa pine) at night.

Later, I walked the site—not really expecting to see much because of the chill, but I did find three Iphthiminus lewisii (family Tenebrionidae) on the trunk of a recently fallen P. ponderosa and saw two more on a pine stump.

Iphthiminus lewisii on trunk of recently fallen Pinus ponderosa (ponderosa pine) at night.

With three days of travel—the first two mostly rainy and today dry but still quite cool, I’m hoping the forecasted warmer, sunnier conditions for tomorrow and onward come true and I can get this collecting trip in high gear.

A waxing gibbous moon illuminates my camp at 7900’ in Jacob Lake Campground, Kaibab National Forest, Arizona.

Day 4

After morning coffee (and moving to a different campsite—long story!), I went back north a bit to the area near Lefevre Overlook that had been only lightly impacted by the widespread burns that occurred here a few years ago and is showing signs of recovery. It felt “early season” with patchy clouds, moderate temps in the low of mid 70s, and plenty of moisture in the soil with lots of plants in bloom. Insects were not abundant, but with continued beating I collected a couple of Anthaxia sp. (family Buprestidae) and a variety of small clytrines, cryptocepahlines, and curculionids off of Purshia standsburyana (Standsbury’s cliffrose—family Rosaceae) in flower.

Purshia standsburyana (Standsbury’s cliffrose).

As the morning warmed, insects seemed to become a bit more active, although the day continued to feel “early season.” Persistent visual searches and beating of a variety of plants turned up a few small black/yellow Pidonia? (family Cerambycidae), one Acmaeodera diffusa?, and one Anthaxia sp. on flowers of Hymenopappus filifolius (fine-leaved hymenopappus—family Asteraceae); one Agrilus sp. prob. malvastri, another Pidonia?, and one A. diffusa? on flowers of Sphaeralcea ambigua (desert globemallow); several small clytrine and cryptocephaline leaf beetles and curculionid weevils on Quercus gambelii (Gambel oak—family Fagaceae); and several A. diffusa? and a few Anthaxia sp. on flowers of Tetraneuris acaulis (four-nerve daisy—family Asteraceae). Sweeping through the area where the T. acaulis was growing produced nothing further of interest, so I placed a sweet red wine-baited jug trap and a yellow bottle trap in the area—hopefully they will each attract a variety of interesting beetles over the next few months.

Sweet red wine-baited jug trap hanging in a tree and a yellow bottle trap sunk into the ground below.
Phrynosoma hernandesi (greater short-horned lizard).

A bit further down, the mountain escaped the fire that ravaged many other parts of the mountain. I expected to see much more insect activity, but the opposite instead was the case. This may be related to the lower incidence and diversity of flowering plants, which were likely triggered at the previous spot by the fire. I did find a small area where Sphaeralcea ambigua was blooming abundantly and sweeping through them produced a fine series of Agrilus sp. prob. malvastri along with a few bees (for Mike). I walked up the slope a ways, but there was virtually nothing in bloom—unlike what I had seen at the previous spot. Eventually, I did find a few Hymenopappus filifolia and Tetraneuris acaulis, but the only I beetle I saw on any of them was a single Acmaeodera diffusa? on the latter.

Escobaria vivipara (viviparous foxtail cactus).

Without anything luring me further up the slope, I worked my way back down towards the car and was about ready to call it a day when I decided to take a gander a bit down the slope—just in case I might see something of interest. Nothing captured my attention, however, so I worked my way over a bit to take a different route back. That’s when I saw it! All day long I had been keeping my eye out for large Utah junipers (Juniperus osteosperma—family Cupressaceae) with thigh-sized main limbs—one of which was dying (not dead), possibly a result of woodboring beetles, and before me was just such a tree!

Juniperus osteosperma (Utah juniper) with dying main limb.

The presence of large oval emergence holes in the lower part of the limb confirmed that this tree was, indeed, under attack by such, and stripping the bark around the holes revealed fresh workings from the larvae indicating that some of them may still be inside the tree. 

Large oval emergence hole caused by a species of longhorned beetle.
Fresh frass packed in the galleries by longhorned beetle larvae.

I hiked back to the car to get my chainsaw—brought along on the trip just for such an eventuality—and cut a portion of the infested limb out of the tree so I could bring it home and attempt to rear out the adults. My hope is that the species infesting the tree is Semanotus juniperi (family Cerambycidae), a very uncommonly encountered longhorned beetle that breeds in the large limbs of juniper in this area, so I’ll be keeping my fingers crossed. Hauling out the infested bolt along with the chainsaw was a struggle, but eventually I made it back to the car and celebrated a successful last act to what had up to that point been a not too spectacular first full day in the field.

Bolt of Juniperus osteosperma (Utah juniper) infested with longhorned beetles.

I needed a break after working the chainsaw and hauling out the wood, so I went back to the campsite to enjoy an end-of-the-day “beverage” and eventually dinner. However, part of me was thinking Inshould have collected more of that infested juniper to increase my odds of successfully rearing whatever was infesting it, so I resolved to do exactly that in the morning.

Dinner on the grill at Jacob Lake Campground.

Before dinner as I was gathering kindling, I had noticed a large, fallen ponderosa pine behind the campsite and returned after dark to see if there might be any nocturnal beetles attracted to it. It seemed recently dead, but as I inspected the upper branches more closely I saw that it had been dead long enough that whatever would have been attracted to it had already come and gone. I thought maybe I could find beetles under the bark and began peeling the bark in the trunk. The bark was still intact but was just loose enough that I could peel it off in large sheets. Doing so, however, revealed only a couple of click beetles (family Elateridae). I didn’t see any other fallen or dead trees in the area, but on the trunks of large living ponderosa pines I found a tenebrionoid and several individuals of a Lecontia discicollis (burnt conifer bark beetle (family Boridae). [Edit: thanks to Alex Harmen for the identification on this one!] Nearby was a cut stump of a very large ponderosa pine, and peeling back the thick bark at the base of the stump revealed yet another elaterid species and two darkling beetles. I was hoping to find more Zopherus uteanus, but no such luck!

Lecontia discicollis (burnt conifer bark beetle) on trunk of Pinus ponderosa (ponderosa pine) at night.

Day 5

After a morning of the coffee ritual and observing campsite wildlife, I made an ice run at the nearby service station.

Callospermophilus lateralis (golden-mantled ground squirrel).
Lophocampa ingens (family Erebidae) larva.
Meanwhile, a century later…

I was having second thoughts about taking only a single bolt of wood from cerambycid-infested Utah juniper that I found at the end of yesterday’s collecting and decided to go back to get more of the infested wood to improve the likelihood of successfully rearing adults from the wood. It would not be an easy job—I’d have to haul the chainsaw down the the tree, cut off the two remaining infested bolts, cut the bolts down to cartable pieces, and then haul the chainsaw and wood bolts back to the car, likely requiring several trips. It was worth it, however, because as I was stripping bark to see where I needed to make the initial cut, I encountered an intact cadaver of one of the beetles that had died while trying to emerge from the wood. I carefully extracted the abdomen from the tree and found the head and pronotum in the bark and placed them in a vial, and they without a doubt represent the species I was hoping they were—Semanotus juniperi, a super rare species that very few people have ever collected [tip of the hat to Ron Alten for sharing his knowledge about this beetle with me and enabling me to find this beetle for myself!]. I’ll be able to put the beetle back together when I get home, and I’m hopeful I’ll rear at least a small series of beetles from the wood I’ve collected.

I love the smell of chainsaw in the morning!

Having carted the additional bolts of wood back to the car and confident in my ability to recognize the work of this species, I looked for—and found—a few more trees that the beetle had infested. However, in each case the workings were old and no beetles were encountered chopping into the wood. I also spent some time looking for much smaller emergence holes in the thick trunk bark of several junipers and carefully shaving the bark to see if I could find another rare longhorned beetle species, Atimia vandykei. I found several galleries that might have been this species, but no beetles or larvae were encountered, leading me to think I might have been a bit late and the adults have mostly emerged. A large, recently dead Pinus edulis (Colorado pinyon pine) also caught my attention, as it looked fresh enough to still contain wood boring beetles that might have infested it. The wood was very hard and difficult to chop into, but I didn’t encounter any larvae of any kind and elected not to collect a bolt or two for rearing. Lastly, I spent some time looking for dead Gambel oaks, which in this area could host Xylotrechus rainei (family Cerambycidae), a recently described species that few have collected. I found several dead stems of the shrubby oak species that contained workings consistent with those of Xylotrechus, but in each case the workings were old and no beetles or larvae were present. This string of “failures” might have seemed like the makings of a bad day, but the success with S. juniperi overshadowed those failures and I left the site happy (though exhausted!).

The author sports his new field hat (an early Fathers Day gift!).

I returned to the first stop of yesterday to see if I could find junipers in this area infested with S. juniperi, reasoning that since the area had burned a few years ago some of the still-living trees might be stressed, thus making them more vulnerable to infestation. I also wanted to see if the bottle and jug traps I placed yesterday had caught anything of interest (even though it had only been one day). Along the way I picked up a single Anthaxia sp. on the flower of Tetraneuris acaulis and a few Acmaeodera diffusa? on flowers of Sphaeralcea ambigua. There wasn’t anything of interest in either trap, so I set about looking for infested junipers. I found only one, but again the workings were old and no larvae or adults were encountered. I also examined a few dead stems of Gambel oak, but none showed signs of infestation. (I suspect they had been killed by the fire and the damage to the wood by the fire made the stems unsuitable for infestation.) I did, however, find a small more recently dead Colorado pinyon pine that showed signs of recent infestation all along the trunk and collected it for rearing. By then, the day had warmed considerably and I was already exhausted from the morning’s chainsaw session, leading to a loss of motivation to keep looking. I needed a change of pace and decided to head higher up the mountain back into the ponderosa pine forest for some more “traditional” woodboring beetle collecting.

Callophrys gryneus (juniper hairstreak) on flowers of Hymenopappus filifolia (fine-leaved hymenopappus).

The forest at this site is dominated by ponderosa pine, and the strategy here was straightforward—look for large, dead or dying trees, either standing or recently wind-thrown, and inspect the examine the trunks for woodboring beetles. I came to this spot two years ago in early July, so this time was about four weeks earlier in the season. I had two specific targets (beyond woodboring beetles in general)—Chalcophora (family Buprestidae) and Monochamus (family Cerambycidae). Both of these genera are the subject of molecular studies being conducted by other researchers that I know, and I’ve been promising to send them fresh specimens killed and preserved in ethanol. The first wind-thrown tree I encountered was still green-needled, and I expected to find buprestid beetles all over it. Instead, all I saw were a few small Enoclerus sp. (family Cleridae). Large dead standing trees dotted the open forest, and I carefully approached and circled each one looking for beetles, paying special attention to those in full sunlight. Time after time, however, I was frustrated. Each time, I chopped into the wood a bit to examine what was going on underneath the bark, and while workings were plentiful I never encountered any larvae.


4.2 mi N Jacob Lake on Hwy 89A, Kaibab National Forest, Arizona.

Finally, I approached a large standing dead tree, and perched on its trunk in the sunlight was Chalcophora angulicollis (western sculptured pine borer). I’m happy to give this specimen up for DNA sequencing, especially since I already collected a specimen at this same spot two years ago (before the molecular study began).

Chalcophora angulicollis (western sculptured pine borer) on trunk of dead Pinus ponderosa (ponderosa pine).

I peeled a good portion of bark of the tree it was perched on but found nothing except a large elaterid larva (I wonder if it was Alaus melanops [western eyed elater], a predator or woodboring beetle larvae).

Alaus melanops (western eyed elater) collected from under bark of dead Pinus ponderosa (ponderosa pine).

For a long time afterwards that would remain the only beetle I encountered until I found a large standing tree in the early stages of death (needles pale green but not brown) with a female Dicerca tenebrosa (family Buprestidae) searching the trunk, occasionally stopping to probe a crack or crevice with her ovipositor. Patiently waiting at the trunk rewarded me with a second individual within a short period of time.

Dicerca tenebrosa on trunk of dead Pinus ponderosa (ponderosa pine).

Another long period of nothingness ensued as I zigzagged from dead tree to dead tree, eventually retracing my steps to the only two trees on which I had found buprestids. Even that failed to produce additional beetles, but as I was standing at one of them I heard a woodpecker persistently pecking and searching and pecking on a nearby tree. The tree looked perfectly healthy at first glance, but I figured the woodpecker had to know something that I didn’t (I’m always willing to learn from locals!) and walked over to the tree. It was immense, and a closer look at the crown revealed a few browning needles scattered throughout the crown. Then a closer look at the trunk revealed several of the same small Enoclerus sp. I saw earlier crawling on it. Woodpeckers and checkered beetles don’t lie, and this tree was clearly under stress and under attack by woodboring beetles. An initial circling of the trunk revealed no other beetles, but then I noticed the gangly antennae and legs of a male/female pair of Monochamus clamator, the male apparently mate-guarding the female. I’ll be happy to contribute one of these specimens to my colleague for DNA sequencing and keep the other for my collection.

Monochamus clamator on trunk of stressed Pinus ponderosa (ponderosa pine).

By that time, the sun was starting to get rather low in the horizon and I was utterly exhausted after a full day of walking, chopping, chainsawing, and hauling. I passed by the two previously successful trees on the way back to the car, to no avail, and headed back to camp (stopping at the nearby market for a celebratory milkshake before going into the campground).

Caenochrysis sp. (family Chrysididae) on trunk of dead Pinus ponderosa (ponderosa pine).

Day 6

The day’s plan was to head north to the area around Coral Pink Sand Dunes system in southern Utah, but before I left I decided to go down the east slope of the Kaibab Plateau a bit to see it I could find a good transition zone (one that hadn’t burned) from the higher elevation ponderosa pine forest to the juniper/pinyon woodland just below. A few miles east of Jacob Lake, right as I hit 7000’ (there was even a sign to that effect) while descending into the canyon, I saw a small pulloff with ponderosa pine forest (and even some fir) on the east-facing slope to the right and juniper/pinyon/oak woodland on the west-facing slope to the left—perfect! It struck me as a good-looking spot to set some traps, so I set a blue bottle trap in an open area on the juniper/pinyon/oak side of the highway and hid a sweet red wine-baited jug trap in the woodland right above it. I then started beating the patches of Gamble oak hoping for Brachys (family Buprestidae) but finding only a single Anthaxia sp. and several species of clytrine leaf beetles and curculionid weevils. There was a patch of Sphaeralcea ambigua in flower near the pulloff, off which I collected several Acmaeodera diffusa? plus another of those small black /yellow lepturines (Pidonia? sp.) that I found yesterday, and I swept some chunky black dermestids and a couple of bees (for Mike) from the flowers of Hymenopappus filifolia. Next came a long period of nothingness! I found another Utah juniper with damage on the main trunk by Semanotus juniperi, but once again it was old with no larvae or adults encountered after another exhausting chopping session. At this point, I turned my attention to the ponderosa pine forest across the highway, as I’d noticed some large dying/dead trees here and there that I wanted to check for buprestids. Despite checking every dead or declining pine within eyeshot, I didn’t find a single beetle! This, combined with the earlier fruitless chopping session, sapped my motivation, and I started heading back towards the car. Only a few A. diffusa? on the flowers of Opuntia polyacantha var. erinacea and Heterotheca hirsutissima (harsh false goldenaster—family Asteraceae) momentarily captured my attention on the way back.

Opuntia polyacantha var. erinacea (Mojave pricklypear)—yellow-flowered form.
Heterotheca hirsutissima (harsh false goldenaster).

As I neared the car, I saw a nicely blooming Purshia stansburyana and beat a nice series of Anthaxia (Haplanthaxia) caseyi from it. I’m not sure if the population in this area is assignable to any of the currently recognized subspecies, so I’ll be interested to study them closer and compare them to other unassignable populations I’ve found in other parts of Utah and Arizona. Right before I reached the car, I saw another juniper that begged “chop me.” I complied and then hated myself for it, as the result was the same as all the other junipers I’ve chopped into since finding that first one with Semanotus juniperi in it. With that, I said goodbye to the Kaibab Plateau and headed north towards Ponderosa Grove Campground near Coral Pink Sand Dunes north of Kanab, Utah.

Clytrine leaf beetle on flower of Purshia stansburyana (Stansbury’s cliffrose).

I reached the area around Coral Pink Sand Dunes (Kane Co., Utah) by mid-afternoon. My favorite campground in that area is Ponderosa Grove Campground—it’s large, spacious campsites are not only well shaded by the namesake, unique-for-the-area grove of massive ponderosa pines, but it is also located right across the road from Moquith Mountain Wilderness Study Area, a BLM-managed area with easy access to the northern portion of the Coral Pink Sand Dunes system (for those who may be asking, this portion of the dune system lies outside the boundaries of Coral Pink Sand Dunes State Park, which protects the only known habitat of the highly vulnerable Coral Pink Sand Dunes tiger beetle, Cicindela albissima).

After picking a campsite (I was glad to see only a few sites occupied) and unloading my gear, I hightailed it to the dunes with one objective—find Chrysobothris nelsoni (family Buprestidae). I collected this species before on my previous trip here back in late June of 2023, but I spent a day and a half looking for it and managed to only a small handful of specimens. I thought the earlier timing might be better, but Norm Woodley was here a couple of weeks before and did not see the species, giving me reason to be also skeptical.

Looking across the dunes at Moquith Mountain Wilderness Study Area.

On the way to the dunes, I set out a yellow bottle trap and sweet red wine-baited jug trap in the sandy juniper/pine woodland bordering the dune. Upon entering the dunes, I immediately started seeing the host plant, Eriogonum alatum (winged buckwheat—family Polygonaceae), a distinctive plant with a basal rosette of linear leaves and, on some plants, a tall stem bearing the inflorescence. One plant in particular, right at the dune entrance, spoke to me saying “look at me.” I don’t know why, but I went over to it, tapped the basal rosette over my net, and off fell a nice large female C. nelsoni! Well, that was fast. I looked at a couple more plants and saw on a second adult (this one a smaller male) sitting head down at the base of the inflorescence stem—two specimens on the first three plants I looked at!

Chrysobothris nelsoni on basal rosette of Eriogonum alatum (winged buckwheat).
Chrysobothris nelsoni on basal rosette of Eriogonum alatum (winged buckwheat).
Chrysobothris nelsoni on stem base of Eriogonum alatum (winged buckwheat).

I found quite a few more (although not quite at that same frequency) over the next hour or so before they seemingly just disappeared. I noted the lateness of the hour and wondered if they have a ‘bedtime’—perhaps they burrow into the sand around the base of the plants for the night—and started back towards camp, picking up a few Eleodes caudifera lumbering across the surface of the sand along the way. I decided at that point, now that I had a nice series of the beetle, to bring the “big camera” over the next morning and try to get much nicer photos of the beetles on their host plant that what I can achieve with this iPhone.

Eleodes caudifera lumbers across the sand.

It had been a long day by that point—I was both famished and exhausted and needed a bit of time to rest and refuel. I had brought two salmon filets along with me, which should have been enough for two meals, but I was so ravenously hungry that I cooked and ate both.

Chill time at Ponderosa Grove Campground.

The rest and food gave me a bit of a spark, and as sunset approached I was inspired to set up the blacklights. Conditions were not close to ideal—a waxing gibbous, nearly full moon along with cool(ish) temperatures are usually enough to kill blacklighting. However, it was warmer than the past several nights at Jacob Lake (1300 ft higher elevation), so I compromised by setting up only the two ultraviolet lights (which I can run right off the car batter) but not the mercury-vapor lamp (which would have required hauling out and running the generator).

Not long after I turned the lights on, I noticed the bright, unmistakable glow of a mercury-vapor lamp at the far other end of the campground. I was like “That has to be an entomologist!” so hiked on down to introduce myself. As I entered the campsite, a man approached me and said “Hello, Ted.” Now I’m thinking okay we’ve met before, but I’m a dummy with poor social skills because I don’t recognize him. He said “My name is Mike.” The omission of his last name had me doubly thinking that I was an idiot because the mention of his name still wasn’t enough to trigger my memory of his last name or who he even was. When I asked him his last name, he said Rashko. I recognized that name instantly—he is the person who discovered Acmaeodera rashkoi, recently described by Rick Westcott and one of the species I had targeted on this trip. I told him of my plan and asked him how he knew who I was. It turns out he is a longtime follower of this blog and had seen my license plate (MOBUGS) earlier in the day. He was traveling from his home in Oregon to Flagstaff to meet his family and stopped here to spend the night and see what he could collect. We had a wonderful time chatting about the art of collecting and about colleagues we know (especially Rick, whose ears were burning I’m sure). I showed him the nice series of C. nelsoni that I’d gotten earlier in the day and told him we could go back in the morning before he left so that he could get some. We got so involved talking that we forgot to even look at his sheet to see what insects had come in until a Polyphylla uteanus (Coral Pink Dunes June beetle—family Scarabaeidae) smacked into his head and bumbled its way over to the sheet (Mike let me keep it). A large female Monochamus clamator also landed on the sheet (which Mike kept).

Eventually we said our goodbyes, and I wandered back to my campsite where I was mildly optimistic that my ultraviolet lights had brought in something. Sadly, there were no beetles of interest, but there was a stunning Hyalophora gloveri (Glover’s silk moth—family Saturniidae) and eventually two small white sphingid moths—all of which I kept for Rich Thoma back home. Success for me that night, however, was not yet out of sight. I made a round to inspect the massive trunks of the ponderosa pines dotting the campground and was rewarded with a nice little series of Zopherus uteanus.

Zopherus uteanus on trunk of Pinus ponderosa (ponderosa pine) at night.

I also found a few of the weirdly explanate Embaphion glabrum (family Tenebrionidae) crawling on the ground beneath the trees. A second round to look at the trees yielded no additional beetles, however, so I turned the lights off and turned in.

The “Big Dipper” prominently graces the western sky during the wee morning hours.

Day 8

Just as I had promised the night before, I wandered down to Mike’s campsite in the morning to see if he wanted to look for Chrysobothris nelsoni. He was already out looking for “beetles in the bush” around his campsite, and together we worked our way over to the dunes. It didn’t take long for me to see one of the beetles in its host plant, which I pointed out to Mike so he could see it and try to catch it himself. That beetle got away, but almost immediately he saw another one on a neighboring plant and succeeded in capturing it. We saw several more, but unlike the previous evening they were faster and more difficult to catch with the rising late-morning temps. Mike caught a couple more (and I one for the record) but had to leave, so we exchanged contact info and said our goodbyes.

Morning on the dunes.
The author (left) with Mike Raschko (right).

Back at the campsite, I looked around a bit to see what else I might collect. There was a nice diversity of plants in bloom, many which could be expected to be visited by buprestid beetles—Sphaeralcea parviflora (small-leaf globemallow), Opuntia aurea (golden pricklypear cactus), Hymenopappus filifolia, etc., but no beetles were seen. There were clumps of Gamble oak and Amelanchier utahensis (Utah serviceberry—family Rosaceae), but beating them produced nothing. The only thing I found was a recently windthrown branch of Utah juniper, which, when I stripped back some of the bark, proved to be in the early stages of infestation by longhorned beetles, probably a Callidium sp. The lack of insect activity (except for C. nelsoni) presented a quandary—should I stay another night (as planned), or should I pack up and head to my next destination (either Leeds Canyon, Utah or Kyle Canyon, Nevada)? I decided it was already too late in the day to head somewhere else—by the time I got there it would be late in the day, and I didn’t relish the idea of searching for an available campsite late with no reservation at such a late hour. I decided I might as well stay put and make the best of it—which I could do by going back over to the dunes and look for more C. nelsoni. The day’s heat, however, was not only making the beetles very difficult to catch but also starting to get to me. Fortunately, distant thunderclouds came closer and closer until they were directly overhead. Rather than rain, however, it was virga, so I got the best of both worlds—an immediate cooling off that was not only comfortable for me but also settled the beetles down without the rain that would have sent me scurrying back to camp.

Virga brings relief from the heat to the dunes.

In the end, I succeeded in collecting another nice series of specimens.

Chrysobothris nelsoni on basal rosette of Eriogonum alatum (winged buckwheat).

By then it was mid-afternoon and I was famished, so I headed back to camp to refresh, refuel, rehydrate, and catch up on my field notes. Comically, as I was writing my notes, I happened to be watching one of the common sagebrush lizards (Sceloporus graciosus) that had been hanging around the campsite as it bit at and then rejected some type of insect that was crawling on the logpile next to the firepit. I got up to see what it had rejected, and it turned out to be Danosoma brevicorne (family Elateridae), which I am happy to add to the collection.

Common sagebrush lizard (Sceloporus graciosus).
Danosoma brevicorne on woodpile.

After some chill time at the campsite [it was actually very busy—I wrote my field notes from earlier, charged all my devices using an inverter hooked up to the car battery, and downloaded photos from my “big camera” memory card to the computer], I went back to the dunes to see if I could get “big camera” photo of the beetles. It was touch-and-go at the start—shortly after reaching the dunes it started sprinkling and the wind started picking up. All I could do was wait it out, hope conditions improved, and be ready to bolt if the skies opened up. Just as quickly as it started up, however, it blew over, and I was able to start looking for beetles. As with every other time during this visit, it didn’t take long. The first beetle I found was a bit hidden within the crown of the plant, but I was able to carefully move the leaves out of the way without disturbing the beetle and got a nice series of shots. I was happy with the photos, but I wanted photos that were a little less “cluttered.” The perfect opportunity for such arose when I saw a beetle sitting out near the tip of a leaf. It was not in the best position—other leaves were partially blocking it from view, so I carefully grabbed the leaf at the base and gently pulled until it detached from the plant. Fortunately, the beetle wasn’t phased by the tugging and continued to sit calmly on the leaf. I wanted a blurred pink sand background, which I thought would look spectacular behind the brilliant green color of the beetle, so I stuck the base of the leaf in the sand to prop it up and adjusted the angle and distance in concert with my camera and flash settings until I achieved the desired effect. I’m super happy with how some of these photographs turned out, so look for them to appear in a future post. I’m probably lucky that the mini-storm moved through when it did, as the cool conditions likely calmed the beetles down and made them more willing subjects. By the time I finished photographing the second beetle, the sinking sun signaled a dinner bell, and I walked back to the campground super satisfied with how this visit had turned out.

After dinner, I turned the ultraviolet lights on despite the just-shy-of-full moon and customary coolish temperatures. I didn’t expect anything to show up, but you don’t know if you don’t try (nothing ever showed up!).

Mustard makes everything fancy in the field.

While waiting for nothing to come to the lights, I made several rounds of the large ponderosa pines, expanding the circuit to include some further toward the west end of the campground. As with the night before, I found a half dozen Zopherus uteanus crawling on the trunks (one was up too high to reach, though, and when I knocked it down with a stick it disappeared into the thick vegetation below the tree). I’d really like to know what these guys are up to during the day (hiding in the leaf litter at the base of the trees?) and what the larvae are doing (no idea!), but they certainly seem to be associated with pines (I’ve also collected Zopherus concolor at night on the trunks of Colorado pinyon pine). I also found a few more tenebrionid darkling beetles, including another Embaphion glabrum, crawling on the sandy ground beneath the trees. I expanded my search to include the trunks of some of the large Utah junipers, finding a few more tenebrionids on them.

Embaphion glabrum in ponderosa pine grove at night.

With midnight approaching, I soaked in my last bit of experience at the place before retiring for the night and leaving the next morning.

A waxing gibbous, almost full moon rises high above the campground.

Day 9

I left Ponderosa Grove Campground about 9:30 am with the plan to spend the next couple of days in Kyle Canyon and Lee Canyon northwest of Las Vegas. The drop down from the Colorado Plateau via the Virgin River Canyon was long the most spectacular stretches of freeway I’ve ever seen, but the sad reward waiting at the bottom was searing +100°F heat (the highest I saw registered on my car’s thermometer was 108°F!). Fortunately, turning off I-11 and heading west on Kyle Canyon Road gradually gained elevation. Before reaching the canyon proper, however, I had one of the trip’s top goals to take care of—setting a bottle trap for the recently described Acmaeodera raschkoi (recall that I encountered the namesake of this species, Mike Raschko, just two days prior at Ponderosa Grove Campground). I found the type locality no problem (where the modest gain in elevation had reduced the temperature to only 100°F!) and also found the trap that Mike had just set the day before we met.

Lower Kyle Canyon Rd—type locality of Acmaeodera raschkoi.

I placed yellow bottle trap about 59 m to the west from Mike’s trap and a blue bottle trap about 45 m to the east. There was no reason to stay at the locality and try to collect, as things were super dry, the only green vegetation seen besides the Yucca jaegeriana (eastern Joshua tree—family Asparagaceae) dotting the landscape was a small Cylindropuntia ramosissima (branched pencil colla—family Cactaceae).

Cylindropuntia ramosissima (branched pencil colla—family Cactaceae).

I had originally planned to camp lower down in the canyon at Kyle Canyon Campground, but by the time I got there it was already nearly filled to capacity. There were a couple of campsites still available, but the campground as a whole had a noisy, crowded vibe that wasn’t my cup of tea. I decided to take a chance and head further up the mountain to Hilltop Campground to see if I could find something more to my liking. It was a short drive up the mountain, and this campground also was nearly filled to capacity. However, it was a much quieter vibe, and among the three campsites still available was one that was isolated from all the others on the side of the mountain with spectacular views across the canyon and down into the desert below. Also, the 8300 ft of elevation offered much cooler conditions—a welcome change from the searing temperatures I had endured earlier in the day. It was perfect!

View from Hilltop Campground, Spring Mountains, Nevada.

I quickly set up camp and began looking around. Not far from the campsite I found a wind-thrown branch Utah juniper, and slicing into the bark I found a dead adult Semanotus sp. prob. caseyi amplus (not as exciting as S. juniperi, but still a nice find) and a small scolytid bark beetle boring an oviposition gallery. I did a bit of chopping into the bark and found small new cerambycid galleries (probably either Callidium or Semanotus) and some very large frass-filled galleries at the larger end that may be Semanotus juniperi. I found the frass-plugged entrance hole to the sapwood —a sign that the beetle had not yet emerged—and chipped away on either side of the gallery into the sapwood until I saw the large cerambycid larvae sitting it it. This confirms that the wood is actively infested, and I’ll cut it up and bring it home for rearing. At the empty campsite next to mine, I found a small, recently dead Abies concolor (white fir—family Pinaceae), and closer inspection of the trunk revealed an adult Dicerca tenebrosa tenebrosa, and on a return trip to the tree I saw another one (but too high up to capture ☹️). I also hung a red wine-baited jug trap just south of the campsite. There wasn’t much else going on in the area and it was getting late, so I got dinner started.

Sunset at Hilltop Campground.

I didn’t even consider setting up lights—the combination of a full moon and the cool, windy conditions that typify sites at this high of an elevation made the chance of success highly unlikely (in fact, it got so cool that I needed to pull on a fleece pullover). That did not, however, mean I could not do any night collecting—examining tree trunks has become a favorite strategy of mine that can be done on almost any night regardless of temperatures or wind. Trees under stress or recently dead (as well as recent wind throws or woodpiles) are especially good to look at, and I had noticed several during my earlier foray that I made a point to check. At the same time, I have also learned not to ignore living, seemingly healthy trees, as these can also harbor interesting beetles. Of course, the very first tree I headed for was the recently dead white fir on which I had collected Dicerca tenebrosa a few hours earlier, and I was rewarded right off the bat with an unusually small Zopherus uteanus. I wondered if this might be an interesting locality for the species, but I saw the species was recorded from Kyle Canyon in a review of the genus by Triplehorn (1972) (at which time it was still placed in the family Tenebrionidae). I quickly found another individual, this time on the trunk of a large Utah juniper—interestingly, the first non-pinaceous host on which I’ve found the species (although still a gymnosperm).

Zopherus uteanus on trunk of large Juniperus osteosperma (Utah juniper) at night.

What I was most anxious to check, however, was a large Pinus monophylla (single-leaf pinyon pine) that I had seen nearby with a long, twisting scar from a lightning strike gashing down the trunk—surely there would be something on that (there wasn’t despite checking it repeatedly for the next couple of hours). What did produce beetles, though, was a large, recently wind-thrown branch from that very tree laying on the ground nearby—two species and multiple specimens of each, one (Oeme? sp.) searching frenetically back and forth along the branch, and the other (Haplidus? sp.) in the form of a mating pair. I found it interesting that my inspection of the branch earlier during the day produced nothing.

Oeme? sp. on fallen branch of Pinus monophylla (single-leaf pinyon pine) at night.
Haplidus? sp. mating pair on fallen branch of Pinus monophylla (single-leaf pinyon pine) at night.

At this point, the night was already a reasonable success, but the greatest success lay just ahead. I had noticed a few large Utah junipers that showed evidence of infestation by Semanotus juniperi—or at least that what I presumed based on what I had learned about that species in the Kaibab Plateau a few days earlier. Not that I thought I would see adults of that species, but I figured the trees would have to be under some stress and might attract other beetles (I had already collected Z. uteanus off of one a few minutes prior). The first one I checked had nothing on it, but when I checked the second one I noticed a large black longhorned beetle on the trunk underneath some shredded bark. I carefully removed the overlying bark to get a better look at it and quickly realized that it was, indeed, S. juniperi! As I prepared to take a photograph, I noticed movement a bit higher on the trunk, and there sat a second individual! In the fraction of a second that followed, the memory of all the chainsaw work I had done a few days earlier in an effort to rear proper specimens flashed through my mind, yet here before me now were two live adults on their host in the wild. I presume the tree is attracting the adults and that there are already numerous individuals inside of it, as I found large swaths of larval galleries under the bark but no obvious adult emergence holes. The tree itself looks healthy and shows no outward sign of stress, and were it not for the evidence of larval galleries I wouldn’t have even suspected it was infested. As with Z. uteanus, this species also has been recorded from Kyle Canyon (Hammond & Williams 2013).

Semanotus juniperi on trunk of large Juniperus osteosperma (Utah juniper) at night.

Over the next hour, I continued inspecting trees in the area, but especially those on which I’d found beetles, and eventually that effort was rewarded with a third S. juniperi on the same tree I collected the previous two. The final collection of the night occurred just after that, when I found not one but two more Z. uteanus, this time on the trunk of P. monophylla. I made one more quick round of the trees, but as it was now midnight I accepted the bounty of the night and turned in.


Days 10 & 11

Unfortunately, I had to end the trip rather abruptly due to a confluence of circumstances at home. The final straw was the rapid decline of Berlioz—our 20-year-old cat (only lifelong cat lovers will understand the bond between a man and his cat). It was not a surprise, and in the morning as soon as I was able to break camp, I left Kyle Canyon and spent the next two days blasting north on I-15 and west on I-70. Sadly, I wasn’t able to make it home before he passed. I’ve had many cats over the course of my life, but King Berlioz was the best!

R.I.P. Berlioz – 10/31/2004 to 6/11/25.
65 hours driving almost 4,000 miles.

©️ Ted C. MacRae 2025

A rare opportunity to do fall insect collecting in western North America

Madam and I are staying in South Lake Tahoe this week, so today I took advantage of this rare opportunity to do some fall insect collecting in western North America. In many insect groups, most species have finished their business for the season by the time autumn arrives. This is especially true for my beloved longhorned beetles, whose spring/early summer flush is now but a distant memory. There are a few genera of longhorned beetles, however, that wait precisely until autumn before making their appearance—Megacyllene and Tragidion being the best-known examples in eastern North America, and Crossidius being the best known out west. I adore autumn collecting, both in the east and in the west—its cooler temperatures and lower humidity make conditions incredibly pleasant, and the longer shadows cast by a low-hanging sun make the landscape—gently morphing from monotonous tones of greens to dazzling shades of amber, tawny, and gold—even more stunning.

Ericameria nauseosa (rubber rabbitbrush) blooming in the valley below a smoke-shrouded eastern slope of the Sierra Nevada.

I figured my best shot for Crossidius was in the sagebrush habitat below the eastern slope of the Sierra Nevada, so I took the Kingsbury Grade over Dagget Pass and found some good looking habitat near Genoa with nice stands of Ericameria nauseosa (rubber rabbitbrush)—host of Crossidius coralinus—in peak bloom. It took quite a bit of work to find the beetles, and for the first half-hour the only insects I saw were non-native honey bees (eventually I did see a few native bees as well, which I collected for Mike Arduser). I also checked some Asclepias speciosa (showy milkweed) that I found hoping to find Tetraopes femoratus (red-femured milkweed borer) but saw only Polistes dominula (European paper wasp).

Polistes dominula (European paper wasp) on Asclepias speciosa (showy milkweed) planted in sagebrush habitat.

After quite a bit of seeching, I finally found a Crossidius adult, although it was not C. coralinus but rather the all-black C. ater.

Crossidius ater on inflorescence of Ericameria nauseosa (rubber rabbitbrush) in sagebrush habitat.

Very quickly afterwards, however, I found the first C. coralinus, and over the next half-hour I collected about a dozen males and females of C. coralinus and one more C. ater.

Crossidius coralinus temprans on inflorescence of Ericameria nauseosa (rubber rabbitbrush) in sagebrush habitat.

Crossidius coralinus has been divided into several subspecies based on differences in markings and coloration across its geographical range—in this area, the species is assignable to subspecies C. coralinus temprans. I did a final check of the stands near where I parked and found one more adult C. coralinus, after which I decided I’d seen enough and went to another spot to look for different species of Crossidius.

Newly-hatched stink bug (superfamily Pentatomoidea) nymphs on inflorescence of Ericameria nauseosa (rubber rabbitbrush) in sagebrush habitat.

After having such good luck with C. coralinus on E. nauseosa, I went a few miles further north to Jack’s Valley Habitat Management Area where I saw a different species of rabbitbrush (Chrysothamnus viscidiflorus, or yellow rabbitbrush) in bloom.

Chrysothamnus viscidiflorus (yellow rabbitbrush) blooming in the valley below a smoke-shrouded eastern slope of the Sierra Nevada.

Yellow rabbitbrush is the host of a different species of Crossidius in this area—C. hirtipes, and I hoped I might be able to find this species as well. It didn’t take long, as I found a small female hiding in the inflorescence of one of the first rabbitbrush plants that I looked at. As sometimes happens, however, my early success was followed by an extended time without seeing another beetle.

Sagebrush habitat below the east slopes of a smoke-shrouded Sierra Nevada.

The plants were sparse in the sagebrush habitat, and I zigzagged my way from plant to plant going west from the trailhead. Despite the lack of beetles, there were other insects active on the flowers, including assassin bugs and digger wasps.

Zelus tetracanthus (four-spurred assassin bug) on inflorescence of Chrysothamnus viscidiflorus (yellow rabbitbrush) in sagebrush habitat.
Sphex ichneumoneus (great golden digger wasp) on inflorescence of Chrysothamnus viscidiflorus (yellow rabbitbrush) in sagebrush habitat.

Eventually, I came upon an area where the plants were more abundant, and I started encountering beetles more regularly, including several big males perched atop the flowers and another C. ater adult hiding inside the flowers on one of these same plants.

Crossidius hirtipes immaculatus on inflorescence of Chrysothamnus viscidiflorus (yellow rabbitbrush) in sagebrush habitat.

Bees—other than honey bees—were scarse, but I managed to pick up a few for Mike, and I also collected a few Eleodes sp. (desert stink beetles) that were, for some reason, crawling and perching on the rabbitbrush flowers. I’ve never thought of darkling beetles—at least those in the genus Eleodes—as anthophilous, and maybe this habit isn’t strict anthophily, per se, but it was remarkable to see so many of these beetles associated with the plants and hardly a single individual crawling about on the ground.

Eleodes sp. (desert stink bug) on inflorescence of Chrysothamnus viscidiflorus (yellow rabbitbrush) in sagebrush habitat.

After a couple hours worth of effort, I’d secured about a dozen C. hirtipes, which, like C. coralinus, has been assigned to several subspecies (populations in this area being assignable to subspecies C. hirtipes immaculatus)—enough for my purposes, and I left to head back up into the mountains.

Smoke and clouds combine above the eastern slope of the Sierra Nevada.

I’d hoped to find one more spot up in the mountains, but the sinking sun and rising elevation conspired to make conditions much too cool for all but the most persistent insects. I did stop at a powerline cut, where recently-cut Jeffrey pine (Pinus jeffreyi) trunks, stumps, and branches provided a perfect attractor for wood-boring beetles. Sadly, none were seen, even on the undersides of the logs and branches. Thus, my rare chance for fall insect collecting in western North America came to a close.

©️ Ted C. MacRae 2022

Big, black (and red), and beautiful!

While I may have already declared Plinthocoelium suaveolens (bumelia borer) as North America’s most beautiful longhorned beetle, any short list of top candidates for this title must also include the species Crossidius coralinus. Like most other members of this strictly North American genus, these gorgeous beetles emerge as adults during late August and September to feed on the profusion of yellow blooms put forth by their larval host plants, Ericameria nauseosa (gray rabbitbrush). Across the Great Basin and adjacent areas, the relatively large size, spectacularly long antennae, and stunning value contrast between red/black or orange/black beetles, yellow flowers, and blue skies combine to make the sight of C. coralinus a highlight on any fall insect collecting trip. If beauty alone isn’t enough, the species also exhibits an unusual level of polytopism across its range. Red in some areas (e.g., C. c. temprans), orange in others (e.g., C. c. monoensis), bigger or smaller, and varying degrees of development of the black areas that cover the basal edge and apical portion of the elytra, the species segregates into several described subspecies and many more unnamed but locally distinct populations. This post features photos of individuals from several populations that field-mate Jeff Huether and I encountered during last August’s Great Basin collecting trip.

Crossidius coralinus coralinus

Crossidius coralinus ssp. coralinus (male) | Montezuma Co., Colorado

Crossidius coralinus coralinus

Crossidius coralinus ssp. coralinus (female) | Montezuma Co., Colorado

One of the most impressive populations I’ve encountered is illustrated by the male and female individuals shown in the above photos, which were seen near the city of Cortez in southwestern Colorado. Linsley & Chemsak (1961) assigned specimens from this area to the nominate subspecies, characterizing them as “moderate-sized”; however, some of the individuals that we encountered at this site were truly gargantuan (exceeding 20 mm in length). Note how extensive the black areas are in these individuals, especially the female.

Crossidius coralinus jocosus (male) | Costilla Co., Colorado

Crossidius coralinus ssp. jocosus (female?) | Costilla Co., Colorado

Crossidius coralinus jocosus (female) | Costilla Co., Colorado

Crossidius coralinus ssp. jocosus (female) | Costilla Co., Colorado

On the other side of the state near Fort Garland (southeastern Colorado) we encountered a population that Linsley & Chemsak (1961) considered representative of the subspecies C. c. jocosus. In contrast to the larger size and extensive black markings of the nominotypical population we found near Cortez, individuals in this population were considerably smaller in size and exhibited less extensively developed black areas of the elytra. Their small size also made them a little harder to notice—perhaps that is the reason we found so few individuals (~7 total at several sites along Hwy 160). We did note also, however, that the gray rabbitbrush flowers seemed to be well past their prime, so perhaps an earlier appearance of the rains upon which plant flowering and beetle emergence rely had us on the tail end of their activity period.

Crossidius coralinus coralinus

Crossidius coralinus ssp.? (male) | San Juan Co., Utah

Crossidius coralinus coralinus

Crossidius coralinus ssp.? (female) | San Juan Co., Utah

Linsley & Chemsak (1961) noted several populations across middle and southern Utah, but the only one to which they assigned a name was C. c. coccineus in Washington Co. (southwestern Utah). While we didn’t visit Washington Co. on this trip, we did look for these beetles at several sites north of Monticello in San Juan Co. (southeastern part of the state). Geography would place this population close to nominotypical populations, and while the beetles in this population resembled them in size they clearly differed in the greatly reduced black areas of the elytra. Note the male especially, with the black area reduced to little more than a sutural stripe in the apical half of the elytra. Linsley & Chemsak (1961) related specimens collected just a few miles further south from ours to an unnamed population in adjacent Wayne Co. (near Hanksville), both of which seem to be close to C. c. coccineus due to their robust size and the reduced black elytral markings of the male.

Crossidius coralinus ssp. (female) | Nye Co., Nevada

Crossidius coralinus ssp.? (female) | Nye Co., Nevada

Crossidius coralinus is found commonly along the western edge of the Great Basin in the form of C. c. temprans. However, Linsley & Chemsak (1961) presented very few records of the species further east in Nevada. We stopped at several spots in central Nevada while traveling along Hwy 6, but despite an abundance of gray rabbitbrush stands in peak bloom we found but a single male and a single female, the latter shown in the above photograph. Geographically this female should be assignable to C. c. temprans, but the black area of the elytra is not nearly so expanded as is typical for that subspecies. The only record from central Nevada in Linsley & Chemsak (1961) is a single male from White Pine Co. (a little further to the east), but they related that specimen to an unnamed population near Marysvale in Piute Co., Utah of smaller size and with the black area of the elytra distinctly expanded in both sexes.

The author photographing insects on flower head of Ericameria nauseosus.

The author photographs insects on gray rabbitbrush in San Juan Co., Utah.

There are those who say “Subspecies, schmubspecies!” I concede they may be right for a large number of named subspecies, possibly including C. coralinus, and while the basin and range topology of the Great Basin and discontinuous distribution of host plants within that geography provide ideal conditions for the development of distinctive, geographically based populations, I suspect C. coralinus has sufficient mobility to allow gene flow across its range (with the possible exception of populations in California’s Central and Owens Valleys). Moreover, the inability of Linsley & Chemsak (1961) to segregate the central Great Basin populations into discrete taxonomic units suggests that the subspecies concept may not be applicable. Nevertheless, it cannot be denied that distinctive, localized populations of this species do exist. Moreover, I hesitate to dismiss subspecies in problematic taxa such as C. coralinus, because doing so makes it easier to ignore variability and presume (possibly incorrectly) no geographic component. Variability is interesting and should be thoroughly evaluated to determine its basis regardless of its basis. Geographically based variability is especially interesting because it suggests the existence of distinct genetic traits that contribute to the genetic diversity of species. Such traits are valuable to protect, and the use of subspecies provides a convenient shorthand for referring to the populations that contain them in both taxonomic and conservation contexts.

REFERENCE:

Linsley, E. G. & J. A. Chemsak. 1961. A distributional and taxonomic study of the genus Crossidius (Coleoptera, Cerambycidae). Miscellaneous Publications of the Entomological Society of America 3(2):25–64 + 3 color plates.

© Ted C. MacRae 2014

Black is beautiful!

Most species in the genus Crossidius exhibit varying amounts of yellow/red/orange coloration on the body. However, one species—Crossidius ater—dispenses with such adornments and remains all-black throughout its expansive range across the Great Basin and surrounding areas. Despite this, they are no less pretty than their more gaudily colored relatives. The adults in these photos were seen on flowers of yellow rabbitbrush (Chrysothamnus viscidiflorus) at two locations in White Pine County, Nevada on 30 August 2013 (all photos w/ natural sky background).

Crossidius ater | 7.2 mi SW NV318 on US6, White Pine Co., Nevada.

Crossidius ater | 7.2 mi SW NV318 on US6, White Pine Co., Nevada.

Crossidius ater | Ely,  1.2 mi S Jct US6/50/93, White Pine Co., Nevada.

Crossidius ater | Ely, 1.2 mi S Jct US6/50/93, White Pine Co., Nevada.

Crossidius ater | Ely,  1.2 mi S Jct US6/50/93, White Pine Co., Nevada.

Crossidius ater | Ely, 1.2 mi S Jct US6/50/93, White Pine Co., Nevada.

© Ted C. MacRae 2014

A Crossidius hirtipes subspecies blend zone…

…or, “There’s something fishy going on here!”

After a day in the vicinity of Yearington, Nevada  looking for (and eventually finding) a population of Crossidius hirtipes longhorned beetles assignable to subspecies “rubrescens“, field mate Jeff Huether and I dropped a little further south to look for two additional subspecies—C. h. immaculipennis and C. h. macswaini. Like C. h. rubrescens, populations assignable to these two subspecies are also restricted to a small area in west-central Nevada. Chemsak & Linsley (1959) described C. h. immaculipennis from specimens collected 10 miles north of Smith (Lyon Co.) and two years later (Linsley & Chemsak 1961) described C. h. macswaini from 19 miles SE of Wellington (Douglas Co.). We were still under the smoke plum from the now 9-day old Rim Fire in the nearby Sierra Nevada, which came and went during much of the day (top photo) and eventually settled in to create some amazing sunset landscapes (bottom photo).

19 mi S of Wellington, Toiyabe National Forest, Nevada.

Rim Fire smoke moves in and out of the area | 19 mi S of Wellington, Toiyabe National Forest, Nevada.

As had happened the day before with C. h. rubrescens, success did not come quickly or easily. We first searched for the type locality of C. h. immaculipennis, but many parts of the area have been converted to agriculture, and in the precise area 10 miles north of Smith we found only vast fields of dark green, irrigated alfalfa and not the rabbitbrush chaparral habitat required by these beetles. We did a little searching in surrounding areas and eventually found one rather nice-looking patch of ground with healthy stands of Chrysothamnus viscidiflorus in full bloom. Mindful of the previous day’s experience with finding the beetles often ensconced down within the inflorescences—especially as cooler temperatures set in, we took care to carefully inspect the blossoms in case the cool morning temperatures were inhibiting the beetles from coming back up for the day. Alas, we saw no beetles despite spending a considerable amount of time searching.

Crossidius hirtipes immaculipennis (male) | 6.3 mi W of Wellington, Nevada.

Crossidius hirtipes immaculipennis (male) | 6.3 mi W of Wellington, Nevada.

Rather than beat a dead horse, we decided to drive a short distance south and west to the town of Wellington, where a fellow cerambycid collector had found the subspecies a few years earlier. He had provided us with some detailed notes on the specific localities where he found the beetles, and these proved to be of great help as we passed through endless, seemingly acceptable chaparral habitat trying to decide exactly where we should stop and invest more time to look for the beetle. We stopped at one of the sites indicated in the notes and immediately found a beetle on one of the first plants we checked, and eventually after a gap in finding any more we found an area where good numbers of the plants were supporting decent numbers of the beetles. Chemsak & Linsley (1959) distinguished C. h. immaculipennis by its reddish legs, pale color, and complete lack of elytral markings in the male, exemplified by the male shown in the photo above.

Crossidius hirtipes macswainei? (female) | 6.3 mi W of Wellington, Nevada.

Crossidius hirtipes macswainei? (female) | 6.3 mi W of Wellington, Nevada.

As we searched the plants and found more and more individuals, I noticed an occasional adult that seemed to be a little more yellowish than reddish and with distinct sutural maculae. I didn’t think much about it then, chalking it up to individual variability, but after returning home and having a chance to look at the specimens more closely I was surprised to determine that these few beetles actually are a better fit for the second subspecies we planned to search for that day—C. h. macswainei. We had found both subspecies at the same site and didn’t even realize it. Okay, I know what you’re thinking… subspecies must exhibit allopatric geographic distributions (cannot occur together at the same place and time). It is, thus, tempting to declare that the two “subspecies” are actually not distinct, but rather represent distinctive extremes of individual variation in a single interbreeding population. However, one must also consider the possibility that the two subspecies represent reproductively isolated populations and, thus, qualify as distinct species. I’m not qualified to make that judgement, but I will note that most of the individuals encountered were assignable to C. h. immaculipennis  and the rest to C. h. macswainei, but that no “intergrades” were found.

Crossidius hirtipes macswainei (male) | 19 mi S of Wellington, Nevada.

Crossidius hirtipes rhodopus? (male) | 19 mi S of Wellington, Nevada.

After collecting adequate series from W of Wellington, we traveled further south of town to the type locality of C. h. macswainei (not knowing we already had it!). The holotype and most of the paratypes were collected 19 miles S of Wellington (Linsley & Chemsak 1961), but a number of paratypes had also been collected 14 miles south of town, so we stopped there first in an unsuccessful bid to find the subspecies before moving on to the type locality a few miles further south. Within a few minutes of arriving, I found the individual shown in the photo immediately above, presumed that I had found the subspecies we were looking for, and popped it into a vial alive as a photo backup if we did not find any other individuals with which I could attempt field photographs. Ironically, that is exactly what happened—despite Jeff and I scouring every plant we could find in about a 1-mile stretch along each side of the road, we never found another beetle. Later that evening I took the above individual out of its vial for photographs, but it never really “perked up” to look fully natural, resulting in “bum” antennae that give away the staged nature of the photograph. Again, it was not until I got back home and could look at the specimen closely before I realized that it did not at all fit the description of C. h. macswainei, but instead seemed to be a good match for the subspecies C. h. rhodopus, known from only a short distance further south but—until now, at least—apparently restricted to the Mono Basin in east-central California (see this post for more details about this subspecies). Jeff has since reported to me that some of the beetles he collected at the “C. h. rubrescens” locality (see this post) also are a match for C. h. macswainei, adding yet another wrinkle to those that resulted from this day’s collecting. Such inconsistencies with the published literature may tempt some to scrap all of Linsley & Chemsak’s subspecies, but considering that those two authors examined more than 12,000 specimens during the course of their studies such a reaction would be both premature and presumptuous. What is needed is more study—more specimens from more localities, hopefully augmented with DNA sequence analysis. For the latter goal we did our part, dropping a specimen or two from every locality in which we found beetles into ethanol for just such purpose. Until such studies are done, I prefer to withhold judgement about whether C. hirtipes is comprised of one highly polytopic population, several subspecifically distinct populations, or perhaps even multiple distinct species.

14 mi SE of Wellington, Toiyabe National Forest, Nevada.

Evening haze creates a spectacular sunset | 14 mi SE of Wellington, Toiyabe National Forest, Nevada.

Even though we found only a single beetle at the second locality, our persistence in searching until the day ran out was rewarded by a most spectacular sunset caused by thick haze from the nearby Rim Fire in California. It would also be our last day in Nevada before dropping south into California and spending the next several days in successful bids for C. coralinus monoensisC. c. caeruleipennisC. h. nubilus, and C. h. rhodopus.

REFERENCE:

Chemsak, J. A. & E. G. Linsley. 1959. Some new species and subspecies of Crossidius from western North America. Journal of the Kansas Entomological Society 32(4):176–183.

Linsley, E. G. & J. A. Chemsak. 1961. A distributional and taxonomic study of the genus Crossidius (Coleoptera, Cerambycidae). Miscellaneous Publications of the Entomological Society of America 3(2):25–64 + 3 color plates.

Copyright © Ted C. MacRae 2014

GBCT Beetle #4—Crossidius hirtipes rubrescens

Crossidius hirtipes rubrescens (male) | Lyon Co., Nevada

Crossidius hirtipes rubrescens (male) | Lyon Co., Nevada

Linsley & Chemsak (1961) characterize the longhorned beetle species Crossidius hirtipes as “probably the most diverse species within the genus.” Occurring throughout the Great Basin and adjacent areas, the variability it expresses in punctation, color, elytral pattern and pubescence have resulted in the recognition of 16 named subspecies and numerous additional distinct but unnamed populations. During our Great Basin Collecting Trip (GBCT) in late August, we targeted ten of the named subspecies (representing the majority of populations found across the southern half of the species’ distribution) and succeeded in finding eight of them. I’ve already featured C. h. immaculatus, occurring across northern Nevada and northeastern California and the first beetle we found on the trip, and the photo above shows a male of what we consider to represent C. h. rubrescens—one of three C. hirtipes subspecies restricted to a very small area in west-central Nevada. Linsley & Chemsak (1961) described this subspecies from a series taken just north of Yearington on Chrysothamnus greenei (now Ericameria greenei), noting that it differs from other subspecies by its pinkish cast to its coloration and its pale appendages.

We almost did not find this subspecies.  We had stopped at several places along the road as we approached Yearington from the north and finally stopped at a spot 2.6 miles north of town with good stands of E. greenei. Although we found a few C. coralinus temprans on the plants, we did not see C. hirtipes. While we were searching we noticed a much smaller yellow-flowering asteraceous plant that at first we thought might be something in the genus Haplopappus but which I now believe represents a variety of Chrysothamnus viscidiflorus—the normal host plant for most subspecies of C. hirtipes (a plant voucher was collected and has been sent off for identification). We searched these plants as well but didn’t see any beetles on them, and after a while we decided we’d given the locality a good enough look and that we should move on. We began walking back towards the car, and as we approached the car I happened to look down and saw a mating pair of C. hirtipes sitting on a C. viscidiflorus flower. The pair split and bolted right when I saw them, but we managed to capture one of them and decided maybe we should look around a little more. The beetles were scarce, and another hour of searching produced only a handful—mostly in a small area further north of the area we had been searching. We then checked a couple of other nearby spots but found only a few host plants and no beetles, so we decided to go back to the site and search again. While none were seen in the original spot, we found much better stands of the plant in the adjacent area even further to the north and managed to collect a decent though not large series of adults before the setting sun caused the beetles to retreat and end our day. The individual in the photo above (recognizable as a male by its relatively longer antennae and immaculate elytra) was photographed as the setting sun turned the smoke-filled sky to a soft, burnt orange color that nicely compliments the color of the beetle.

REFERENCE:

Linsley, E. G. & J. A. Chemsak. 1961. A distributional and taxonomic study of the genus Crossidius (Coleoptera, Cerambycidae). Miscellaneous Publications of the Entomological Society of America 3(2):25–64 + 3 color plates.

Copyright © Ted C. MacRae 2013

Under Blood Red Skies

At the start of my recent Great Basin collecting trip, I found myself presented with a rather unique and unanticipated photographic opportunity. As I landed in Reno, Nevada, the then 6-day old Rim Fire was already well on it’s way to becoming the largest wildfire on record in the California Sierra Nevada. As acre after acre of the Sierra’s dramatic coniferous forest succumbed to the blaze, an enormous plum of smoke drifted northward for several hundred miles over eastern California and western Nevada, blanketing the area in a thick haze that turned the sun’s hot glare to a soft glow and limited visibility to under a mile. It was like a thick overcast foggy day, only without the cool, damp humidity. This was of little consequence to our business at hand—collecting beetles (although it did make pointless most attempts to photograph the area’s stunning landscape). At day’s end, however, a dramatic transformation took place in the sky as the sun sank lower and lower, turning to an increasingly red globe as it strained to shine through the ever thicker layer of smoke and haze. Then, for a few brief moments, the sun floated—a dark red globe under blood red skies—before the thick bottom layers of haze finally extinguished its fading light.

I’ve just begun trying to incorporate setting suns into my photography, having made to this point only a few attempts over Midwestern landscapes. I’m not really sure what gave me the idea, but I thought it might be fun to try incorporating the spectacular sun and unusual sky I was seeing as backgrounds in full-flash insect macrophotographs. Perhaps it seemed a logical progression from the natural sky background macrophotographs that I’ve put a lot of effort into perfecting this year. It was certainly a learning experience, but the basic principle is the same as it is for blue sky background—finding the right combination of camera and flash settings to balance flash illumination of the subject with ambient illumination of the background. The most difficult thing was, surprisingly, getting the sun in the desired position within the composition, as it does not appear through the viewfinder as the discrete ball that is seen in the photos. Rather, it appears as a large, amorphous, blinding flash that comes and goes as one pans across it, leading to a lot of guesswork regarding its actual position within the composition.

I gave a sneak preview of one of these photos in Sunset for another great collecting trip, and several of the photos I’ve shared since then have featured the remarkably colored sky in the background. Here are some other attempts that I was happy with:

Agrilus walsinghami (female) | Washoe Co., Nevada

Agrilus walsinghami (female) | Washoe Co., Nevada

Agrilus walsinghami (male) | Washoe Co., Nevada

Agrilus walsinghami (male) | Washoe Co., Nevada

Crossidius coralinus temprans (male) | Pershing Co., Nevada

Crossidius coralinus temprans (male) | Pershing Co., Nevada

Initially hot yellow (previous photo), the sun turns to soft yellow...

Initially hot yellow (previous photo), the sun turns to soft yellow…

...then yellow-red...

…then yellow-red…

...and finally blood-red!

…and finally blood-red!

Crossidius hirtipes macswainei (female) | Lyon Co., Nevada

Crossidius hirtipes macswainei (female) | Lyon Co., Nevada

Sunset over Toiyabe National Forest | Lyon Co., Nevada

Sunset over Toiyabe National Forest | Lyon Co., Nevada

Gray rabbitbrush (Ericameria nauseosa) | Lyon Co., Nevada

Gray rabbitbrush (Ericameria nauseosa) | Lyon Co., Nevada

Copyright © Ted C. MacRae 2013